Diptera: Sarcophagidae

Ann. soc. entomol. Fr. (n.s.), 2011, 47 (3–4) : 303-329

ARTICLE

A review of subtribe Boettcheriscina Verves 1990 (Diptera: Sarcophagidae), with descriptions of a new species and genus from China Wan-Qi Xue (1), Yuriy G. Verves (2) & Jing Du (3) (1)

Institute of Entomology, Shenyang Normal University, Shenyang, 110034, China (2) Postbox 23, Kyiv, 03118, Ukraine (3) College of life sciences, Nankai University, Tianjin, 300071, China

Abstract. A review of subtribe Boettcheriscina with a key to genera is present. Subtribe consists from 18 genera and 72 species, widely distributed in all zoogeographical regions, except Antarctic. The description of a new genus, Fanzideia, n. gen., with type species Fanzideia cygnocerca, n. sp., is presented. 14 new combinations of specific names are given: Boettcherisca krathonmai (Pape & Bänzinger 2000), n. comb.; B. talomoensis (Magpayo & Kano 1986), n. comb.; B. yuwanensis (Sugiyama 1990), n. comb.; Johnstonimyia paineiana (Baranov 1934), n. comb.; Lioproctia (Coonoria) aureolata (Pape & Kurahachi 2000), n. comb.; L. (C.) kurahashii (Shinonaga & Tumrasvin 1979), n. comb.; L. (C.) saprianovae (Pape & Bänzinger 2000), n. comb.; L. (C.) serracudo (Pape & Kurahachi 2004), n. comb.; L. (C.) sumbaensis (Shinonaga 2004), n. comb.; L. (C.) sundaensis (Shinonaga 2004), n. comb.; L. (C.) vietnamensis (Shinonaga & Thinh 2003), n. comb.; Rosellea suthep (Pape & Banziger 2003), n. comb.; Latystyla czernyi (Böttcher 1912), n. comb.; L. paularnaudi (Lehrer 1981), n. comb. 16 new synonyms are established, among them 3 generic: Macabiella Lehrer 1994, n. syn. = Latistyla Strukan 1970; Leigongshanophaga Lehrer & Wei 2010, n. syn. = Rosellea Rohdendorf 1937; Ussuriphalla Lehrer 2010, n. syn. = Takanoa Rohdendorf 1965, and 13 specific: Burmanomyia parvatia Lehrer 2008, n. syn. = Sarcophaga beesoni Senior-White 1924; Burmanomyia guanyina Lehrer & Wei 2010, n. syn. = Sarcophaga beesoni Senior-White 1924; Liproctia kunlunea Lehrer 2008, n. syn. = Sarcophaga pattoni Senior-White 1924; Phallosphaera huangdinia Lehrer 2008, n. syn. = Sarcophaga gravelyi Senior-White 1924; Phallosphaera jimmuana Lehrer 2008, n. syn. = Sarcophaga gravelyi Senior-White 1924; Rosellea fuxingia Lehrer 2010, n. syn. = Sarcophaga khasiensis SeniorWhite 1924; Rosellea manipuriella Lehrer 2010, n. syn. = Sarcophaga khasiensis Senior-White 1924; Rosellea longwangiana Lehrer & Wei 2010, n. syn. = Sarcophaga khasiensis Senior-White 1924; Sarcophaga (Johnstonimyia) hugoi Pape 1996, n. syn. = Johnstonimyia lopesi Shinonaga et Kano 1990; Sarcophaga gorokaensis Sugiyama, Shinonaga & Kano 1988, n. syn. = Johnstonimyia gressitti Kano & Lopes 1981; Sarcophaga (Lioproctia) imita Pape 1996, n. syn. = Johnstonimyia imitatrix Lopes 1959; Takanoa vervesiana Lehrer 2010, n. syn. = Sarcophaga hakusana Hori 1954; Ussuriphalla qirimia Lehrer 2010, n. syn. = Takanoa rugosa Rohdendorf 1969. Résumé. Une revue de la sous-tribu Boettcheriscina Verves 1990 (Diptera: Sarcophagidae), avec la description d’une nouvelle espèce d’un nouveau genre de Chine. Une révision de la soustribu Boettcheriscina est présentée, ainsi qu’une clef des genres. Cette sous-tribu comprend 18 genres et 72 espèces largement distribuées dans toutes les régions zoogéographiques, sauf l’Antarctique. La description d’un nouveau genre, Fanzideia, n. gen., est présentée, avec Fanzideia cygnocerca, n. sp. comme espèce-type. 14 nouvelles combinaisons de noms spécifiques sont apportées : Boettcherisca krathonmai (Pape & Bänzinger 2000), n. comb.; B. talomoensis (Magpayo & Kano 1986), n. comb.; B. yuwanensis (Sugiyama 1990), n. comb.; Johnstonimyia paineiana (Baranov 1934), n. comb.; Lioproctia (Coonoria) aureolata (Pape & Kurahachi 2000), n. comb.; L. (C.) kurahashii (Shinonaga & Tumrasvin 1979), n. comb.; L. (C.) saprianovae (Pape & Bänzinger 2000), n. comb.; L. (C.) serracudo (Pape & Kurahachi 2004), n. comb.; L. (C.) sumbaensis (Shinonaga 2004), n. comb.; L. (C.) sundaensis (Shinonaga 2004), n. comb.; L. (C.) vietnamensis (Shinonaga & Thinh 2003), n. comb.; Rosellea suthep (Pape & Banziger 2003), n. comb.; Latystyla czernyi (Böttcher 1912), n. comb.; L. paularnaudi (Lehrer 1981), n. comb. 16 nouvelles synonymies sont établies, parmi lesquelles 3 génériques, Macabiella Lehrer 1994, n. syn. = Latistyla Strukan 1970; Leigongshanophaga Lehrer & Wei 2010, n. syn. = Rosellea Rohdendorf 1937; Ussuriphalla Lehrer 2010, n. syn. = Takanoa Rohdendorf 1965; et 13 spécifiques: Burmanomyia parvatia Lehrer 2008, n. syn. = Sarcophaga beesoni Senior-White 1924; Burmanomyia guanyina Lehrer & Wei 2010, n. syn. = Sarcophaga beesoni Senior-White 1924; Liproctia kunlunea Lehrer 2008, n. syn. = Sarcophaga pattoni Senior-White 1924; Phallosphaera huangdinia Lehrer 2008, n. syn. = Sarcophaga gravelyi Senior-White 1924; Phallosphaera jimmuana Lehrer 2008, n. syn. = Sarcophaga gravelyi Senior-White 1924; Rosellea fuxingia Lehrer 2010, n. syn.

E-mail: xwqfl[email protected], fl[email protected], [email protected] Accepté le 10 septembre 2011

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W.-Q. Xue, Y. G. Verves & J. Du

= Sarcophaga khasiensis Senior-White 1924; Rosellea manipuriella Lehrer 2010, n. syn. = Sarcophaga khasiensis Senior-White 1924; Rosellea longwangiana Lehrer & Wei 2010, n. syn. = Sarcophaga khasiensis Senior-White 1924; Sarcophaga (Johnstonimyia) hugoi Pape 1996, n. syn. = Johnstonimyia lopesi Shinonaga et Kano 1990; Sarcophaga gorokaensis Sugiyama, Shinonaga & Kano 1988, n. syn. = Johnstonimyia gressitti Kano & Lopes 1981; Sarcophaga (Lioproctia) imita Pape 1996, n. syn. = Johnstonimyia imitatrix Lopes 1959; Takanoa vervesiana Lehrer 2010, n. syn. = Sarcophaga hakusana Hori 1954; Ussuriphalla qirimia Lehrer 2010, n. syn. = Takanoa rugosa Rohdendorf 1969.

Keywords: Sarcophagidae, review, Fanzideia cygnocerca, descriptions.

S

ubtribe Boettcheriscina had been established by Verves (1990: 540). It belongs to tribe Sarcophagini and subfamily Sarcophagidae. 18 genera and 72 known species of this subtribe widely distributed in all zoogeographical regions, exceptly Antarctic. There are grey flies of middle or great sizes (5.0–20.0 mm). In both sexes acr [acrostichal bristles] 0+1; dc [dorsocentral bristles] 2+3–4+7, the anterior postsutural pairs sometimes rather weak and distinctly shorter than posterior ones. The main differences from other subtribes are based on such specific features of male postabdomen: - VII+VIII segment not more than 1.5x as long as wide; - hind ventral angle of epandrium not elongate; - cerci and different parts of aedeagus often covered with small spines; - vesica well developed; - juxta well developed, with or without lateral arms; - lateral styli widened and have different appendages, often covered by spines. Larvae are shizophagous in general, but can be facultative parasites or predators of different invertebrate (terrestrial snails, insects etc.) and vertebrate animals. Terminology follows Merz & Haenni (2000) with few modifications proposed by Povolný & Verves (1997), Verves (2000), Verves & Khrokalo (2006, 2009).

Key to the genera and subgenera of Boettcheriscina 1. 5th abdominal sternite with ventral protuberance ......... 2 - 5th abdominal sternite without ventral protuberance .... 3 2. Protuberance of 5th sternite digitate (Fig. 13, a). Lateral styli with short spines (Fig. 13, c) ............................ ................................................. Phallosphaera Rohdendorf - Protuberance of 5th sternite conic (Fig. 15, a). Lateral styli haired, without spines (Figs 15, c–d) ..................... .......................................................... Rosellea Rohdendorf 3. Antennae, palpi and legs red. Lateral styli spinous; parastyli well developed (Figs 14, a–b) .... Robackina Lopes - Antennae, palpi and legs black ..................................... 4 4. Abdomen with black spots and bands, wings with 3 black spots. Apical part of cerci narrow and pointed. Juxtal filament widened, bilobed, without lateral processes (Fig. 3) ........................... Dasysceloctis Enderlein - Abdomen with checkering pattern or longitudinal medial black stripe, wings without spots ................................. 5

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5. Juxtal filament distinctly shortened than elongate lateral styli ............................................................................. 6 - Juxtal filament so long or longer than lateral styli ........ 8 6. Ventral processes elongate and narrow, hook-shaped, parastyli present (Fig. 18, a, c) .... Sclerophalla Rohdendorf - Ventral processes short or poor developed, parastyli absent .......................................................................... 7 7. Apical part of cerci with spines (Fig. 5, a). Lateral styli directed apically (Figs 5, c–d) ........... Johnstonimyia Lopes - Apical part of cerci without spines. Lateral styli directed ventrally (Fig. 12) ................................. Nyikamyia Lehrer 8. Ventral processes elongate and narrow, bristle-shaped .... 9 - Ventral processes short and wide or poor developed .... 11 9. Vesica petiolate, trilobed. Pregonites bifurcated apically. Juxta well sclerotized. Ventral processes elongate, narrow, strongly curved, with a pair of apical digitate appendages. Apical part of cerci with dorsal row of spines (Figs 16, a–c) ................................. Sabiella Verves - Vesica not petiolate. Pregonites hook-shaped. Juxta poorly sclerotized, membranous. Apical part of cerci haired, without spines ................................................. 10 10. Lateral juxtal processes narrow, bristle-shaped. Vesica straight, lancet-shaped (Fig. 20, b) ... Ziminisca Rohdendorf - Lateral juxtal processes very widened at basis, triangle. Vesica curved, claw-shaped (Fig. 17, d–e) .................. ......................................................... Saputaramyia Verves 11. Cerci short and wide ................................................... 12 - Cerci narrow and elongate ........................................... 13 12. Cerci spinous in apical part (Fig. 6, a–b). Lateral styli and vesica covered with numerous spines; juxtal processes short (Fig. 6, d–e). Propleuron haired ......... ....................................................... Kramerea Rohdendorf - Cerci without spines, covered with hairs (Fig. 4, d–e). Lateral styli and vesica without spines; juxtal processes elongated and bifurcated (Fig. 4, a). Propleuron bare .............................................................. Fanzideia n. gen. 13. Lateral styli short, parastyli well developed, skittlesshaped (Fig. 19, b–c). 4th abdominal sternite with long erected hairs. Propleuron bare ........ Takanoa Rohdendorf - Lateral styli elongate; if shortened, than parastyli absent. 4th sternite shortly haired ............................................. 14 14. Juxta without spines .................................................... 15 - Juxta with numerous spines. Ventral processes short or absent; vesica without spines ....................................... 17 15. Vesica wide, covered with numerous spines ................. 16 - Vesica narrow, hook-liked, without spines (Fig. 11). Propleuron bare ..................................... Latistyla Strukan

A review of subtribe Boettcheriscina

16. Paraphallus almost straight (Fig. 1, a). Propleuron more or less haired ............................. Boettcherisca Rohdendorf - Apical tip of paraphallus hook-liked, curved dorsally (Fig. 10). Propleuron bare .................... Lucyphalla Lehrer 17. Vesica paired, petiolate, widened apically (Fig. 2, b–c). Cerci with a tuft of short spines on subapical dorsal protuberance (Fig. 2, a). Abdomen without checkering pattern, with golden pollinosity and longitudinal median dark stripe at tergites. Propleuron bare ........... .............................................. Chrysosarcophaga Townsend - Vesica not petiolate. Cerci haired, without spines Figs 7, a–b). Abdomen with gray checkering pattern ............ .......................................................... Lioproctia Enderlein a. Genae entirely white haired. Ventral processes absent; vesica well sclerotized, multipointed (Figs 7, d–e). Propleuron more or less haired, sometimes bare ........................................................ s/g Burmanomyia Fan - Genae in fore parts black haired. Ventral processes well developed; vesica bipointed. Propleuron haired ........... b. b. Juxta so long as lateral styli. Medial stylus very short (Fig. 8). Head with white or silver pollinosity, parafacials covered with irregular hairs ................... s/g Coonoria Fan - Juxta distinctly longer than shortened lateral styli. Medial stylus elongate, spinous apically (Figs 9, a–c). Head with golden pollinosity, parafacials with a distinct row of short setae ............................... s/g Lioproctia s. str.

Review of the genera and species Genus Boettcherisca Rohdendorf 1937 Rohdendorf 1937: 51, 270. Type species: Myophora peregrina Robineau-Desvoidy 1830, by original designation. Boettcherisca: Fan 1992: 688; 2002: 86; Fan & Pape 1996: 248; Hardy 1932a: 45; Kano et al. 1967: 17; Kano & Shinonaga 1977: 32; Kano & Sugiyama 1983: 43; Kurahashi & Kano 1984: 27; Kurahashi & Ohtaki 1989: 291; Lehrer 2003: 37, 126; Lopes 1961a: 69; Lopes et al. 1977: 565; Nandi 1992a: 34; 2002: 204; Rohdendorf 1963: 3, 10; 1965: 684, 694; Verves 1986a: 161; 1990: 541; Verves & Khrokalo 2006: 77, 175. Notochaetomima Rohdendorf 1937: 273, as subgenus of Boettcherisca. Type species: Boettcherisca septentrionalis Rohdendorf 1937, by original designation. Athyrsiola Baranov 1938: 174. Type species: Athyrsia atypica Baranov 1934 [= Sarcophaga invaria Walker 1859], by original designation. Sarcophaga (Boettcherisca): Pape 1996: 309.

16 species are distributed in Palaearctic, Oriental, Australasian/Oceanian and Madagascan regions.

Boettcherisca bengalensis Nandi 1992 Boettcherisca bengalensis Nandi 1992a: 35. Type locality: India: West Bengal: Midnapore, Arabari Forest. Holotype (♂) deposited in Zoological Survey of India, Kolkata, India. Boettcherisca bengalensis: Nandi 2002: 221; Verves 2001: 242. Sarcophaga (Boettcherisca) bengalensis: Pape 1996: 309.

Oriental region: India: Gujarat, Maharashtra, West Bengal. Flies were collected by bush-sweeping.

Boettcherisca cabrerai Kano & Sugiyama 1983. Boettcherisca cabrerai Kano & Sugiyama 1983: 43. Type locality: Philippines: Luzon: Los Banos, Mt. Maquiling. Holotype (♂) deposited in National Science Museum, Tokyo, Japan. Boettcherisca cabrerai: Verves 1990: 542. Sarcophaga (Boettcherisca) cabrerai: Pape 1996: 309.

Oriental region: Philippines: Luzon. The adult flies were found in mountain jungle at altitude about 500 m a. s. l. and attracted by dead fish and meat.

Boettcherisca dumoga (Sugiyama & Kurahashi 1988) Sarcophaga (Boettcherisca) dumoga Sugiyama & Kurahashi 1988: 49. Type locality: Indonesia: Sulawesi: Sulawesi Utara, Toraut, Taman National Park, Dumoga-Bone. Holotype (♂) deposited in Bogor Museum, Java, Indonesia. Sarcophaga (Boettcherisca) dumoga: Pape 1996: 309. Sarcophaga dumoga: Blackith & Blackith 1988: 301. Boettcherisca dumoga: Verves 1990: 542.

Oriental region: Indonesia: Sulawesi. The flies were reared from spoiled meat baited with fly trap which was set for canopy sampling at 40 m above ground. Development from newborn 1st stage larva to adult is continued 15–19 days.

Boettcherisca formosensis Kirner & Lopes 1961 Boettcherisca formosensis Kirner & Lopes 1961: 65. Type locality: China: Taiwan: 20th km of Taipei-Taokian road. Holotype (♂) deposited in Institute Oswaldo Cruz, Rio de Janeiro, Brazil. Boettcherisca formosensis: Fan 1992: 690; 2002: 86; Fan & Pape 1996: 248; Kano & Sugiyama 1983: 46; Lopes 1961a: 79; So & Dudgeon 1989a: 349; 1989b: 113; 1990: 337; Verves 1986a: 161; 1990: 542. Sarcophaga (Boettcherisca) formosensis: Lin & Chen 1999: 117; Pape 1996: 310; Sugiyama et al. 1987: 70.

Oriental region: China: Guangdong, Taiwan, Zhejiang. Larvae necrophagous; adult flies were collected at altitudes to 1000 m a. s. l.

Boettcherisca invaria (Walker 1859) Sarcophaga invaria Walker 1859: 103. Type locality: Indonesia: Maluku: Aru Is. Holotype (♂) deposited in Natural History Museum, London, United Kingdom. Sarcophaga invaria: Blackith & Blackith 1988: 301; Sugiyama et al. 1988a: 284. Sarcophaga (Boettcherisca) invaria: Pape 1996: 310; Shinonaga 2004: 282. Boettcherisca invaria: Kano & Sugiyama 1983: 45; Kurahashi & Kano 1984: 27; Lopes & Kano 1979a: 305; Verves 1990: 542. Athyrsiola atypica Baranov 1934: 183. Type locality: Solomon Is.: Santa Isabel I., Festiva. Holotype (♂) deposited in Natural History Museum, London, United Kingdom. Athyrsiola atypica: Baranov 1938: 174. Boettcherisca atypica: Lopes 1961a: 81; 1967: 165.

Oriental region: Indonesia: Sulawesi. Australasian/ Oceanian region: Indonesia: Maluku (Aru Is.); Papua

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New Guinea: Bismarck Arch. (Manus, New Ireland), New Guinea; Solomon Is.: Santa Isabel I., Tulagi I. Flies attracted to destroyed animal matters in the forests and not more than 3 km inside of them. Larvae necrophagous; development from newborn 1st stage larva to adult is continued 15–19 days.

Boettcherisca javanica Lopes 1961 Boettcherisca javanica Lopes 1961a: 79. Type locality: Indonesia: Java: Soekaboemie, “Croisiere du Nirvana”. Holotype (♂) deposited in Institute Oswaldo Cruz, Rio de Janeiro, Brazil. Boettcherisca javanica: Kano & Sugiyama 1983: 45; Verves 1990: 542. Sarcophaga (Boettcherisca) javanica: Bänzinger & Pape 2004: 1677; Pape 1996: 310; Shinonaga 2004: 282.

Oriental region: Indonesia: Bali, Java, Kalimantan, Sumatra; Malaysia: West Malaysia; Singapore; Philippines; Thailand. Flies were collected in evergreen forests.

Boettcherisca karnyi (Hardy 1927) Sarcophaga karnyi Hardy 1927: 454. Type locality: Indonesia: Java: Bogor [as “Buitenzong”]. Holotype (♂) deposited in Bogor Museum, Java, Indonesia. Sarcophaga karnyi: Blackith & Blackith 1988: 301; Hardy 1932a: 45; Ho 1938: 115; Pape 1990: 110; Senior-White 1931: 74; Senior-White et al. 1940: 273. Sarcophaga (Boettcherisca) karnyi: Bänzinger & Pape 2004: 1677; Pape 1996: 310; Shinonaga 2004: 282. Boettcherisca karnyi: Kano & Sugiyama 1983: 47; Kurahashi & Kano 1984: 27; Lopes 1958: 17; 1961a: 78; 1967: 165; Lopes et al. 1977: 565; Nandi 1990: 117; 2002: 208; Verves 1990: 542; 2001: 243. Sarcophaga fuscicauda: Senior-White 1924: 252; in part [misidentification: not Sarcophaga fuscicauda Böttcher 1912]. Sarcophaga peregrina: Hall & Bohart 1940: 129 [misidentification: not Sarcophaga peregrina Robineau-Desvoidy 1830] Sarcophaga “near peregrina”: Bohart & Gressitt 1951: 136.

Oriental region: Cocos Is.: West I.; India: Andaman Is.; Indonesia: Bali, Java, Kalimantan, Sulawesi, Sumatra; Malaysia: West Malaysia, East Malaysia (Sarawak); Myanmar; Philippines: Tawi Tawi; Thailand. Australasian/Oceainian region: Guam I.; Indonesia: Moluccas (Ambon, Ceram); Marchall Is.: Ailinglapalap, Ebon, Majuro, Namorik, Ujelang; Marianas: Agrihan, Pagan, Rota, Tinian; Micronesia: Caroline Is. (Elato, Ifaluk, Kusaie, Lukunor, Ngulu, Nomwin, Pingelap, Ponape, Pulo Anna, Ulithi), Truk Is. (Wena), Yap Is. (Map, Yap); Palau Is.: Angaur, Babelthuap, Koror, Malakal, Ngaiangl, Ngerkabesang, Ngurukdabel, Peleliu, Ulebsehel; Wake I. Flies attracted to destroyed animal matters in the forests and not more than 3 km inside of them, and on sand beach. Larvae developed in decaying vegetable, dead terrestrial snails, insects (orthopteran Gryllotalpa sp. and beetle Xylotrupes sp.), fishes, frogs, lizards, and beef. Larvae development from newborn 1st stage larva to adult is continued 16 days in general.

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Boettcherisca koimani Kano & Shinonaga 1977 Boettcherisca koimani Kano & Shinonaga 1977: 323. Type locality: Indonesia: Flores Is.: Maumere, 10 m a. s. l. Holotype (ff) deposited in Bogor Museum, Java, Indonesia. Boettcherisca koimani: Kano & Sugiyama 1983: 45; Verves 1990: 542. Sarcophaga (Boettcherisca) koimani: Pape 1996: 310.

Oriental region: Indonesia: Flores Is. Adult flies prefer the forests; larvae are developed on decayed meat and fishes in laboratory conditions.

Boettcherisca krathonmai Pape & Bänzinger 2000, n. comb. Sarcophaga (Boettcherisca) krathonmai Pape & Bänzinger 2000: 204. Type locality: Thailand: Ranong Province, Phya Nak Falls, Khlong Naka Wildlife Sanctuary. Holotype (♂) deposited in Museum of Natural History, Stockholm, Sweden. Sarcophaga krathonmai: Bänzinger & Pape 2004: 1677.

Oriental region: Malaysia: West Malaysia (Perak); Thailand. Flies were collected from flowers of Rafflesia cantleyi Solms-Laubach 1910 and Sapria ram Bänziger & B. Hansen 1997 in lowland evergreen forests. Female are registered during larviposition on carrion; larvae bred on liver in laboratory conditions.

Boettcherisca nathani Lopes 1961 Boettcherisca nathani Lopes 1961a: 79. Type locality: India: Tamil Nadu: Korumbagaram. Holotype (♂) deposited in Institute Oswaldo Cruz, Rio de Janeiro, Brazil. Boettcherisca nathani: Fan & Pape 1996: 248; Kano & Sugiyama 1983: 45; Kano et al. 1999: 135; Kurahashi & Kano 1984: 27; Kurahashi & Ohtaki 1989: 291; Kurahashi et al. 1991: 111; Kurahashi et al. 1995: 573; Nandi 1992b: 187; 2002: 205; Verves 1990: 542; 2001: 243. Sarcophaga nathani: Bänzinger & Pape 2004: 1677; Sugiyama & Kano 1984: 351; Sugiyama et al. 1988b: 360. Sarcophaga (Boettcherisca) nathani: Pape 1996: 310; Pape & Bänzinger 2000: 206; Shinonaga & Thinh 2003: 332. Boettcherisca chianshanensis Ma 1964: 58. Type locality: China: Liaoning, Chianshan. Holotype (♂) deposited in University of Liaoning, China.

Palaearctic region: China: Liaoning. Oriental region: China: Yunnan; India: Gujarat, Karikal, Maharashtra, Tamil Nadu, West Bengal; Indonesia: Java, Kalimantan, Sumatra; Malaysia: West Malaysia; Nepal; Pakistan; Philippines: Palawan; Thailand; Vietnam. Hemisynanthropic species; larvae are developed in decaying animal matter and human faeces, responsible for intestinal myiasis of man (Kurahashi et al. 1995). Adult flies feed on flowers of Sapria ram; known to be mechanical transmitters of agents of disease.

Boettcherisca nepalensis Kano & Sugiyama 1983 Boettcherisca nepalensis Kano & Sugiyama 1983: 45. Type locality: Nepal: Bara, Ratanpur. Holotype (♂) deposited in National Science Museum, Tokyo, Japan.


Boettcherisca nepalensis: Nandi 2002: 218; Verves 1990: 542. Sarcophaga birganjensis Sugiyama 1988 in: Sugiyama et al. 1988b: 357. Figures 1–3 1, Male genitalia of Boettcherisca peregrina (a, aedeagus, lateral view; b, distiphallus, ventral view; c, gonites, lateral view; d, epandrium, cercus & surstylus, lateral view); orig. 2, Male genitalia of Chrysosarcophaga superba (a, cercus & surstylus, lateral view; b, aedeagus, lateral view; c, distiphallus, ventral view); after Lopes & Kano 1978. 3, Male genitalia of Dasysceloctis congensis, lateral view; after Curran 1934.

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Unnecessary new name for Boettcherisca nepalensis Kano & Sugiyama 1983. Sarcophaga (Boettcherisca) birganjensis: Pape 1996: 309.

Oriental region: Nepal. Adult flies were collected in the garden of native house in jungle and attracted by human feces.

Boettcherisca peregrina (Robineau-Desvoidy 1830) (Fig. 1, a–d). Myophora peregrina Robineau-Desvoidy 1830: 356. Type locality: Australia: New South Wales: Port Jackson near Sydney. Holotype (♂) deposited in National Museum of Natural History, Paris, France; probably lost. Boettcherisca peregrina: Artamonov 1980a: 32; 1987: 179; Buei et al. 1978: 125; Cantrell 1981: 237; Early & Goff 1986: 520; Evenhuis 1985: 385; Fan 1992: 689; 2002: 86; Fan & Pape 1996: 248; Feng et al. 1990: 63; Ferrar et al. 1975: 10; Goff et al. 1989: 91; 1991: 537; Jiang 2002: 1445; Kamimura & Arakawa 1986: 163; Kani & Iwata 1981: 13; Kani et al. 1981: 173; Kano et al. 1967: 17; Kano & Sugiyama 1983: 47; Kano et al. 1999: 136; Kurahashi & Kano 1984:27; Kurahashi & Ohtaki 1989: 291; Lehrer 2003: 126; Lopes 1958: 29; 1961a: 71; 1961b: 422; Lopes et al. 1977: 565; Mihara et al. 1988: 131; Nandi 1992a: 36; 2002: 212; Park 1977: 253; Rohdendorf 1937: 270; 1963: 10; 1964: 81; Rudzinski & Kozanek 1991: 330; Sakurai 1979: 263; Smithers 1998: 27; Sun & Ren 1995: 50; Verves 1986a: 161; 1986b: 547; 1990: 542; 2001: 243; 2003: 10; 2007: 75; Verves & Khrokalo 2006: 123; 2009: 276, 291. Sarcophaga peregrina: Blackith & Blackith 1988: 301; Chigusa et al. 1994: 153; 2005a: 355; 2005b: 249; Hall & Bohart 1948: 127; Hanski 1981: 197; Hardy 1927: 453; 1932a: 45; Ho 1938: 115; Hori 1951: 3; 1952: 77; 1967: 60; James 1947: 55; Johnston & Hardy 1923: 121; Johnston & Tiegs 1922a: 87; 1922b: 182; 1922c: 177; Joshi 1973: 77; Kano 1950: 854; 1951: 225; 1957: 291; Kano et al. 1951: 115; Lopes 1939: 561; 1941: 55; Maeda 1960: 67; Moribayashi et al. 2001: 643; Mungomery 1947: 35; Otranto & Stewens 2002: 1347; Park 1962: 39; Quo 1952: 76; Salem 1946: 186; Sankaran & Syed 1972: 57; Séguy 1941: 131; Sherman et al. 2000: 58; Sugiyama et al. 1987: 75; 1988a: 290; 1988b: 361; Zumpt 1964: 61. Sarcophaga (Parasarcophaga) peregrina: Hardy 1943: 29. Sarcophaga (Prionophalla) peregrina: Reed 1974: 195. Sarcophaga (Boettcherisca) peregrina: Bänzinger & Pape 2004: 1677; Kano & Shinonaga 1994: 263; Lin & Chen 1999: 117; Pape 1996: 310; Shinonaga & Thinh 2003: 332; Yoneda et al. 1998: 51. Sarcophaga fuscicauda Böttcher 1912b: 169. Type locality: China: Taiwan. Holotype (♂) deposited in German Entomological Institute, BerlinDahlem, Germany. Sarcophaga fuscicauda: Bezzi 1928: 189; Böttcher 1913: 379; Bryan 1934: 417; Buxton 1929: 145; Eysell 1915: 2; 1926: 4; Greene 1925: 11; Hennig 1941: 185; Ho 1934: 31; 1936: 264; 1938: 119; Illingworth 1926: 263; Johnston & Tiegs 1922b: 182; Kang 1988a: 211; 1988b: 217; Malloch 1930: 483; Patton & Evans 1929: 484; Senior-White 1924: 252; 1927: 77; 1930: 74; Senior-White et al. 1940: 272. Sarcophaga hutsoni Parker 1923: 127. Type locality: Sri Lanka: Peradeniya. Holotype (♂) deposited in Natural History Museum, London, United Kingdom. Sarcophaga meriani Zumpt 1951: 182. Type locality: Seychelles. Holotype (♂) deposited in South African Institute for Medical Research, Johannesburg, South Africa. Sarcophaga meriani: Orian 1962: 1. Sarcophaga (Prionophalla) meriana: Reed 1974: 195; incorrect subsequent spelling of Sarcophaga meriani Zumpt 1951.

Palaearctic region: China: Anhui, Beijing, Gansu, Hebei, Heilongjiang, Henan, Hubei, Jangsu, Jilin,

308

Liaoning, Neimenggu, Ningxia, Shaanxi, Shandong, Shanghai, Shanxi, Sichuan, Tianjin, Xizang; Japan: Hashido Is., Hokkaido, Honshu, Kyushu, Shikoku, Tsushima Is.; North Korea; Russia: Far East (Southern Primorye); South Korea. Oriental region: Bangladesh; Bhutan; Cambodia; China: Fujian, Guangdong, Guangxi, Guizhou, Hainan, Hunan, Jiangxi, Taiwan, Yunnan, Zhejiang; India: Andaman Is., Assam, Bihar, Dadra and Nagar Haveli, Delhi, Himachal Pradesh, Karnataka, Kerala, Madhya Pradesh, Maharashtra, Manipur, Meghalaya, Mizoram, Nagaland, Nicobar Is., Orissa, Pondicherry, Sikkim, Tamil Nadu, Tripura, Uttar Pradesh, West Bengal; Indonesia: Java, Kalimantan, Riou Arch., Sulawesi; Japan: Ryukyu Is.: Okinawa (Yaeyama I.); Malaysia: Borneo (Sarawak, Sabah), West Malaysia (Perak); Laos; Myanmar; Nepal; Singapore; Sri Lanka; Thailand; Vietnam. Australasian/Oceanian region: Australia: New South Wales, Northern Territory, Queensland, South Australia, Western Australia; Bonin [= Ogasawara] Is.: Chihi Jima, Myianohama, Ogiura, Okumara, Omura, Sakaiura; Eastern Samoa; Fiji; French Polynesia: Society Is.; Guam; Hawaiian Is.: Hawaii, Kauai, Lanai, Maui, Molokai; Indonesia: Irian Jaya; Kiribati: Gilbert Is.; Marianas: Saipan I.; New Zealand: North I.; Niue I.; Norfolk I.; Papua New Guinea: Bismarck Arch. (New Britain), New Guinea; Volcano Is.: Iwo Jima I.; Western Samoa. Madagascan: Mauritius; Reunion; Seychelles: Aride I., Mahe I., North I., Poivre I., Silhouette I. The larvae bred from dead vertebrate and invertebrate (insects, snails) animals, garbage, animal dung and human feces. They are occasional parasites of living earthworms, and locust Chortoicetes terminifera (Walker 1870); facultative predators of lepidopteran pupae: nymphalid, Euploea corinna MacLeay 1827, and pyralid, Crocidolomia binotalis Zeller 1852; produced facultative aural, nasal, nasocomial, urogenital, and wound myiasis (sometimes larvae are developed in different cancerous growths) of men and tissue myiasis of mammals (Chigusa et al. 1994, 2005a, b; James 1947; Jiang 2002; Kamimura & Arakawa 1986; Kani & Iwata 1981; Kani et al. 1981; Nandi 2002; Patton & Evans 1929; Segal et al. 1968; Senior-White 1924; Senior-White et al. 1940; Sun & Ren 1995; Uni et al. 2006; Verves & Khrokalo 2006, 2009; Yoneda et al. 1998). Larvae develop during 4–7 days in summer and 7–11 days in winter, pupae – 8–11 days in summer and 1–15 days in winter in India. Larvae are the members of arthropod succession patterns in corpses and have an important forensic value. Flies feed on garbage, corpses, feces, flowers and fallen fruits. Adult flies extensively distributed in the eusynanthropic as well as the semisynanthropic and asynanthropic zones


at altitudes to 1300 m a. s. l.; known as a mechanical carrier of pathogenic microorganisms.

Boettcherisca septentrionalis Rohdendorf 1937 Boettcherisca (Notochaetomima) septentrionalis Rohdendorf 1937: 273. Type locality: Russia: Primorskiy Kray: environs of Vladivostok, Iman. Holotype (♂) deposited in Zoological Institute, St Petersburg, Russia. Boettcherisca septentrionalis: Artamonov 1980a: 32; 1987: 110; 1992: 48; Chigusa et al. 2006: 139; Fan 1992: 690; Fan & Pape 1996: 248; Kano et al. 1967: 19; Kano & Sugiyama 1983: 46; Kurahashi & Kano 1984: 27; Kurahashi & Suenaga 1996: 247; Lopes 1961a: 78; Mitsui 2002: 275; Nandi 2002: 216; Verves 1986a: 161; 1990: 542; Verves & Khrokalo 2006: 175. Sarcophaga septentrionalis: Kano 1951: 223; Shinonaga 2006: 265. Sarcophaga (Boettcherisca) septentrionalis: Pape 1996: 311.

Palaearctic region: China: Liaoning; Japan: Hachido Is., Honshu, Kyushu; Russia: Far East (South Kurily Is., South Primorye); South Korea. Oriental region: Bhutan. The time of larval development in corpses of small mammals and birds, rarely feces, is continued 5–6 days, and pupal development in soil – 11–38 days. A case of wound myiasis of Emberiza elegans Temminck 1836 nestling and hospital-acquired oral human myiasis due to larvae of this species were registered too (Chigusa et al. 2006). Adult flies prefer forests and bushes along river banks at altitudes to 1000 m a. s. l.

Boettcherisca talomoensis (Magpayo & Kano 1986), n. comb. Sarcophaga (Boettcherisca) talomoensis Magpayo & Kano 1986: 75. Type locality: Philippines: Mindanao: Davao, Mt Talomo. Holotype deposited in Tokyo Medical & Dental University, Tokyo, Japan. Sarcophaga (Boettcherisca) talomoensis: Pape 1996: 311.

Oriental region: Philippines: Mindanao.

Boettcherisca timorensis Kano & Shinonaga 1977 Boettcherisca timorensis Kano & Shinonaga 1977: 324. Type locality: Indonesia: Timor I., Kupang. Holotype (♂) deposited in Bogor Museum, Java, Indonesia. Boettcherisca timorensis: Kano & Sugiyama 1983: 45; Verves 1990: 542. Sarcophaga (Boettcherisca) timorensis: Pape 1996: 311.

Oriental region: Indonesia: Timor. The adults were in the forests using decayed meat and fishes at bait, and colonized in the laboratory.

Boettcherisca yuwanensis (Sugiyama 1990), n. comb.

Genus Chrysosarcophaga Townsend 1932 Townsend 1932: 441. Type species: Chrysosarcophaga superba Townsend 1932, by original designation. Chrysosarcophaga: Lopes & Kano 1978: 223; Townsend 1938: 19; Verves 1990: 542. Sarcophaga (Chrysosarcophaga): Pape 1996: 313.

A single species is distributed in Australasian/ Oceanian region.

Chrysosarcophaga superba Townsend 1932 (Fig. 2, a–c). Chrysosarcophaga superba Townsend 1932: 441. Type locality: Papua New Guinea: Bougainville I. Holotype (♀) deposited in American Museum of Natural History, New York, USA. Chrysosarcophaga superba: Curran 1936: 62; Lopes & Kano 1978: 223; Townsend 1938: 19; Verves 1990: 542. Sarcophaga (Chrysosarcophaga) superba: Pape 1996: 313.

Australasian/Oceanian region: Papua New Guinea: Bougainville I.; Solomon Is.: Guadalcanal I., Malaita I., Suta I. Flies were collected in mountains at altitudes to 1200 m a. s. l.

Genus Dasysceloctis Enderlein 1928 Enderlein 1928: 35. Type species: Dasysceloctis maculipennis Enderlein 1928; by original designation. Dasysceloctis: Dear 1980: 807; Lehrer 2002: 50; 2003: 164; Townsend 1931: 377; Zumpt 1972: 206. Mufindia Verves 1990: 541. Type species: Sarcophaga tanzaniae Zumpt 1972, by original designation. Mufindia: Lehrer 2002: 48. Sarcophaga (Dasysceloctis): Pape 1995:6; 1996: 314. Sarcophaga (Mufindia): Pape 1996: 364.

Three species are distributed in Afrotropical region.

Dasysceloctis congensis Curran 1934 (Fig. 3) Poecilometopa congensis Curran 1934: 23. Type locality: Democratic Republic of Congo: Burunga, 1°30’S, 29°18’E. Holotype (♂) deposited in American Museum of Natural History, New York, USA. Poecilometopa congensis: Zumpt 1953a: 16; Dasysceloctis congensis: Arnaud & Owen 1981: 185; Lehrer 2003: 164; Dasysceloctis maculipennis [misidentification: not Dasysceloctis maculipennis Enderlein 1928]: Dear 1980: 807 (in part); Townsend 1938: 54 (in part); Zumpt 1953a: 16; 1972: 206 (in part). Sarcophaga (Dasysceloctis) maculipennis: Pape 1995: 20 (in part); 1996: 314 (in part).

Afrotropical region: Democratic Republic of Congo; Kenya; Tanzania.

Dasysceloctis longanota Lehrer 2005

Sarcophaga yuwanensis Sugiyama 1990: 421. Type locality: Japan: Ryukyu Is., Amami-Oshima I., Kagoshima Prefecture, Mt Yuwan. Holotype (♂) deposited in the National Science Museum (Natural History), Tokyo, Japan. Sarcophaga (Boettcherisca) yuwanensis: Pape 1996: 311.

Dasysceloctis longanota Lehrer 2005: 14. Type locality: Kenya: Mt. Elgon, 10250 ft [= Mt. Longanot, 2700 m a. s. l.], 1°05”N, 34°40” E. Holotype (♂) deposited in Natural History Museum, London, United Kingdom.

Oriental region: Japan: Ryukyu Is. (Amami-Oshima I.).

Afrotropical region: Kenya.

309


Dasysceloctis maculipennis Enderlein 1928 Dasysceloctis maculipennis Enderlein 1928: 35. Type locality: Tanzania: Lake Nyasa, Langenburg. Lectotype (♂; designated by Zumpt 1972: 208) deposited in Museum fur Naturkunde der Humbolt Universitat, Berlin, Germany. Dasysceloctis maculipennis: Dear 1980: 807 (in part); Lehrer 2003: 167 (holotype examined); Townsend 1938: 54 (in part); Zumpt 1953a: 16; 1972: 206 (in part). Sarcophaga (Dasysceloctis) maculipennis: Pape 1995: 20 (in part); 1996: 314 (in part).

Poecilometopa maculipennis: Rohdendorf 1963: 13. Sarcophaga (Liopygia) tanzaniae Zumpt 1972: 174. Type locality: Tanzania: Mufindi. Holotype (♂) deposited in South African Institute for Medical Researches, Johannesburg, South Africa. Sarcophaga (Liopygia) tanzaniae: Dear 1980: 813. Mufindia tanzaniae: Verves 1990: 542. Sarcophaga (Mufindia) tanzaniae: Pape 1996: 364.

Afrotropical region: Democratic Republic of Congo; Tanzania.

Figure 4 Male genitalia of Fanzideia cygnocerca (a, 5th abdominal sternite, ventral view; b, aedeagus, lateral view; c, gonites, lateral view; d, cercus & surstylus, lateral view; e, cerci, dorsal view); orig.

310


Genus Fanzideia n. gen. Type species: Fanzideia cygnocerca n. sp.

Etymology. The generic name Fanzideia is referring to Prof. Fan Zide, in honor of his contribution on entomology. Diagnosis. Antennae elongate; 1st flagellomere about 4.5 times as long as pedicel; 4 pairs of posterior dc, 6 rows of hair-liked pre acr, and only one prescutellar pair of post acr developed; katepisternal setae 1+1+1; middle hollow of propleuron bare, 3rd costal section is subequal to 5th one in the length; vein R1 bare, costal spine absent; posterior surfaces of hind coxae haired; 3rd abdominal tergite without median marginal setae. ♂: 4th abdominal sternite without dense setae on posterior half, 5th abdominal sternite broad and shortened, without medial processes, its lateral lobes slender and elongate, at inner margins with 2–3 rows of spine-like setae. Cerci in profile straight and broad, vesica and lateral styli strongly sclerotized, medial process menbranous, enlarged; juxtal processes elongated and bifurcated.

Remarks. This genus belongs to subtribe Boettcheriscina by widened styli and a presence of 4 pairs of posterior dc. It differs from all other genera of this subtribe by such features: propleuron bare; 5th abdominal sternite shortened and wide; juxta without spines, with elongated and bifurcated lateral arms; vesica strongly sclerotized, lateral styli elongated, band-shaped, and distinctly widened at apex. A single species is distributed in Oriental region (south China).

Fanzideia cygnocerca n. sp. (Fig. 4, a–e). Etymology. This specific name is formed from the Greek words, cygn - swan and cerca - cercal plate, referring to male cercal plate swan-head shaped distally. Description. ♂. Body length 10.0–10.5 mm. Head. Eyes bare; frons about 2.8 times as wide as antenna; frontal vitta black, about 2.0 times as wide as fronto-orbital plate; both interfrontal and ocellar setae short, inner vertical seta well developed, fr 10 pairs, proclinate orb absent; fronto-orbital plate and parafacial with golden pruinosity; one row of parafacial setae presents; parafacial about 1.2–1.4 times as wide as antenna; antenna black; arista long plumose, the longest hairs about 1.8 times as wide as 1st flagellomere; vibrissal angle situated behind frontal angle in profile; genal height about 2/7 of eye height, gena with bluish gray pruinosity, genal hairs black; 3 rows of black postocular setae present; occiput covered by yellowish-white hairs; proboscis shortened, prementum about 3.0 times as long as broad, palpus black, slightly longer than prementum. Thorax. Black in ground color, with gray pruinosity, scutum with three longitudinal black vittae; acr 0+1, dc 4+4, ial 0+3, pra subequal to posterior notopleural seta in length; lateral margins of scutellum with black hairs; subapical and basal scutellar setae well developed; middle hollow of propleuron bare; katepisternal setae 1+1+1. Wings. Hyaline, veins fuscous, basicosta yellowish to white, dorsal and ventral surfaces of R4+5 from base to crossvein r-m with short hairs; cell r4+5 open; both calypters white to yellowish; halter brown, orange at base.

Legs. Wholly black; fore tibia with 2 short ad in basal part, and 1 pv near the middle; mid femur with a row of short and strong av, becoming pectinated apically, 2 rows of long fringe-like pv in basal half, those cilia in distal half short and pectinated; mid tibia with 1 ad, 2 pd and 1 p; hind femur with a complete row of av, shorter than its diameter, only with a short and thin setae row on posteroventral surface, without distinct pv; hind tibia with a row of 7 av in middle part, in preapical part with a distinct av, 2 ad, and 2 pd, with a row of long hairs at distal 3/5 of posteroventral surface, and with apical pv; tarsi slightly longer than tibiae, 4th tarsomere shortened, claws and pulvilli elongate, distinctly longer than 5th tarsomere. Abdomen. Black in ground color, elongate oviform in dorsal view, with chessboard shaped versicolor patches; tergite 3 without median marginal setae, tergite 4 with distinct median marginal bristles. Cercus broad in lateral view, its distal half slightly swan-head, cerci sharply shopped off at apex in dorsal view; surstylus slender; pregonite wide at base and with apical kneeshaped edge; basiphallus short; distiphallus as wide as long; paraphallus broad; ventral processes absent; harpes small, but well developed, stock-shaped; distal part of vesica bottle-plug shaped; juxtal filament broad, elongate and membranous, its lateral margin reversed; juxtal processes well sclerotized and bifurcated, inner margin of bifurcation with numerous fine hairs; lateral styli long and mid broad, distinctly widened at apex. ♀. Unknown. Holotype. ♂, China: Jianfengling, 1412 m a. s. l., Hainan Province, 21. V. 2004 (Chun-tian Zhang). Paratype. China: ♂, same data as holotype. Both type specimens (holotype and paratype) were deposited in collection of Institute of Entomology, Shenyang Normal University, China.

Genus Johnstonimyia Lopes 1959 Lopes 1959: 48. Type species: Sarcophaga kappa Johnston & Tiegs 1921, by original designation. Johnstonimyia: Kano & Lopes 1981a: 295; Verves 1990: 542. Sarcophaga (Johnstonimyia): Pape 1996: 338.

10 species are distributed in Oriental and Australasian/Oceanian regions.

Johnstonimyia bezzii Kano & Lopes 1981 (Fig. 5, a–d). Johnstonimyia bezzii Kano & Lopes 1981a: 295. Type locality: Vanuatu: Santo I. Holotype (♂) deposited in Museu National, Rio de Janeiro, Brazil. Johnstonimyia bezzii: Lopes & Kano 1979b: 659. Sarcophaga (Johnstonimyia) vanuatu Pape 1991: 215 [unnecessary new name for Johnstonimyia bezzii Kano & Lopes 1981], n. syn. Sarcophaga (Johnstonimyia) vanuatu: Pape 1996: 340.

Australasian/Oceanian region: Vanuatu: Espiritu Santo I., Santo I.

Johnstonimyia fatua Lopes 1967 Johnstonimyia fatua Lopes 1967: 147. Type locality: Papua New Guinea: Bismarck Arch.: Manus I., Lorengau. Holotype (♂) deposited in Zoological Museum, Copenhagen, Denmark. Sarcophaga (Johnstonimyia) fatua: Pape 1996: 339.

Australasian/Oceanian region: Papua New Guinea: Bismarck Archipelago (Manus I.).

311


312


Johnstonimyia gressitti Kano & Lopes 1981 Johnstonimyia gressitti Kano & Lopes 1981a: 297. Type locality: Papua New Guinea: NE New Guinea: Karimui, S of Goroka. Holotype (♂) deposited in P. B. Bishop Museum, Honolulu, Hawaii, USA. Johnstonimyia gressitti: Verves 1990: 542. Sarcophaga gorokaensis Sugiyama, Shinonaga & Kano 1988a: 288 [unnecessary new name for Johnstonimyia gressitti Kano & Lopes 1981], n. syn. Sarcophaga gorokaensis: Pape 1996: 339.

Australasian/Oceanian region: Papua New Guinea: New Guinea. Flies were collected at altitudes to 1000 m a. s. l.

Johnstonimyia kappa (Johnston & Tiegs 1921) Sarcophaga kappa Johnston & Tiegs 1921: 81. Type locality: Australia: Queensland: Brisbane. Holotype (♂) deposited in Queensland Museum, Brisbane, Australia. Sarcophaga kappa: Hardy 1934: 50; Johnston & Hardy 1923: 124; Johnston & Tiegs 1922b: 181; Sugiyama et al. 1988a: 285, 289. Sarcophaga (Johnstonimyia) kappa: Pape 1996: 339. Sarcophaga (Parasarcophaga) kappa: Hardy 1943: 28. Johnstonimyia kappa: Brown & Shipp 1978: 179; Kano & Lopes 1981a: 297; Lopes 1959: 50; Verves 1990: 542. Sarcophaga illingworthi Parker 1922: 6. Type locality: Australia: Queensland: Gordonvale near Cairns. Holotype (♂) deposited in American Museum of Natural History, New York, USA.

Australasian/Oceanian region: Australia: New South Wales, Northern Territory, Queensland, Western Australia; Papua New Guinea: New Guinea. Larvae are developed in carrion.

Johnstonimyia lincta Lopes 1959 Johnstonimyia lincta Lopes 1959: 52. Type locality: Australia: Queensland: Springsurre. Holotype (♂) deposited in Division of Entomology, Canberra, Australia. Johnstonimyia lincta: Kano & Lopes 1981a: 297; Verves 1990: 542. Sarcophaga (Johnstonimyia) lincta: Pape 1996: 339.

Australasian/Oceanian region: Australia: Queensland. Figures 5–13 5, Male genitalia of Johnstonimyia bezzii (a, cercus & surstylus, lateral view; b, gonites, lateral view; c, aedeagus, lateral view; d, distiphallus, ventral view); after Kano & Lopes 1981. 6, Male genitalia of Kramerea schuetzei (a, cercus & surstylus, lateral view; b, aedeagus, lateral view; c, distiphallus, ventral view); orig. 7, Male genitalia of Lioproctia (Burmanomyia) beesoni (a, cercus & surstylus, lateral view; b, ibid., dorsal view; c, gonites, lateral view; d, aedeagus, lateral view; e, distiphallus, ventral view); orig. 8, Male genitalia of Lioproctia (Coonoria) sumbaensis, lateral view; after Shinonaga 2004. 9, Male genitalia of Lioproctia (s. str.) imitatrix (a, genitalia, lateral view; b, aedeagus, lateral view; c, distiphallus, ventral view); after Lopes 1959. 10, Male genitalia of Lucyphalla nuzzacii, lateral view; orig. 11, Male genitalia of Latistyla czernyi, lateral view; orig. 12, Distiphallus of Nyikamyia barracloughiana, lateral view; orig. 13, Male genitalia of Phallosphaera gravelyi (a, 5th abdominal sternite, ventral view; b, cercus & surstylus, lateral view; c, distiphallus and gonites, lateral view); after Povolný & Verves 1997 and Verves & Khrokalo 2006.

Johnstonimyia lopesi Shinonaga & Kano 1990 Johnstonimyia lopesi Shinonaga & Kano 1990: 487. Type locality: Vanuatu: Efate, 10 km NW of Vila. Holotype (♂) deposited in National Science Museum, Tokyo, Japan. Sarcophaga (Johnstonimyia) hugoi Pape 1996: 339, n. syn. [unnecessary new name for Johnstonimyia lopesi Shinonaga & Kano 1990].

Australasian/Oceanian region: Vanuatu: Efate I.

Johnstonimyia multicolor (Johnston & Tiegs 1922) Sarcophaga multicolor Johnston & Tiegs 1922c: 187. Type locality: Papua New Guinea: Bismarck Archipelago, New Britain I., Rabaul. Holotype (♂) deposited in Australian Museum, Sidney, Australia. Sarcophaga multicolor: Hardy 1927: 456; Sugiyama et al. 1988a: 290. Sarcophaga (Lioproctia) multicolor: Pape 1996: 345. Johnstonimyia multicolor: Lopes 1959: 55; Verves 1990: 542.

Australasian/Oceanian region: Indonesia: Maluku (Aru Is.); Papua New Guinea: Bismarck Archipelago (New Britain I.), New Guinea.

Johnstonimyia notabilis (Kano & Lopes 1969) Burmanomyia notabilis Kano & Lopes 1969: 521. Type locality: Malaysia: Selangor: Ulu Langat. Holotype (♂) deposited in B. P. Bishop Museum, Honolulu, Hawaii, USA. Burmanomyia notabilis: Lopes et al. 1977: 567. Johnstonimyia notabilis: Kano & Lopes 1981a: 297; Verves 1990: 542. Lioproctia notabilis: Lopes & Kano 1979b: 658. Sarcophaga (Lioproctia) notabilis: Bänzinger & Pape 2004: 1677; Pape 1996: 345; Pape & Bänzinger 2000: 201.

Oriental region: Malaysia: West Malaysia (Selangor), Sarawak; Thailand: south part. Flies were collected in lowland evergreen and upland mixed deciduous forests at altitudes to 500 m a. s. l.; they are pollinators of flowers Bulbophyllum putidum (Teijsmann & Binnendijk 1862) J. J. Smith 1912 and Sapria ram. Larviposition was discovered on carrion.

Johnstonimyia paineiana (Baranov 1934), n. comb. Lioproctia paineiana Baranov 1934: 184. Type locality: Solomon Is.: Guadalcanal I., Tenaru. Holotype (♂) deposited in Natural History Museum, London, United Kingdom. Lioproctia paineiana: Sabrosky & Crosskey 1970: 429. Sarcophaga (Lioproctia) paineiana: Pape 1996: 345.

Australasian/Oceanian Guadalcanal I.

region:

Solomon

Is.:

Johnstonimyia taiwanensis (Kano & Lopes 1969). Burmanomyia taiwanensis Kano & Lopes 1969: 522. Type locality: China: Taiwan: Pen-ch’i-hu [= Fenchihu], Chia-i-Hsien. Holotype (♂) deposited in Tokyo Medical & Dental University, Japan. Burmanomyia taiwanensis: Lopes et al. 1977: 566. Johnstonimyia taiwanensis Kano & Lopes 1981a: 297; Verves 1990: 542.

313


Lioproctia taiwanensis: Fan & Pape 1996: 251. Sarcophaga (Lioproctia) taiwanensis: Lin & Chen 1999: 117; Pape 1996: 345; Sugiyama et al. 1987: 77.

Oriental region: China: Taiwan.

Genus Kramerea Rohdendorf 1937 Rohdendorf 1937: 274. Type species: Sarcophaga schuetzei Kramer 1909, by original designation. Kramerea: Fan & Pape 1996: 251; Verves 1986a: 162; Verves 1990: 542; Verves & Khrokalo 2006: 175. Sarcophaga (Kramerea): Pape 1996: 341.

Kramerea schuetzei (Kramer 1909) (Fig. 6, a–c) Sarcophaga schuetzei Kramer 1909: 83. Type locality: Germany: Oberlausitz, Koenigsholz. Lectotype (♂; designated by Povolný 1988: 13) deposited in Staatliche Museum fur Naturkunde, Görlitz, Germany. Sarcophaga schuetzei: Böttcher 1912a: 733; Hori 1953: 83; Séguy 1941: 143; Shinonaga 2006: 265; Sugiyama et al. 1987: 76. Sarcophaga (Kramerea) schuetzei: Lin & Chen 1999: 117; Pape 1996: 341. Kramerea schuetzei: Artamonov 1980a: 32; 1987: 111; 1992: 52; Boldaruev 1952: 56; Egorov 1962: 294; Fan 1992: 665; Fan & Pape 1996: 251; Kano et al. 1967: 34; Lehrer & Dobrivojević 1970: 95; Mitsui 2002: 275; Novotný et al. 1998; Park 1977: 255; Rohdendorf 1937: 275; Rohdendorf & Verves 1978: 257; Verves 1986a: 162; 1998: 54; Tereshkin & Lobodenko 1997; Verves & Khrokalo 2006: 176; Zinovyev 1962: 30. Thyrsocnema schuetzei: Enderlein 1928: 43.

Palaearctic region: Austria; Azerbaijan; Bulgaria; Byelorussia; China: Beijing, Gansu, Heilongjiang, Henan, Jilin, Liaoning, Neimenggu, Shaanxi, Shanxi; Croatia; Czech Republic: Bohemia, Moravia; France; Germany; Hungary; Japan: Hokkaido, Honshu, Kyushu, Shikoku; Kazakhstan; Macedonia; Moldova; Mongolia: Central aimak, East aimak, Uvs Nuur aimak; Netherlands; North Korea; Poland; Russia: European part: Bashkortostan, Belgorod, Kaliningrad, Leningrad, Lypetzk, Moscow, Voroniezh regions, West Siberia: Altay, Tuva regions, East Siberia: Buryatia, Chita regions, Far East: Amur, Khabarovsk, S Kurily Is., Primorye, Sakhalin regions; Serbia; Slovakia; South Korea; Switzerland; Ukraine: Cherkasy, Chernigiv, Crimea, Ivano-Frankivsk, Kharkiv, Kherson, Khmelnytskiy, Kyiv, Zakarpattya, Zhytomyr regions. Oriental region: China: Taiwan. Larvae are developed in corpses of different invertebrate (snails, insects etc) and small vertebrate animals (mice, birds, fishes), decomposing meat, and known as facultative predators of lepidopteran pupae: Aporia crataegi L. 1758, Arctia sp., Cosmotriche potatoria (L. 1758), Dasychira albodentata (Bremer 1864), Dendrolimus pini (L. 1767), D. segregatus (Butler 1877), D. sibiricus (Tschetverikov 1908), D. spectabilis (Butler 1877), Dictyoploca japonica (Moore 1872), Lymantria dispar (L. 1758), L. monacha (L. 1758), Orgya antiqua (L. 1758), Pygaera anastomosis (L. 1758), Selenephera

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lunigera (Esper 1784). The time of larval development is 8–12 days, pupal development in soil – 10–18 days. Adult flies feed on garbage, corpses, feces, sweat of man, mucous secrets from mouth and nose of hoof animals, at haemolymph of wounded insects, destroyed fruits, human food and flowers. They prefer borders of forests, bushes, meadows, parks, gardens, and settlements.

Genus Latistyla Strukan 1970, n. stat. Strukan 1970: 96, as subgenus of Parasarcophaga Johnston & Tiegs 1921. Type species: Sarcophaga czernyi Böttcher 1912, by original designation & monotypy. Macabiella Lehrer 1994a: 15, n. syn. Type species: Parasarcophaga paularnaudi Lehrer 1981, by original designation. Sarcophaga (Macabiella): Pape 1996: 360.

Two species are distributed in Palaearctic (East Mediterranean area) region.

Latistyla czernyi (Böttcher 1912), n. comb. (Fig. 11) Sarcophaga czernyi Böttcher 1912a: 731. Type locality: Croatia: Dalmatia, Dubrovnik [= Raguza]. Holotype (♂) deposited in German Entomological Institute, Berlin-Dahlem, Germany. Sarcophaga czernyi: Séguy 1941: 96. Sarcophaga (Macabiella) czernyi: Pape 1996: 360. Thyrsocnema czernyi: Enderlein 1928: 43. Parasarcophaga (Latistyla) czernyi: Strukan 1970: 96. Parasarcophaga (Rosellea) czernyi: Verves 1986a: 172. Rosellea czernyi: Verves 1990: 542.

Palaearctic region: Croatia; Greece.

Latistyla paularnaudi (Lehrer 1994), n. comb. Parasarcophaga paularnaudi Lehrer 1981: 185. Type locality: Lebanon: Bterran, El Coura, 70 miles SE of Beirut. Holotype (♂) deposited in Department of Entomology, Californian Academy of Sciences, San Francisco, USA. Parasarcophaga (Rosellea) paularnaudi: Verves 1986a: 172. Macabiella paularnaudi: Lehrer 1994a: 15; 1998: 43; 2006b: 18. Rosellea paularnaudi: Verves 1990: 542.

Palaearctic region: Israel; Lebanon.

Genus Lioproctia Enderlein 1928 Enderlein 1928: 26. Type species: Lioproctia aurifrons sensu Enderlein 1928, by original designation [= Lioproctia enderleini Kano & Lopes 1970], misidentification: not Sarcophaga aurifrons Doleshall 1858. Lioproctia: Fan & Pape 1996: 251; Kano & Lopes 1970: 313; Lopes et al. 1977: 566; Nandi 2002: 197; Pape 1995: 9; Verves 1986a: 162; 1990: 541. Sarcophaga (Lioproctia): Pape 1996: 343.

16 species are distributed in Palaearctic, Oriental and Australasian/Oceanian regions.

Subgenus Burmanomyia Fan 1964 Fan 1964: 305, 316, as genus. Type species: Sarcophaga beesoni Senior-White 1924, by monotypy.


Two species are distributed in Palaearctic, Oriental and Australasian/Oceanian regions.

Lioproctia (Burmanomyia) alcicornis (Hardy 1932b) Sarcophaga alcicornis Hardy 1932b: 275. Type locality: Australia: Queensland, Brisbane. Holotype (♂) deposited in University of Queensland, Brisbane, Australia. Sarcophaga alcicornis: Hardy 1934: 50; Lopes 1939: 560; Sugiyama et al. 1988a: 285. Sarcophaga (Chrysosarcophaga) alcicornis: Hardy 1943: 26. Sarcophaga (Lioproctia) alcicornis: Pape 1996: 344. Tricholioproctia alcicornis: Brown & Shipp 1978: 179; Lopes 1955: 249. Lioproctia alcicornis: Lopes & Kano 1979b: 658; Lioproctia (Burmanomyia) alcicornis: Verves 1990: 542.

etc. Description of ♂ Burmanomyia parvatia is based at drawings of ♂ genitalia of Lioproctia beesoni from Nandi 2002. The differences in drawings are very petty and reflected the different styles of painters; they can not be used as reason for designation of two new species.

Subgenus Coonoria Fan 1964 Fan 1964: 305, 316, as subgenus of Burmanomyia. Type species: Sarcophaga pattoni Senior-White 1924, by original designation.

9 species are distributed in Oriental and Australasian/ Oceanian regions.

Lioproctia (Coonoria) aureolata (Pape & Kurahachi 2000), n. comb.

Australasian/Oceanian region: Australia: New South Wales, Queensland; Indonesia: Irian Jaya; Papua New Guinea: New Guinea. Larvae bred in corn, dead grubs, and scrub. Adult flies were collected on altitudes to 1000 m a. s. l.

Sarcophaga (Lioproctia) aureolata Pape & Kurahachi 2000: 519. Type locality: Indonesia: Timor: Nusa Tenggare Timor, Soe. Holotype (♂) deposited in Puslitbang Biologi LIPI, Cibining, Indonesia. Sarcophaga aureolata: Pape & Kurahachi 2004: 160.

Lioproctia (Burmanomyia) beesoni (Senior-White 1924) (Fig. 7, a–e).

Oriental region: Indonesia: Timor. Flies were collected in forest at latitude 800 m a. s. l.

Sarcophaga beesoni Senior-White 1924: 243. Type locality: Myanmar [as “Burma”]: Mohnyin. Holotype (♂) deposited in Natural History Museum, London, United Kingdom. Sarcophaga beesoni: Bänzinger & Pape 2004: 1677; Kano & Shinonaga 1994: 263; Lin & Chen 1999: 116; Senior-White et al. 1940: 248; Sugiyama et al. 1987: 65. Sarcophaga (Lioproctia) beesoni: Pape 1996: 344. Burmanomyia beesoni: Fan 1964: 305; 1992: 663; Kano et al. 1967: 30; Weng & Zhou 1995: 147. Johnsonimyia beesoni: Kano & Shinonaga 1965: 16, pl. 73. Lioproctia beesoni: Fan & Pape 1996: 251; Kano et al. 1999: 133; Lopes et al. 1977: 567; Nandi 2002: 232; Verves 1986a: 162. Lioproctia (Burmanomyia) beesoni: Verves 1990: 542; 2001: 243. Burmanomyia parvatia Lehrer 2008: 12, n. syn. Type locality not given, type material absent. Burmanomyia guanyina Lehrer & Wei 2010a: 1, n. syn. Type locality: China: Guizhou, Longli forest farm, Longli, 1000 m a. s. l. Holotype (♂) deposited in Center for Disease Prevention and Control, Guishou, China.

Palaearctic region: China: Anhui, Henan, Hubei, Hunan, Jangsu, Shanghai, Sichuan; Japan: Kyushu. Oriental region: China: Fujian, Guangdong, Guangxi, Guizhou, Hainan, Hunan, Jiangxi, Taiwan, Yunnan, Zhejiang; India: West Bengal; Myanmar; Nepal; Thailand; Vietnam. Larvae bred in pupae of moth Dendrolimus punctatus (Walker 1855). Adult flies were collected on leaves along mountains path and tops of hills. Taxonomical notes: The original descriotion and drawings of ♂ genitalia of male Burmanomyia guanyina are very detailed and practically not differentiated from similar descriptions and drawings of ♂ genitalia of Lioproctia beesoni by Fan 1992; Kano et al. 1967; Nandi 2002; Rohdendorf 1966; Senior-White 1924

Lioproctia (Coonoria) kurahashii (Shinonaga & Tumrasvin 1979), n. comb. Phallosphaera kurahashii Shinonaga & Tumrasvin 1979: 141. Type locality: Thailand: Kanchana Buri: Sai Yok. Holotype (♂) deposited in National Science Museum, Tokyo, Japan. Phallosphaera kurahashii: Kano & Lopes 1981b: 575. Sarcophaga kurahashii: Bänzinger & Pape 2004: 1677. Sarcophaga (Lioproctia) kurahashii: Shinonaga & Thinh 2003: 335. Sarcophaga (Phallosphaera) kurahashii: Pape 1996: 378.

Oriental region: Thailand: south-east part. Flies were collected in upland evergreen forests.

Lioproctia (Coonoria) lothianensis Sinha & Nandi 2002 Lioproctia lothianensis Sinha & Nandi 2002: 39. Type locality: India: West Bengal: Sundarbans Biosphere Reserve. Holotype (♂) deposited in Department of Zolology, Presidency College, Kolkata, India.

Oriental region: India: West Bengal.

Lioproctia (Coonoria) pattoni (Senior-White 1924) Sarcophaga pattoni Senior-White 1924: 242. Type locality: India: Tamil Nadu: Nilgiri Hills, Coonor. Holotype (♂) deposited in Natural History Museum, London, United Kingdom. Sarcophaga pattoni: Kano & Shinonaga 1994: 263; Lin & Chen 1999: 117; Senior-White et al. 1940: 248; Shinonaga 2004: 283; Sugiyama et al. 1987: 75; 1988b: 361. Sarcophaga (Lioproctia) pattoni: Bänzinger & Pape 2004: 1677; Pape 1996: 345; Pape & Bänzinger 2000: 201. Burmanomyia (Coonoria) pattoni: Fan 1964: 305; 1992: 663. Tricholioproctia pattoni: Lopes 1955: 274.

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Lioproctia pattoni: Fan & Pape 1996: 251; Kano et al. 1999: 133; Lopes & Kano 1979b: 658; Lopes et al. 1977: 567. Lioproctia (Coonoria) pattoni: Nandi 2002: 239; Verves 1990: 542; 2001: 243. Sarcophaga pilipleuris Salem 1946: 191. Type locality: Indonesia: Java: Jakarta [as “Batavia”]. Holotype (♂) deposited in Natural History Museum, London, United Kingdom. Liproctia kunlunea Lehrer 2008: 16, n. syn. Type locality not given, type material absent.

Palaearctic region: China: Henan, Hubei, Sichuan. Oriental region: China: Taiwan, Yunnan; India: Mizoram, Nagaland, Sikkim, Tamil Nadu, Tripura; Indonesia: Java, Sumatra; Malaysia: West Malaysia; Nepal; Philippines; Singapore; Thailand; Vietnam. Larvae bred in dead mammals (rabbits etc), occasionally in human feces. Flies were collected at altitudes to 1600 m a. s. l., in most types of forests and synanthropic habitats. Adults feed on decomposed animal matters and flowers of Sapria ram. Taxonomical notes: Description of ♂ Burmanomyia parvatia is based at drawings of ♂ genitalia of Lioproctia pattoni from Fan 1965, 1992 and Nandi 2002. The differences in drawings are very petty and reflected the different styles of painters; they can not be used as reason for designation of a new species.

Lioproctia (Coonoria) saprianovae (Pape & Bänzinger 2000), n. comb. Sarcophaga (Lioproctia) saprianovae Pape & Bänzinger 2000: 202. Type locality: Thailand: Ranong Province: Phya Nak Falls, Khlong Naka Wildlife Sanctuary. Holotype (♂) deposited in Swedish Museum of Natural History, Stockholm, Sweden. Sarcophaga (Lioproctia) saprianovae: Bänzinger & Pape 2004: 1677.

Oriental region: Thailand: central-west & south parts. Flies were collected in lowland evergreen and upland mixed deciduous forests at altitudes to 810 m a. s. l. Adults feed on flowers of Sapria ram. Female larviposition on carrion was registered.

Lioproctia (Coonoria) serracudo (Pape & Kurahachi 2004), n. comb. Sarcophaga (Lioproctia) serracudo Pape & Kurahachi 2004: 174. Type locality: Indonesia: Timor I. Holotype (♂) deposited in Indonesian Institute of Sciences, Gibinong, Indonesia. Sarcophaga (Lioproctia) serracuda: Shinonaga 2004: 287; incorrect subsequent spelling of serracudo.

Oriental region: Indonesia: Timor I.

Lioproctia (Coonoria) sumbaensis (Shinonaga 2004), n. comb. (Fig. 8) Sarcophaga (Lioproctia) sumbaensis Shinonaga 2004: 288. Type locality: Indonesia: Sumba I.: Praikalittu, Lewapaku, Sumba Timur. Holotype (♂) deposited in National Science Museum, Tokyo, Japan. Sarcophaga (Lioproctia) sumbawensis: Shinonaga 2004: 287; incorrect original spelling of sumbaensis.

Oriental region: Indonesia: Sumba I.

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Lioproctia (Coonoria) sundaensis (Shinonaga 2004), n. comb. Sarcophaga (Parasarcophaga) sundaensis Shinonaga 2004: 291. Type locality: Indonesia: Timor I.: Kupang. Holotype (♂) deposited in National Science Museum, Tokyo, Japan.

Oriental region: Indonesia: Lombok I., Timor I.

Lioproctia (Coonoria) vietnamensis (Shinonaga & Thinh 2003), n. comb. Sarcophaga (Lioproctia) vietnamensis Shinonaga & Thinh 2003: 334. Type locality: Vietnam: Thua Thien Hue Province: Bach Ma. Holotype (♂) deposited in National Science Museum, Tokyo, Japan.

Oriental region: Vietnam. Flies were collected in mountains at altitudes 1000–1200 m a. s. l.

Subgenus Lioproctia (s. str.) 5 species are distributed in Australasian/Oceanian region.

Lioproctia (s. str.) aurescens (Lopes 1967). Johnstonimyia aurescens Lopes 1967: 148. Type locality: Papua New Guinea: Bismarck Arch.: New Britain I., Yalom. Holotype (♂) deposited in Zoological Museum, Copenhagen, Denmark. Lioproctia aurescens: Kano & Lopes 1970: 313; Verves 1990: 542. Sarcophaga (Lioproctia) aurescens: Pape 1996: 344. Sarcophaga compta Walker 1859: 102 [junior primary homonym of Sarcophaga compta Wiedemann 1830]. Type locality: Indonesia: Aru Is. Holotype (♀) deposited in Natural History Museum, London, United Kingdom. Lioproctia compta: Lopes & Kano 1979a: 306.

Australasian/Oceanian region: Indonesia: Moluccas: Aru Is.; Papua New Guinea: Bismarck Arch. (New Britain I.). Flies were collected at altitude about 1000 m a. s. l.

Lioproctia (s. str.) enderleini (Kano & Lopes 1970) Johnstonimyia enderleini Kano & Lopes 1970: 313 [new name for Lioproctia aurifrons sensu Enderlein 1928]. Type locality: Indonesia: Maluku: Ambon I. Holotype not designated. Lioproctia (s. str.) enderleini: Verves 1990: 542. Lioproctia aurifrons sensu Enderlein 1928: 26 [misidentification: not Sarcophaga aurifrons Doleshall 1858]; Lopes et al. 1977: 567. Sarcophaga (Lioproctia) ambon Pape 1995: 5 [unnecessary new name for Lioproctia enderleini Kano & Lopes 1970]. Sarcophaga (Lioproctia) ambon: Pape 1996: 344.

Australasian/Oceanian region: Indonesia: Maluku Arch.: Ambon I.

Lioproctia (s. str.) imitatrix (Lopes 1959) (Fig. 9, a–c) Johnstonimyia imitatrix Lopes 1959: 49, 56. Type locality: Australia: Queensland: Cairns. Holotype (♂) deposited in Instituto Oswaldo Cruz, Rio de Janeiro, Brazil. Lioproctia (s. str.) imitatrix: Verves 1990: 542.


Sarcophaga spinifera: Lopes 1939: 562 [misidentification: not Sarcophaga spinifera Hardy 1932]. Sarcophaga (Lioproctia) imita Pape 1996: 58, 344, n. syn. [unnesessary new name for Lioproctia imitatrix Lopes 1959] .

Australasian/Oceanian region: Australia: Queensland; Indonesia: Irian Jaya; Papua New Guinea: Bismarck Arch., New Guinea.

Lioproctia (s. str.) spinifera (Hardy 1932) Sarcophaga spinifera Hardy 1932b: 277. Type locality: ♂: Australia: Queensland: Brisbane. Holotype (♂) deposited in Queensland University, Brisbane, Australia. Sarcophaga spinifera: Hardy 1934: 50; 1943: 31. Lioproctia (s. str.) spinifera: Verves 1990: 542. Johnstonimyia spinifera: Lopes 1959: 49, 57. Sarcophaga (Lioproctia) spinifera: Pape 1996: 345.

Australasian/Oceanian region: Australia: Queensland.

Lioproctia (s. str.) torvida (Lopes 1959) Johnstonimyia torvida Lopes 1959: 49, 54. Type locality: Australia: Western Australia: S estuary of Fortescue River, Mardie. Holotype (♂) deposited in Division of Entomology, Canberra, Australia. Johnstonimyia torvida: Brown & Shipp 1978: 179. Lioproctia (s. str.) torvida: Verves 1990: 542. Sarcophaga (Lioproctia) torvida: Pape 1996: 345.

Australasian/Oceanian region: Australia: Western Australia.

Genus Lucyphalla Lehrer 2004 Lehrer 2004: 116. Type species: Lucyphalla nuzzacii Lehrer 2004, by original designation.

A single species is distributed in Afrotropical region.

Lucyphalla nuzzacii Lehrer 2004 (Fig. 10) Lucyphalla nuzzacii Lehrer 2004: 116. Type locality: Benin: Porto Novo. Holotype (♂) deposited in Laboratory of Zoology, Tel Aviv University, Israel.

Afrotropical region: Benin.

Genus Nyikamyia Lehrer 1994 Lehrer 1994b: 213. Type species: Nyikamyia barracloughiana Lehrer 1994, by original designation. Nyikamyia: Lehrer 2003: 36, 343. Sarcophaga (Nyikamyia): Pape 1996: 371.


Nyikamyia barracloughiana Lehrer 1994b (Fig. 12) Nyikamyia barracloughiana Lehrer 1994b: 213. Type locality: Malawi: Nyila National Park, Chilinda Camp Riverine scrub. Holotype (♂) deposited in Natal Museum, Pietermaritzburg, South Africa. Nyikamyia barracloughiana: Lehrer 2003: 343. Sarcophaga (Nyikamyia) barracloughiana: Pape 1996: 371.

Afrotropical region: Malawi.

Genus Phallosphaera Rohdendorf 1938 Rohdendorf 1938: 107. Type species: Phallosphaera konakovi Rohdendorf 1938, by original designation. Phallosphaera: Fan 1964: 305, 316; 1992: 665; Fan & Pape 1996: 254; Kano et al. 1967: 91; Kano & Lopes 1981b: 575; Lopes et al. 1977: 572; Nandi 2002: 198, 351; Rohdendorf 1965: 685, 694; Verves 1986a: 173; 1990: 541; Verves & Khrokalo 2006: 77, 176. Yunnanomyia Fan 1964: 305, 316, as subgenus of Phallosphaera Rohdendorf 1938. Type species: Sarcophaga gravelyi Senior-White 1924, by original designation. Sarcophaga (Phallosphaera): Pape 1996: 378.

Four species are distributed in Palaearctic (east hart) and Oriental regions.

Phallosphaera amica Ma 1964 Ma 1964: 62. Type locality: China: Liaoning: Ch’ienshan. Holotype (♂) deposited in University of Liaoning, China. Phallosphaera amica: Fan 1992: 665; Fan & Pape 1996: 254; Kano & Lopes 1981b: 577; Verves 1986a: 174; 1990: 542. Sarcophaga (Phallosphaera) amica: Pape 1996: 378.

Palaearctic region: China: Liaoning.

Phallosphaera gravelyi (Senior-White 1924) (Fig. 13, a–c) Sarcophaga gravelyi Senior-White 1924: 229. Type locality: India: Tamil Nady: Nilgiri Hills, Kallar (as “Kulla”). Holotype (♂) deposited in Natural History Museum, London, United Kingdom. Sarcophaga gravelyi: Bänzinger & Pape 2004: 1677; Kano & Shinonaga 1994: 263; Lin & Chen 1999: 117; Senior-White et al. 1940: 224; Shah et al. 2006: 222; Sugiyama et al. 1987: 70; 1988b: 356, 359. Sarcophaga (Phallosphaera) gravelyi: Pape 1996: 378; Shinonaga 2004: 283; Shinonaga & Thinh 2003: 332. Phallosphaera gravelyi: Artamonov 1978: 54; 1980a: 32; Fan 1992: 664; Fan & Pape 1996: 254; Hsien 1958: 79; Kano et al. 1967: 91; 1999: 134; Kano & Lopes 1981b: 575; Lopes et al. 1977: 573; Mitsui 2002: 275; Nandi 2002: 352; Park 1977: 267; Verves 1986a: 174; 1990: 542; 2001: 243; Verves & Khrokalo 2006: 177. Phallosphaera (Yunnanomyia) gravelyi: Fan 1964: 305, 316. Sarcophaga longicornis Böttcher 1912b: 166 [junior primary homonym of Sarcophaga longicornis Macquart 1843]. Type locality: China: Taiwan: Dafulin (as “Taihorinsho”). Holotype (♂) deposited in German Entomological Institute, Berlin-Dahlem, Germany. Sarcophaga formosana Senior-White 1924: 243 [new name for Sarcophaga longicornis Böttcher 1912]. Sarcophaga formosana: Baranov 1934: 41; Senior-White et al. 1940: 250. Sarcophaga kinoshitai Hori 1954: 45. Type locality: Japan: Honshu: Ishikawa Prefecture, Mt Shiritaka. Holotype (♂) deposited in Museum of Kanazawa University, Japan. Phallosphaera huangdinia Lehrer 2008: 17, n. syn. Type locality not given, type material absent. Phallosphaera jimmuana Lehrer 2008: 18, n. syn. Type locality not given, type material absent.

Palaearctic region: China: Hubei, Liaoning, Shaanxi, Sichuan; Japan: Honshu, Kyushu; Korea; Russia: Kurily Is. (Kunashir I.). Oriental region: China: Fujian, Guangdong, Guizhou, Hunan, Taiwan, Yunnan, Zhejiang; India: Andra Pradesh, Tamil Nadu; Indonesia: Java; Japan: Ryukyu Is.; Nepal; Pakistan: Punjab; Thailand; Vietnam.

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Larvae are developed in dead invertebrate and small vertebrate animals, sometimes as predators of pupae of saturniid butterfly Samia cynthia (Drury 1773). Adult flies prefer borders of forest and bush inhabitates at altitudes to 1300 m a. s. l.; feed on corpses, feces, decomposed fruits and aphid excreta. Taxonomical notes: Description of ♂ Phallosphaera huangdinia is based at drawings of ♂ genitalia of Phallosphaera gravelyi from Fan 1965, 1992, and Verves & Khrokalo 2006; description of ♂ Phallosphaera jimmuana is based at drawings of ♂ genitalia of Phallosphaera gravelyi from Kano et al. 1967. The differences in drawings are very petty and reflected the different styles of painters; they can not used as reason for designation of both new species.

Phallosphaera konakovi Rohdendorf 1938 Phallosphaera konakovi Rohdendorf 1938: 107. Type locality: Russia: Primorye: Sikhote-Alin’ State Reservation. Holotype (♂) deposited in Zoological Institute, Russian Academy of Science, St Petersburg, Russia. Phallosphaera konakovi: Artamonov 1980a: 32; 1985: 19; 1987: 116; 1988: 31; Fan 1992: 664; Fan & Pape 1996: 254; Kano et al. 1967: 93; Kano & Lopes 1981b: 577; Lopes et al. 1977: 573; Verves 1986a: 174; 1990: 542; Verves & Khrokalo 2006: 177. Sarcophaga (Phallosphaera) konakovi: Pape 1996: 378; Shinonaga & Thinh 2003: 332. Sarcophaga konakovi: Bänzinger & Pape 2004: 1677; Shinonaga 2000: 476; 2001: 359. Sarcophaga shiroganensis Kano & Okazaki 1956: 73. Type locality: Japan: Honshu: Tokyo, Hirayama Park. Holotype (♂) deposited in Tokyo Medical and Dental University, Tokyo, Japan.

Palaearctic region: China: Heilongjiang, Jilin, Liaoning, Neimenggu, Shaanxi, Sichuan; Japan: Hokkaido, Honshu, Kyushu; North Korea; Russia: Far East: Kurily Is., Southern Primorye. Oriental region: China: Yunnan; Malaysia: Sabah, West Malaysia; Thailand; Vietnam. Larvae are developed in dead fishes and river crustaceans, sometimes as predators of butterfly pupae (Lymantria dispar); larval cannibalism was registered Figures 14–20 14, Male genitalia of Robackina triplasia (a, aedeagus and gonites, lateral view; b, distiphallus, ventral view); after Lopes 1975. 15, Male genitalia of Rosellea aratrix (a, 5th abdominal sternite, ventral view; b, cercus & surstylus, lateral view; c, aedeagus, lateral view; d, distiphallus, ventral view; e, gonites, lateral view); orig. 16, Male genitalia of Sabiella freyi (a, genitalia, lateral view; b, cerci, dorsal view; c, distiphallus, anterior view); after Zumpt 1972. 17, Male genitalia of Saputaramyia saputaraensis (a, cercus & surstylus, lateral view; b, ibid., dorsal view; c, 5th abdominal sternite, ventral view; d, aedeagus and gonites, lateral view; e, distiphallus, ventral view); after Nandi 1992. 18, Male genitalia of Sclerophalla santosdiasi (a, genitalia, lateral view; b, cerci, dorsal view; c, distiphallus, anterior view); after Zumpt 1972. 19, Male genitalia of Takanoa rugosa (a, hypandrium, cercus, surstylus & gonites, lateral view; b, aedeagus, lateral view; c, distiphallus, ventral view); orig. 20, Male genitalia of Ziminisca semenovi (a, cercus & surstylus, lateral view; b, aedeagus & gonites, lateral view); orig.

too. Adult flies prefer forests, parks and bushes along river shores at altitudes to 1000 a. s. l.; feed on corpses, feces, decomposed fruits and aphid excreta.

Phallosphaera metzgeri Kano & Shinonaga 1964 Phallosphaera metzgeri Kano & Shinonaga 1964: 213. Type locality: Japan: Hokkaido: Sapporo, Maruyama Park. Holotype (♂) deposited in Tokyo Medical and Dental University, Tokyo, Japan. Phallosphaera metzgeri: Kano et al. 1967: 94; Kano & Lopes 1981b: 577; Verves 1986a: 174; 1990: 542; Verves & Khrokalo 2006: 177. Sarcophaga (Phallosphaera) metzgeri: Pape 1996: 378.

Palaearctic region: Japan: Hokkaido, Honshu; South Korea. Mountainous species.

Genus Robackina Lopes 1975 Lopes 1975: 159. Type species: Sarcophaga triplasia Wulp 1896, by original designation. Robackina: Pape 1996: 366 [as synonym of Neobellieria Blanchard 1939]; Verves 1990: 542.

Two species are distributed in Nearctic and Neotropical regions.

Robackina sternalis Reinhard 1939 Emblemasoma sternalis Reinhard 1939: 62. Type locality: USA: Texas: Donna. Holotype (♂) deposited in Canadian National Collection of insects and related arthropods, Ottawa, Canada. Archimimus sternalis: Downes 1965: 942. Robackina sternalis: Lopes 1988: 921; Lopes & Tibana 1988: 325. Sarcophaga (Neobellieria) sternalis: Pape 1996: 367.

Nearctic region: USA: Texas.

Robackina triplasia (Wulp 1896) (Fig. 14, a, b) Sarcophaga triplasia Wulp 1896: 269. Type locality: Mexico: Guerrero: Amula. Holotype (♂) deposited in Natural History Museum, London, United Kingdom. Sarcophaga triplasia: Downes 1965: 960. Sarcophaga (Neobellieria) triplasia: Pape 1996: 367. Robackina triplasia: Lopes 1975: 159; 1980: 229; Verves 1990: 542. Sarcophaga nigra Parker 1914: 38, as var. of Sarcophaga fulvipes Walker 1853 [junior secondary homonym of Myophora nigra RobineauDesvoidy 1830]. Type locality: USA: New York: Niagara Falls. Deposited in Massachusetts Agricultural College, Boston, USA. Sarcophaga dissidua Parker 1917: 157 [New name for Sarcophaga fulvipes nigra Parker 1914]. Sarcophaga fulvipes [misidentification; not Sarcophaga fulvipes Macquart 1842 and Walker 1853]: Aldrich 1916: 182.

Nearctic region: Canada: Ontario, Quebec; Mexico: Durango, Nuevo Leon; USA: Arizona, Arkansas, Florida, Georgia, Indiana, Iowa, Kentucky, Louisiana, Maryland, Mississippi, Missouri, New Mexico, New York, North Carolina, Ohio, Pennsylvania, South Carolina, Tennessee, Texas, Virginia.

Genus Rosellea Rohdendorf 1937 Rosellea Rohdendorf 1937: 242, as subgenus of Parasarcophaga Johnston & Tiegs 1921. Type species: Sarcophaga aratrix Pandellé 1896; by original designation.

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Parasarcophaga (Rosellea): Verves 1986a: 172. Rosellea: Fan & Pape 1996: 256; Lehrer & Martínez-Sanchez 2000: 259; Lopes et al. 1977: 573; Nandi 2002: 197, 372; Peris et al. 1999: 121; Povolný & Verves 1997: 226; Verves 1990: 542; Verves & Khrokalo 2006: 77, 177. Sarcophaga (Rosellea): Pape 1996: 386. Coeisca Rohdendorf 1966: 459, as subgenus of Boettcherisca Rohdendorf 1937. Type species: Sarcophaga khasiensis Senior-White 1924, by monotypy. Leigongshanophaga Lehrer & Wei 2010b: 8, n. syn. Type species: Sarcophaga catoptosa Wei & Yang 2007, by original designation.

Seven species are distributed in Nearctic, Palaearctic and Oriental regions.

Rosellea aratrix (Pandellé 1896) (Fig. 15, a–e) Sarcophaga aratrix Pandellé 1896: 191. Type locality: France: HautesPyrénées: Tarbes. Lectotype (♂; designated by Pape 2004: 23) deposited in Muséum Nationale d’Histoire Naturelle, Paris, France. Sarcophaga aratrix: Allen 1966: 228; Baudet 1985: 176; Bezzi 1907: 471; Böttcher 1912a: 730; Kramer 1909: 83; Pape & Merz 1998: 339; Séguy 1941: 75; Sarcophaga (Rosellea) aratrix: Kara & Pape 2002: 295; Pape 1996: 386; 2004: 23; Pape et al. 2002: 219; Papp 2001: 436. Parasarcophaga (Rosellea) aratrix: Artamonov 1980a: 31; 1987: 112; Blackith & Blackith 1984: 255; 1990: 699; Blackith et al. 1994: 427; Fan 1992: 711; Pape 1987: 162; Rognes 1986: 12; Rohdendorf 1937: 243; 1975: 201; Salaas 1943: 23; Verves 1986a: 172. Rosellea aratrix: Artamonov 1993: 226; Fan & Pape 1996: 256; Peris et al. 1999: 121; Povolný 1997: 100; 1999: 19; Povolný & Verves 1997: 226; Verves 1990: 542; 1998: 54; Verves & Khrokalo 2006: 178. Villeneuvella aratrix: Baranov 1942: 570; Enderlein 1928: 34. Sarcophaga kuntzei Kramer 1905: 13. Type locality: Germany: Oberlausitz. Holotype (♂) deposited in Staatliche Museum für Naturkunde, Görlitz, Germany. Sarcophaga pseudaratrix Baranov 1925: 2. Type locality: Serbia: Topčider near Belgrade. Lectotype (♂; designated by Sabrosky & Crosskey 1970: 432) deposited in U. S. National Museum, Washington, D. C., USA. Sarcophaga pseudaratrix: Sabrosky & Crosskey 1970: 432. Rosellea naumanni Lehrer & Martínez-Sanchez 2000: 258. Type locality: Spain: Alicante: Sierra Salinas (Villena). Holotype (♂) deposited in Unidad de Entomologia, Departamento de Ciencas Ambientales y Recurses Naturales, Universidad de Alicante, Spain.

Nearctic region: Canada: Manitoba; USA: Alaska. Palaearctic region: Albania; Armenia; Austria; Azerbaijan; Belgium; Bulgaria; Byelorussia; China: Liaoning, Neimenggu; Croatia; Czech Republic: Bohemia, Moravia; Denmark; Estonia: Finland; France; Germany; Gruzia (including Abkhazia); Hungary; Ireland; Italy: continental part, Corsica & Sicily; Kazakhstan; Kosovo; Latvia; Lithuania; Moldova; Norway; Poland; Romania; Russia: European part: Bashkortostan, Ivanovo, Krasnodar, Leningrad, Lipetzk, Moscow, Stavropol’, Voroniezh regions, West Siberia: Altay, Tomsk, Tuva regions, East Siberia: Buryatia, Irkutsk regions, Far East: Amur, Kamchatka, Khabarovsk, Magadan regions; Serbia; Slovakia; Slovenia; Spain; Sweden; Switzerland; Turkey; Ukraine: Cherkasy, Chernigiv, Chernivtsi, Crimea, Ivano-Frankivs’k, Kharkiv, Kherson,

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Kirovograd, Kyiv, Mykolayiv, Odesa, Poltava, Sumy, Vinnytsya, Volyn’, Zakarpattya, Zaporizhzhya, Zhytomyr regions; United Kingdom. Larvae are predators of another maggots, developing in small carcasses (insects, snails, birds, mice etc.), and also known as facultative predators of pupae of dendrophilous butterflies (Lymantria monacha) and parasites of adult cerambycid beetle Prionus coriarius (L. 1758). The adult flies visit the corpses, feces, flowering plants and decaying fruits. Flies prefer suburban gardens, humid forests and bogs at altitudes to 2000 m a. s. l. and tend towards culturophily.

Rosellea beckiana Lehrer 1996 Rosellea beckiana Lehrer 1996: 265. Type locality: Israel: Be’er Sheva. Holotype (♂) deposited in Institute of Systematics and Biology of University of Amsterdam, The Netherlands. Rosellea beckiana Lehrer 2006b: 19.

Palaearctic region: Israel.

Rosellea catoptosa (Wei & Yang 2007), n. comb. Sarcophaga catoptosa Wei & Yang 2007: 531. Type locality: China: Guizhou: Leigongshan National Nature Reserve, 800-1000 m a s. l. Holotype (♂) deposited in Centre for Disease Prevention and Control, Anshun City, Guizhou, China. Leigongshanophaga catoptosa: Lehrer & Wei 2010b: 8.

Oriental region: China: Guizhou.

Rosellea gigas (Thomas 1949) Sarcophaga gigas Thomas 1949: 166. Type locality: China: Sichuan: Chungking, Koloshan. Holotype (♂) deposited in National Institute of Health, Nanking, China. Sarcophaga (Rosellea) gigas: Pape 1996: 386. Parasarcophaga (Rosellea) gigas: Kano et al. 1967: 69; Park 1977: 264; Verves 1986a: 172. Rosellea gigas: Fan 1992: 711; Fan & Pape 1996: 256; Verves 1990: 542; Verves & Khrokalo 2006: 123. Sarcophaga koreaensis Park & Kano 1961: 116. Type locality: South Korea: Taegu, Mt Palgong. Holotype (♂) deposited in Tokyo Medical and Dental University, Japan.

Palaearctic region: China: Heilongjiang, Henan, Hubei, Jangsu, Liaoning, Sichuan; Russia: Far East: South Primorye; South Korea. Oriental region: China: Yunnan, Zhejiang. Flies were collected at borders of mountain forests.

Rosellea khasiensis (Senior-White 1924) Sarcophaga khasiensis Senior-White 1924: 246. Type locality: India: Assam: Cherrapunji, Khasi Hills. Holotype (♂) deposited in Natural History Museum, London, United Kingdom. Sarcophaga khasiensis: Bänzinger & Pape 2004: 1677; Kano & Shinonaga 1994: 261; Senior-White et al. 1940: 225; Sugiyama et al. 1988b: 357, 360. Sarcophaga (Rosellea) khasiensis: Pape 1996: 387; Tricholioproctia khasiensis: Lopes 1955: 274; Boettcherisca (Coeisca) khasiensis: Rohdendorf 1966: 459;


Parasarcophaga (Rosellea) khasiensis: Fan 1992: 711; Kano et al. 1967: 70; 1999: 139; Rosellea khasiensis: Fan & Pape 1996: 256; Lopes et al. 1977: 573; Nandi 1991: 40; 2002: 372; Verves 1990: 542; 2001: 243. Rosellea longwangiana Lehrer & Wei 2010a: 2, n. syn. Type locality: China: Guizhou: Ziyun, Bandang, 800 m, 15.07.1984. Holotype (♂) deposited in Center for Disease Prevention and Control, Guishou, China. Rosellea fuxingia Lehrer 2010a: 4, n. syn. Type locality not given, type material absent. Rosellea manipuriella Lehrer 2010a: 6, n. syn. Type locality not given, type material absent.

Palaearctic region: China: Sichuan. Oriental region: Bhutan; China: Guizhou, Yunnan; India: Jammu & Kashmir, Manipur, Meghalaya, Mizoram, Nagaland, Rajastan, Sikkim; Nepal; Thailand; Vietnam. Flies were collected on altitudes about 500–2500 m a. s. l., from bushes and mixed forests. Taxonomical notes: The original descriotion and drawings of genitalia of male Rosellea longwangiana are very detailed and practically are not differentiated from similar descriptions and drawings of ♂ genitalia of Rosellea khasiensis by Fan 1992; Kano et al. 1967; Nandi 2002; Rohdendorf 1966; Senior-White 1924 etc. Description of ♂ Rosellea manipuriella is based at drawings of ♂ genitalia of Rosellea khasiensis from Fan 1992, and the same of ♂ Rosellea fuxingia - at drawings of ♂ genitalia of Rosellea khasiensis from Nandi 2002. The differences in drawings are very petty and reflected the different styles of painters; they can not used as base for designations of the new species.

Rosellea spinipenis (Shinonaga & Tumrasvin 1979) Parasarcophaga spinipenis Shinonaga & Tumrasvin 1979: 144. Type locality: Thailand: Kanchana Buri: near Sai York. Holotype (♂) deposited in National Science Museum, Tokyo, Japan. Sarcophaga spinipenis: Pape 1996: 419 (not assigned to subgenus). Sarcophaga spinipes: Bänzinger & Pape 2004: 1687, incorrect subsequent spelling of spinipenis. Rosellea spinipenis: Verves 1990: 542.

Oriental region: Thailand. Flies were collected in forest by using the decaying meat bait.

Rosellea suthep (Pape & Bänzinger 2003), n. comb. Sarcophaga (Rosellea) suthep Pape & Bänzinger 2003: 52. Type locality: Thailand: Chiang Mai: Doi Suthep, above Sangwal School. Holotype (♂) deposited in Swedish Museum of Natural History, Stockholm, Sweden. Sarcophaga (Rosellea) suthep: Bänzinger & Pape 2004: 1677. Sarcophaga (Parasarcophaga) suthep: Shinonaga & Thinh 2003: 332. Parasarcophaga (Rosellea) aratrix: Kano et al. 1999: 138 (misidentification: not Sarcophaga aratrix Pandellé 1896).

Oriental region: Thailand; Vietnam. Flies were collected at altitudes 1000–1300 m a. s. l., in evergreen hill forests.

Genus Sabiella Verves 1990 Verves 1990: 541. Type species: Sarcophaga freyi Zumpt 1953, by original designation. Sabiella: Lehrer 2002: 53; 2003: 38, 404. Sarcophaga (Sabiella): Pape 1996: 387.

Two species are distributed in Afrotropical region.

Sabiella freyi (Zumpt 1953) (Fig. 16, a–c) Sarcophaga freyi Zumpt 1953b: 77. Type locality: South Africa: Transvaal [now: Mpumalanga]: Sabie. Holotype (♂) deposited in South African Institute for Medical Research, Johannesburg, South Africa. Sarcophaga (Prionophalla) freyi: Dear 1980: 814; Zumpt 1972: 46, 51, 77. Sarcophaga (Sabiella) freyi: Pape 1996: 387; Sabiella freyi: Lehrer 2003: 404; 2006a: 23; Verves 1990: 542. Parasarcophaga (Liosarcophaga) freyi: Rohdendorf 1963: 9.

Afrotropical region: South Africa: Eastern Cape, KwaZulu-Natal, Mpumalanga, Western Cape.

Sabiella mandelania Lehrer 2005 Sabiella mandelania Lehrer 2005: 40. Type locality: South Africa: KwaZulu-Natal: Itala Game Reserve, Louwsburg. Holotype (♂) deposited in Museum für Naturkunde der Humboldt-Universität, Berlin, Germany.

Afrotropical region: South Africa: KwaZulu-Natal.

Genus Saputaramyia Verves 2001 Verves 2001: 243. Type species: Parasarcophaga saputaraensis Nandi 1992, by original designation.

A single species is distributed in Oriental region.

Saputaramyia saputaraensis (Nandi 1992) (Fig. 17, a–e). Parasarcophaga (Liosarcophaga) saputaraensis Nandi 1992b: 189. Type locality: India: Gujarat: Saputara. Holotype (♂) deposited in Zoological Survey of India, Kolkata, India. Parasarcophaga (Liosarcophaga) saputaraensis: Nandi 2002: 253. Sarcophaga (Ziminisca) saputaraensis: Pape 1996: 415. Saputaramyia saputaraensis: Verves 2001: 243.

Oriental region: India: Gujarat.

Genus Sclerophalla Rohdendorf 1963 Sclerophalla Rohdendorf 1963: 5, 9, as subgenus of Parasarcophaga. Type species: Sarcophaga santosdiasi Zumpt 1951, by original designation. Sclerophalla: Lehrer 2002: 56; 2003: 42, 414; Verves 1990: 541.


Sclerophalla santosdiasi (Zumpt 1951) (Fig. 18, a–c) Sarcophaga santosdiasi Zumpt 1951: 180. Type locality: Mozambique: Maputo. Holotype (♂) deposited in South African Institute for Medical Research, Johannesburg, South Africa. Sarcophaga (Prionophalla) santosdiasi: Dear 1980: 813; Sarcophaga (Liopygia) santosdiasi: Pape 1996: 348; Zumpt 1972: 44, 63,180.

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Parasarcophaga (Sclerophalla) santosdiasi: Rohdendorf 1963: 9. Sclerophalla santosdiasi: Lehrer 2003: 414; Verves 1990: 542.

Afrotropical region: Mozambique; South Africa: Kwazulu-Natal.

Genus Takanoa Rohdendorf 1965 Takanoa Rohdendorf 1965: 690, 694. Type species: Sarcophaga hakusana Hori 1954, by monotypy. Takanoa: Kano et al. 1967: 161; Verves 1986a: 178; 1990: 541; Verves & Khrokalo 2006: 77, 178. Sarcophaga (Takanoa): Pape 1996: 410. Ussuriphalla Lehrer 2010b: 12, n. syn. Type species: Takanoa rugosa Rohdendorf 1969, by original designation.

Two species are distributed in southeastern part of Palaearctic region.

Takanoa hakusana (Hori 1954) Sarcophaga hakusana Hori 1954: 49. Type locality: Japan: Honshu: Ishikawa Prefecture, Mt. Hakusan. Holotype (♂) deposited in Kanazawa University, Japan. Takanoa hakusana: Fan & Pape 1996: 257; Kano et al. 1967: 31; Park 1974: 24; 1977: 254; Rohdendorf 1965: 694; Verves 1986a: 178; 1990: 542; Verves & Khrokalo 2006: 178. Sarcophaga (Takanoa) hakusana: Pape 1996: 410. Takanoa vervesiana Lehrer 2010b: 12, n. syn. Type locality not given, type material absent.

Palaearctic region: China: Liaoning; Japan: Honshu, Kyushu; South Korea. Flies were collected in mountain regions. Taxonomical notes: Description of ♂ Takanoa vervesiana is based at drawings of ♂ genitalia of Takanoa hakusana from Verves & Khrokalo 2006. The differences with other drawings of ♂ genitalia of T. hakusana are very petty and reflected the different styles of painters; they can not be used as base for description of a new species.

Takanoa rugosa Rohdendorf 1969 (Fig. 19, a–c) Takanoa rugosa Rohdendorf 1969: 947. Type locality: Russia: Southern Primorye: Shkotov District. Holotype (♂) deposited in Zoological Institute, Russian Academy of Sciences, St. Petersburg, Russia. Takanoa rugosa: Artamonov 1980a: 31; 1988: 30; Verves 1986a: 178; 1990: 542; Verves & Khrokalo 2006: 178. Sarcophaga (Takanoa) rugosa: Pape 1996: 410. Takanoa kolomyietzi Artamonov 1980b: 151. Type locality: Russia: Far East: Primorye, Ussuriysk District, environs of Kamenushka. Holotype (♂) deposited in Institute of Biology, Siberian Branch of Russian Academy of Sciences, Novosibirsk, Russia. Ussuriphalla qirimia Lehrer 2010b: 13, n. syn.

Palaearctic region: Russia: Far East: Southern Primorye. Larvae developed in dead fishes, freshwater crustaceans and mollusks on shores of natural freshwater reservoirs. Flies were collected on shores of streams, rivers and lakes. Taxonomical notes: Description of ♂ Ussuriphalla

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qirimia is based at drawings of ♂ genitalia of Takanoa rugosa from Verves & Khrokalo 2006. The differences with other published drawings of ♂ genitalia of T. rugosa are very petty and reflected the different styles of painters; they can not be used as base for designation of a new species.

Genus Ziminisca Rohdendorf 1965 Ziminisca Rohdendorf 1965: 689, as subspecies of Parasarcophaga Johnston & Tiegs 1921. Type species: Sarcophaga semenovi Rohdendorf 1925, by original designation. Parasarcophaga (Ziminisca): Verves 1986a: 173. Ziminisca: Verves 1990: 541. Sarcophaga (Ziminisca): Pape 1996: 415.

One species is distributed in Palaearctic (Central Asia) region.

Ziminisca semenovi (Rohdendorf 1925) (Fig. 20, a, b) Sarcophaga semenovi Rohdendorf 1925: 121. Type locality: Kazakhstan: “Kulandy-Aral, Turkestan, Aralsee”. Holotype (♂) deposited in Zoological Museum of Moscow Lomonosov University, Russia. Sarcophaga (Ziminisca) semenovi: Pape 1996: 415. Parasarcophaga (Liosarcophaga) semenovi: Rohdendorf 1937: 207. Parasarcophaga (Ziminisca) semenovi: Fan 1992: 703; Rohdendorf 1965: 689; Rohdendorf & Verves 1977: 726; 1978: 256; Verves 1986a: 173. Ziminisca semenovi: Fan & Pape 1996: 257; Verves 1990: 542. Sarcophaga linearis Villeneuve 1936: 8. Type locality: China: Xinjiang: Djungaria, Bugas. Holotype (♂) deposited in Stuttgarter Museum zur Naturkunde, Germany.

Palaearctic region: China: Gansu, Jilin, Ningxia, Xinjiang; Kazakhstan; Kyrgyzstan; Mongolia: BayanUlegey aimak; Russia: North Caucasus (Dagestan); Turkmenistan; Uzbekistan. Larvae developed in decaying meat in laboratory conditions. Acknowledgements. This study was supported by the National Nature Science Foundation of China (grant No. 30870330), The Ministry of Science and Technology of the People’s Republic of China (grant No. 2006FY110500) and the Foundation of Experimental Centre of Shenyang Normal University (grant No. SY200609).We are very grateful to Prof. A. Z. Lehrer, Tel Aviv University, Israel, and Dr. T. Pape, University of Copenhagen, Denmark, for accordance of copies of their very valuable works; to Dr. A. C. Pont, Oxford University Museum of Natural History, UK, and to Mr D. M. Ackland, of the same Museum for their kindly help over many years and valuable suggestions; to Mr. Chun-tian Zhang and Mr. Shu-chong Bai for providing the type specimens of Fanzideia cygnocerca.

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