and Lombok (Wallace's Line), while the remaining taxa were recorded either to the east or the west of this strait. ..... Marcia hiantina (Lamarck, 1818). 0 0 0. +.
Phuket Marine Biological Center Special Publication no. 19(2): 397-402 (1999) 397
MOLLUSCS FROM BEACHES ON BALI, LOMBOK, SUMBAWA, AND KOMODO ISLANDS, INDONESIA Jorgen Hylleberg Aarhus University, Department of Marine Ecology. Finlandsgade 14, 8200 Aarhus N, Denmark
ABSTRACT A total of 131 taxa of gastropods, bivalves and cephalopods were recorded by participants in the 9th TMMP Congress and Workshop. The material does not represent a systematic collection; rather the identifications reflect a limited number of study sites and taxonomic interests of the participants. The families Conidae, Littorinidae, Neritidae, Muricidae, and Olividae are well represented among the gastropods, and Cardiidae is well represented among the bivalves. Only 3 taxa (all cardiids) were recorded on all 4 islands; 10 species of gastropods (littorinids and lottidids) were recorded on both sides ofthe strait between Bali and Lombok (Wallace's Line), while the remaining taxa were recorded either to the east or the west of this strait.
INTRODUCTION Whenever an opportunity appeared, Mollusca were collected and identified during the 9th TMMP Congress and Workshop, 18-29 August 1998 in Indonesia. Most collections were made on Lombok and Komodo, while collections were rather sporadic on Sumbawa and Bali (Fig. 1).
Hence, the material does not represent a systematic collection along the shores, which otherwise would be very desirable. Records were mainly made during short stays at the sites, often where the bus incidentally stopped.
FLORES SEA
BALI SEA
INDIAN OCEAN 10'
1:3300000 L -_ _ _ _~------------------------------------~----~
115'
120'
Figure 1. The study area in Indonesia. Samples were collected on Bali, Lombok (including the Gili Islands), Sumbawa, and Komodo. Travel from Lombok eastward to Komodo by bus and ferry. From Lombok (Mataram city) westward to Bali (Denpasar) by air.
398
Tropical Marine Mollusc Programme (TMMP)
Many of the taxa are from the intertidal zone, while a few species have been observed subtidally during snorkelling (e.g. around the Gili Islands north of Lombok). However, many subtidal taxa were encountered because shells were washed up on the beaches, probably during storms. The identified material is biased towards taxonomic interests ofthe participants. This bias can be considered a weakness because the checklist only gives a glimpse of the incredible biodiversity of these Indonesian islands. However, an advantage is that the names are reliable, and we really need verified records from the area. I submit the data in the form of a checklist based on information I received from participants (alphabetical order, last part ofthe name as written according to the country traditions): Farnis Boneka, UNSRAT, Manado, Indonesia - Nguyen Chinh, RIA 3, Nha Trang, Vietnam - Rathe R. Jensen, Zoological
Taxonomic account GASTROPODA PROSOBRANCHIA CERITHIDAE Rhinoclauis uertagus (Linne, 1758) Pseudouertagus aluco (Linne, 1758) Terebralia sulcata (Bruguiere, 1792) COLUMBELLIDAE Pyrene testudinaria Link, 1807 CONIDAE Conus coronatus Gmelin Conus figulinus L . Conus imperialis L. Conus flauidus Lamarck Conus liuidus Hwass Conus rattus Hwass Conus coffeae Gmelin Conus music us Hwass Conus boeticus Reeve Conus planorbis Born Conus striolatus Kiener CORALLIOPHILIDAE Rapa rapa (L.)
Museum of Copenhagen, Denmark - Alan Kohn, University of Washington, USA Markus Lasut, UNSRAT, Manado, Indonesia - Jaruwat Nabhitabhata, Department of Fisheries, Thailand- Anuwat N ateewathana, Department of Fisheries, Thailand - David Reid, British Museum of Natural History, UK - Tan Koh Siang, National University of Singapore - Bernard Tursch, University Libre, Brussels, Belgium - Jacques Vidal, Museum National d'Histoire Naturelle, Paris, France.
CHECKLIST The checklist indicates family, species & authorship followed by the localities: (1) Bali, (2) Lombok, (3) Sumbawa, (4) Komodo. The last column shows identifiers of Prosobranchia, Opisthobranchia, Bivalvia and Cephalopoda; (0) = no record. Families are listed in alphabetical order.
1
2
3
4
Identifier(s)
0 0 +
0 0 0
+ + 0
0 0 0
Jorgen Hylleberg Farnis Boneka Farnis Boneka
+
0
0
0
Farnis Boneka
0 0 0 0 0 0 0 0 0 0 0
+ + + + + + + + + + +
0 0 0 0 0 0 0 0 0 0 0
0 + 0 0 0 0 0 0 0 0 0
Alan Alan Alan Alan Alan Alan Alan Alan Alan Alan Alan
0
0
0
+
Hylleberg, Reid,
Kohn Kohn Kohn Kohn Kohn Kohn Kohn Kohn Kohn Kohn Kohn
Phuket Marine Biological Center Special Publication no. 19(2): 397-402 (1999) 399
COSTELLARIIDAE Vexillum rugasum (Gmelin, 1791) CYMATIDAE Cymatium lotorium Linne, 1758 CYFRAEIDAE Cypraea caputserpentis (Linne, 1758) Cypraea moneta (Linne, 1758) Cypraea annulus (Linne, 1758) Cypraea lynx (Linne, 1758) Cypraea limacina Lamarck 1810 Cypraea tigris L. Cypraea staphylaea L. FISSURELLIDAE Hemitama pauhi (Quoy & Gaimard, 1834) Scutus unguis (Linnaeus) HALIOTIDAE Haliatis sq uamala Reeve LOTTIIDAE Patel/aida saccharina (L.) Patel/aida striata Quay & Gaimard, 1834 LITTORINIDAE Peasiella fasciata Reid & Mak, 1998 Peasiella canoida.l is (Pease, 1868) Peasiella roepstor{fiana (Nevill, 1885) Littararia earinifera (Menke, 1830) Littararia intermedia (Philippi , 1846) Littararia lutea (Philippi, 1847) Littararia seabra (Linne, 1758) Littararia pallescens (Philippi , 1846) Littararia undulata (Gray, 1839) Nadilittarina millegrana (Philippi, 1848) Nadilittorina sundaica (Altena, 1945) Nadilittarina trochoides (Gray, 1839) Nadilittorina vidua (Gould, 1859) Tectari"s teclumpersicum (L. ) MURICIDAE Morula anaxares (Kiener, 1836) Morula ceylonica (Dall, 1923) Morula fusca (Kuster, 1862) Morula granulata (Duclos, 1832) Morula margariticala (Broderip, 1833) Morula purpureocincta (Preston, 1909) Thais distinguenda (Dunker & Zelebor, 1866) Thais marginatra (Blainville, 1832) Thais muricoides (BJainville, 1832) Thais squamosa (Pease, 1868) Thais turbilloides (Blainville, 1832) Drupa (Drupina) grossularia Riiding, 1798
0
0
+
0
Farnis Boneka
+
0
0
0
Farnis Boneka
+ + + + +
0 0 0 0 0 0 0
0 0 0 0 0 0 0
+
Hylleberg, Boneka Jorgen Hylleberg Jorgen Hylleberg Jorgen Hylleberg Farnis Boneka David Reid David Reid
+ +
0 0
0
+
+
David Reid A. Kohn, D. Reid
+
0
0
0
David Reid
+
+ +
0 0
0 0
D. Reid, J . Hylleberg David Reid
+
+ + + + + + + + + + + + +
0
0
0 0 0 0 + 0 + + 0 0 0 0 0 +
0 0 + 0 0 0 0 0 0 0 0 0 0 0
David Reid David Reid David Reid David Reid D. Reid, F. Boneka David Reid D. Reid, F. Boneka D. Reid, F. Boneka D. Reid, F. Boneka D. Reid, F. Boneka David Reid Reid, Boneka D. Reid, F. Boneka David Reid
0 0 0 0 0 0 0 0 0 0 0 0
+ + + + + + + + + + +
0 0 0 0 0 0 0 0 0 0 0 0
0 0 0 0 0 0 0 0 0 0 0 0
Tan Koh Tan Koh Tan Koh Tan Koh Tan Koh Tan Koh Tan Koh Tan Koh Tan Koh Tan Koh Tan Koh Tan Koh
0 +
0
0 0 0 0 0 + 0 + + + + + +
+
0 0 0 0 +
0
Siang Siang Siang Siang Siang Siang Siang Siang Siang Siang Siang Siang
400
Tropical Marine Mollusc Programme (TMMP)
NACELLIDAE Cellana testudinaria (Linnaeus, 1758) Cellana radiata orientalis (Pilsbry 1891) NASSARIDAE Nassarius gaudiosus (Hinds, 1844) NATICIDAE Mamilla melanostoma (Gmelin, 1791) Polinices (Polinices) mammilla (L.) NERITIDAE Nerita albicilla Linne, 1758 Nerita antiquata Recluz, 1841 Nerita costata Gmelin, 1791 Nerita chamaeleon L. Nerita exuvia L. Nerita georgina Recluz, 1841 Nerita histrio L. Nerita planospira Anton, 1839 Nerita plicata Linne, 1758
Nerita polita Linne, 1758 Nerita reticulata Karsten, 1789 Nerita undata L. Neritina reticulata Sowerby, 1838 Smaragdia souverbiana (Montrouzier, 1863) OLIVIDAE Oliva bulbiformis Duclos, 1840 Oliva cameola (Gmelin, 1791) Oliva elegans Lamarck, 1811 Oliva oliva (Linnaeus, 1758) Oliva reticulata (Roding, 1798) Oliva sericea (Roding, 1798) Oliva tigridella Duclos, 1835 Oliva todosina Duclos, 1835 OVULIDAE Calpumus verrucosus (L. ) [On soft coral] PATELLIDAE Scutellastra flexuosa (Quoy & Gaimard, 1834) PLANAXIDAE Planaxis sulcatus (Born, 1778) STROMBIDAE Strombus aurisdianae Linne, 1758 Lambis lambis (L.) Strombus erythrinus Dillwyn, 1817 TEREBRIDAE Terebra nebulosa Sowerby TONNIDAE Tonna cepa (Roding, 1798)
0 +
+ 0
0 0
+ 0
J. Hylleberg, D. Reid David Reid
0
+
0
0
Farnis Boneka
0 0
+ +
0 +
+ 0
Markus Lasut Markus Lasut
0 0 0 0 0 0 0 0 +
+ 0 + + + + + + +
+ + 0 0 0 0 0 0 0
+ + 0 0 + 0 0 0 +
0 0 0 0 0
+ 0 0 + +
0 + + 0 0
+ + + 0 0
D. Reid, F. Boneka David Reid David Reid David Reid David Reid David Reid David Reid David Reid D. Reid, J. Hylleberg F.Boneka D. Reid, J. Hylleberg David Reid David Reid David Reid David Reid
+ + 0 0 + 0 0 +
0 0 + + + + 0 0
0 0 0 0 0 0 + 0
0 0 0 0 + 0 0 0
Bernard Bernard Bernard Bernard Bernard Bernard Bernard Bernard
0
0
0
+
A. Kohn, D. Reid
0
+
0
0
David Reid
0
+
0
0
Farnis Boneka
0 0 0
0 + +
+ 0 0
0 0 0
Jorgen Hylleberg Alan Kohn Farnis Boneka
0
0
0
+
Jorgen Hylleberg
+
0
0
0
Farnis Boneka
Tursch Tursch Tursch Tursch Tursch Tursch Tursch Tursch
Phuket Marine Biological Center Special Publication no. 19(2): 397-402 (1999) 401
TROCHIDAE Troehus maculatus Linne, 1758 Teetus fenestratus (Gmelin, 1791) Monodonta labio (Linne, 1758) Monodonta canalifera Lamarck, 1816 TURBINIDAE Lunella cinerea (Born, 1778) Turbo petholatus L OPISTHOBRANCHIA CEPHALASPIDEA Hydatina amplustre (1.) Bulla ef. ampulla (L. ) NOTASPIDEA Pleul'obranehus forskalii SACOGLOSSA Berthelinia s p. (bivalved) Costasiella sp. Dolabella auricularia ElysiR.lla pusilia Bergh, 1872
0 0 0 0
+ + 0 +
0 0 0 0
+ + + 0
J orgen Hy lie berg Farnis Boneka Farnis Boneka David Reid
0 0
0 +
0 0
+ 0
D. Reid, F. Boneka AJan Kahn
0 0
+ +
0 0
0 0
Kathe R. Jensen Kathe R. Jensen
0
+
0
0
Kathe R. Jensen
0 0 0 0
+ + 0 0
0 0 0 0
0 0 + +
Kathe Kathe Kathe Kathe
+ + + + 0 + 0 0 + +
+ + 0 0 0 + + 0 + +
+ + 0 0 0 + + + + +
Jacques Vidal J. Vidal, J. Hylleberg Jacques Vidal J. Vidal, J. Hylleberg Jorgen Hylleberg J. Vidal, J. Hylleberg Jacques Vidal J. Vidal, J. Hylleberg Jacques Vidal Jacques Vidal
+
+
+
J. Vidal, J. Hylleberg
0
+
+
David Reid
+
0
0
David Reid
+
0
0
Jorgen Hylleberg
0
0
+
David Reid
BIVALVIA CARDIIDAE Fulvia papyracea (Bruguiere, 1789) 0 Fragum unedo (Linne, 1758) + Fragum SCl'uposum (Deshayes, 1855) 0 Fragum fragum (Linne, 1758) 0 Laevicardium cf. biradiatum (Bruguiere) + Maoricardium setosum (Redfield, 1846) + Nemocardium lyratum (Sowerby, 1841) + Vasticardium angulatum (Lamarck, 1819) 0 Vastieardium orbita philippinense (Hedley, 1899)+ Vasticardium /lavum subrugosum (Sow., 1840) 0 Vastieardium pectini(orme Born, 1780 = rugosum Lamarck , 1819 0 HEMIDONACIDAE Hemidonax donaciformis (SchrDter) 0 LUCINIDAE Anodontia philippiana (Reeve, 1858) 0 MACTRIDAE M aetra maeulata 0 MESODESMATIDAE Ataetodea striata (Gmelin , 1791) 0 PECTINIDAE Comptopallium radula (1.) 0 PINNIDAE Pinna cf. bicolor Gmelin, 1791 0
R. R. R. R.
Jensen Jensen Jensen Jensen
0
0
+
David Reid
0
0
+
Kathe R. Jensen
402
Tropi-cal ilI/arine Mollusc Programme (TMMP)
TRIDACN1DAE Trid.aclw maxim.a (ROding 1798) VENERIDAE Anomalodiscus squamosus (L.) Antigona (Periglypta) lacerata, (Hanley, 1830) Circe scripta (Linne, 1758) Marcia hiantina (Lamarck, 1818) Pilar pellucidus (Lamarck, 1818) Tapes literalus (Linne, 1758)
+
0
0
0
Jorgen Hylleberg
0
0
0 0
0 + 0
+ 0 0 0
0 0
0 0
0 + 0 + + +
David Reid Nguyen Chinh D. Reid, J. Hylleberg David Reid David Reid J. HylJeberg, D. Reid
0 0 0
CEPHALOPODA NAUTILIDAE Nautilu.s pompilius Linne, 1758 SEPIIDAE Sepia pharaonis Ehrenberg
0
0
0
+
Jorgen Hylleberg
0
0
0
+
Jaruwat
Sepia lalimanus Quoy & Gaimard, 1832
0
0
0
+
Nabhitabhata A. Nateewatbana
