Lepidoptera

(Lepidoptera)

INSECTS OF KOREA Series 11

By Vladimir Kononenko and Hui Lin Han

Series Editor : Park, Kyu-Tek Ass. Editor : Byun, Bong-Kyu

KOREAN NATIONAL ARBORETUM & CENTER FOR INSECT SYSTEMATICS KOREA 2007

INSECTS OF KOREA Series 11 Copyright 2007ⓒ by the Center for Insect Systematics, Korea All rights reserved. No part of this book may be reproduced or translated in any form, by print, photoprint, microfilm, microphiche, disk, CD-room or any other means without written permission from the publisher. Editor in Chief: Kyu-Tek Park Photographs: Vladimir S. Kononenko, H. L. Han, M. Y. Chae Plates and cover page design: Vladimir S. Kononenko Part of the series “Insects of Korea” should be cited as following example: Kononenko, V. S. and H. L. Han, 2007. Atlas genitalia of the Noctuidae in Korea (Lepidoptera). In Park, K. T. (ed.): Insects of Korea [11], pp. 464.

Authors Vladimir Stepanovich Kononenko Laboratory of Entomology, Institute of Biology and Soil Science, Far Eastern Branch of Russian Academy of Sciences, Vladivostok 22, Russia 690022 E-mail: [email protected] Hui Lin Han School of Forestry, Northeast Forestry University, Harbin, Heilongjiang, 150040 China E-mail: [email protected] ISBN: 978-89-88154-55-7-94490 ISBN 978-89-88154-04-5 (Series)

FOREWORD Our planet’s essential goods and services depend on the variety and variability of genes, species, populations and ecosystems. Biological resources provide us foods, clothes, medicines, housing materials, and spiritual nourishment. The natural ecosystems of forests, pasture and rangelands, deserts, rivers, lakes, and seas contain most of the earth’s biodiversity. Farmer’s fields and gardens are also great importance as repositories, while gene banks, arboretums, botanical gardens, zoos, and other germplasm repositories make a small but significant contribution. The current decline in biodiversity is largely the result of human activity and represents a serious threat to human development. The loss of the world’s biological diversity, mainly from the habitat destruction, overharvesting, pollution and inappropriate introduction of foreign plant and animals, has continued. Biological resources constitute a capital asset with great potential for yielding sustainable benefits. Urgent and decisive action is needed to conserve and maintain species and ecosystems, with a view to the sustainable management and use of biological resources. Capacities for the assessment, study and systematic observation and evaluation of biodiversity need to be reinforced at national and international levels. Effective national action and international cooperation are required for the in situ protection of ecosystems, for the ex situ conservation of biological and genetic resources and for the enhancement of ecosystem functions. The conservation of biodiversity is not only just a matter of protecting wildlife in nature reserves, but also fundamental element to the success of the development process. It is also about safeguarding the natural systems of the earth that are our life-support systems; purifying the waters; recycling the oxygene, carbon, And other essential elements; maintaining the fertility of the soil; providing the food from the land, and seas; yielding medicines; and safeguarding for the genetic richness. On the point of view of environmental sustainable use and related activities including UNCED program, the development of biodiversity conservation technology became the urgent work we will have to do. As one of the representative institute for the researches on biological resources in Korea, we, Korea National Arboretum, have conducted various projects to investigate the biological diversity in the forest ecosystem, including plants, insects and microorganisms since established in 1999. In this aspect, I am pleased to publish this book for understanding the insect diversity in the forest ecosystem. This series could not be published without the devotion of editors and authors. I would like to express my thanks to them. I hope that this book will be helpfully used, not only for domestic, but also foreign workers, to identify species of noctuid moths, and to understand the noctuid fauna of the Korean peninsula and the adjacent territories.

August 2007 Park, Chong Seo Director General, Korea National Arboretum 1

EDITORIAL PREFACE This book is an extended volume of the “Illustrated Catalogue of Noctuidae in Kononenko”, Ahn, Ronkay, 1998), as the series number 11 of “Insects of Korea”. The book presents the extensive faunal revision of the Noctuidae in the Korean peninsula with illustrations of the male and female genitalia of all available species of Noctuidae inhabiting the region, since almost 10 years after publication of the comprehensive catalogue. The book is a result of the international collaborative work between Russian specialist, Dr. V. S. Kononenko, Institute of Biology and Soil Science, the Far Eastern Branch of Russian Academy of Science, Vladivostok, Russia and Korean workers and especially Chinese specialist, Dr. Hui Lin Han, who studied the Korean Noctuidae during last six years for his course studies and Post-Doct. in Korea, and now a staff of School of Forestry, Northeast Forestry University, Harbin, Heilongjiang, China. Although the study of the Lepidoptera of Korean peninsula has been started from the end of 19C, as well as in some other Asian countries. The faunistic study or the taxonomic works on Lepidoptera, including noctuids, in the Korean peninsula is far from a complete inventory, especially in its northern part. The main reasons are the great diversity of the family including over 1,000 species in the region, lack of specialists, literature, materials, and information from the adjacent territories of the peninsula. It is well known that the insect fauna in the Korean peninsula is closely related to that of south of the Russian Far East and North East China. For these reasons, I have supported the authors to make international collaborative works during last 5 years, and to publish this book. The main purpose of the publication is to help an easy identification of the species with illustratations of the male and/or female genitalia for all Korean species of Noctuidae. It is no doubt that the most reliable tool for identification of the species is the structure of the male and female genitalia and they play important roles in the taxonomy and systematic of the Noctuidae. In these reasons, this publication, “Atlas genitalia of Noctuidae in Korea” will be very helpful for the identification of the species, but also the systematic study of Noctuidae for home workers or foreign researchers who have interests in the taxonomic study of Noctuidae. In this publication, 1,044 species of Noctuidae are enumerated for the Korean peninsula, among them 79 species including 15 newly reported species for Korea are added to the faunal list since the publication of the Illustrated Catalogue of Korean Noctuidae (1998), and four species will be described new to science. We hope that this book will be helpfully used for the study of preservation of insect diversity, for the identification of noctuid-species, and for understanding the noctuid fauna of the Korean peninsula and the adjacent territories.

Park, Kyu-Tek Editor

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AUTHOR’S PREFACE About 10 years have been passed since the publication of “Illustrated Catalogue of Noctuidae in Korea” by Kononenko, Ahn and Ronkay (1998), in which 961 species of the Korean Noctuidae are illustrated and catalogued. The publication has inspired all researchers for the lepoidopteran fauna in Korea and the other countries, and initiated further faunistic and biodiversity researches in the region. During the last decade, numerous species have been newly discovered from the Korean peninsula and neighboring regions or described new taxa for science, and the subfamily (tribe) Nolinae has been revised increasing the number of species occurring in Korea. At the same time, our knowledge on noctuids of the Palaearctic region as a whole, as well as that of the Pacific region of Asia, has been improved considerably. The Noctuidae has become a subject of an intensive study by many lepidopterists in Europe, Asian countries, and North America. The taxonomy, nomenclature, and classification of the Noctuidae have undergone with dramatic changes. Some subfamilies and genera or other taxonomic groups have been often changed in their taxonomic status and nomenclature. The higher classification of the Noctuoidea has newly proposed by Lafontaine and Fibiger (2006), using morden cladistic methodology, the discovery of new morphological characters, especially in genitalia, and the combination of comprehensive morphological characters of adults and immature, and the molecular analysis. The new system changed subfamilies, tribal and subtribal division of the Noctuidae. The present book aims two goals: The first is to revise the checklist of the Noctuidae in Korea. It is summarized with all recent information on the noctuids-fauna of the Korean peninsula, based on recent faunistic and biological data, and literatures for the fauna of Korea during last 10 years. It was also rearranged all taxa, a total of 1,042 species, according to the latest taxonomic and nomenclatural revision of Noctuoidea by Lafontaine and Fibiger (2006). The second is to provide all available illustrations of the male and female genitalia for the easy identification of the Noctuidae in Korea and adjacent territories. Due to the high diversity of the Noctuidae and their often-uniform external appearance in many groups, it is one of the difficult group for identification in Lepidoptera. The correct identification of them can only be possible by the examination of genitalia and comparing genitalia preparates with authentical preparates or illustrations. The both series: the catalogue preiously published in 1998 and this publication would be helpfully used for students and museum researchers in identification of collections, and also for ecologists, applied entomologists in agriculture or plant quarantine field. With the development of the molecular study, the illustrations of the genitalia are helpful with special significance for the correct identification of specimens which are fixed in alcohol for further molecular examination. We planed to illustrate the genitalia of Korean Noctuidae, following after the Illustrated Catalogue of Korean Noctuidae, however, this project has practically been postponed for almost 10 years. At present, with the development of digital photography and new techniques, it become possible to enumerate all data and figures in short period. In the course of the preparation of this book, we dissected made or female genitalia and made permanent slide of more than 3,000, and the best of 3

them are illustrated here. This book is a result of the friendship and scientific cooperation between of Korean, Russian and Chinese lepidopterists. For this work of the Atlas genitalia of Korean Noctuidae, we have been supported not only for financial support, but also constant help and encouragement from the former president of the Center for Insect Systematics and the editor of the series “Insects of Korea”, Dr. Kyu-Tek Park, and also a financial support for the printing this book by Korea National Arboretum. Thus, this publication is dedicated to Professor Kyu-Tek Park who is retiring from the position of professor, Kangwon National University in this August, 2007. We hope that this book will helpfully be used by home and foreign researches for further study of insect diversity in Pacific Asia region, and be comprehensive guidance for the identification of species, together with the “Illustrated Catalogue of Noctuidae in Korea”.

Authors July, 2007

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ACKNOWLEDGEMENTS The preparation and publication of this book would be impossible without unstinting helps from many lepidopterist colleagues, both professional and amateur. We are cordially thank following persons for their help in providing us information, valuable advices and comments, and for the loan of photographs of type specimens or preparates : Dr. D. Stüning, Dr. W. Speidel (ZMFK, Bonn, Germany), Mr. G. Behounek (Grafing, Germany), Mr. M. Fibiger (Denmark, Sorø), Dr. K. Mikkola, Dr. J. D. Lafontaine (Agriculture Canada, Research Branch, Ottava, Canada); Dr. L. Ronkay (HNHM, Budapest), Mr. G. Ronkay (Budapest, Hungary); Mr. S. Sugi, (Japan, Tokyo), Mr. K. Eda (Japan), Dr. M. Owada (NSM, Tokyo), Dr. S.-I. Yoshimatsu (NIAST, Tsukuba, Japan), Dr. A. Matov, (ZI, St. Petersburg, Russia); Mr. J. C. Sohn (Republic of Korea), Dr. C. Nguyen (NIPP, Hanoi, Vietnam). We are greatly thankful to Dr. M. Owada, Dr. J. Holloway, and Malaysian Natural Society and Southdene Sdn. Bhd., Dr. M. Lödl, Mr. M. Fibiger, Mr. H. Hacker, and Dr. H. Inoue for the permission to use their illustrations for this book. Our great thanks to Dr. L. Ronkay and Mr. G. Ronkay for fruitful discussion of many critical taxa and their kind help in course of study of the Korean collection in HNHM. We are indebted to Mr. M. Fibiger and Dr. D.J. Lafontaine for many valuable advices concerning classification of the Noctuoidea; the late Prof. C.M. Naumann (ZMFK, Bonn, Germany), for his kind support of museum work for the preparation of this book. Also thanks to Mr. J.C. Sohn for valuable information about new records of additional species in Korea. We are deeply grateful to Prof. Dr. K.T. Park, the editor of the series “Insects of Korea”, the former chief of the Center of Insects Systematic in Kangwon National University, initiator and organizer of numerous lepidopterological projects for his support and encouragement and for his efforts for the publication of this book. With special warm feeling, we remember the late Dr. S.B. Ahn, who was very enthusiastic Noctuidae researcher in Korea and a co-author of the Illustrated Catalogue of Noctuidae in Korea. Authors are also indebt to graduate students, Kangwon National University; Miss Min-Young Kim, and Eun-Mi Ji for their help in preparation of this book, especially to Miss Min-Young Chae for the preparation of excellent genitalia preparates. The museum work for this project has been financially supported by KOSEF in supporting for the first author’s visiting to Korea since 1994. The first author in part was supported by grants from the Finnish Academy in the year 2003, Deutsche Forschungsgemeinschaft (DFG grant Nr. 436 RUS 17/47/04) in the years 2004, by Ernst Mayer Traveling grant from the Museum of Comparative Zoology Harvard University, USA in the year 2004. We wish to express our thanks the Center for Insect Systematics and the Korea National Arboretum, who gave a financial support for the printing of this publication. Finally, we thank the publisher, Miss K. H. Cho, the president of Jeunheng-Sa publishing company, for her heartful contribution in printing this publication.

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CONTENTS

Foreword ∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙1 Editorial Preface ∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙2 Author’s Preface ∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙3 Acknowledgements ∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙5 Introduction ∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙9 Materials and Methods ∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙11 Synopsis ∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙14 Genitalia morphology and terminology ∙∙∙∙∙∙∙∙∙∙∙∙15 Classification ∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙20 Systematic list ∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙29 Family Micronoctuidae ∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙29 Family Noctuidae ∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙29 Subfamily Rivulinae ∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙29 Subfamily Hypenodinae ∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙30 Subfamily Araeopteroninae ∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙31 Subfamily Eublemminae ∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙31 Subfamily Herminiinae ∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙39 Subfamily Hypeninae ∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙45 Subfamily Phytometrinae ∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙48 Subfamily Aventiinae ∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙50 Subfamily Erebinae ∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙50 Subfamily Calpinae ∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙51 Subfamily Catocalinae ∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙53 Subfamily Stictopterinae ∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙65 Subfamily Euteliinae ∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙65 Subfamily Nolinae ∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙66 Subfamily Arctiinae ∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙74

Subfamily Lymantriinae ∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙75 Subfamily Plusiinae ∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙75 Subfamily Eustrotiinae ∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙79 Subfamily Bagisarinae ∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙83 Subfamily Acontiinae ∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙84 Subfamily Pantheinae ∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙85 Subfamily Raphiinae ∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙85 Subfamily Balsinae ∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙86 Subfamily Acronictinae ∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙86 Subfamily Metoponiinae ∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙89 Subfamily Sinocharinae ∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙90 Subfamily Agaristinae ∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙90 Subfamily Cuculliinae ∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙90 Subfamily Oncocnemidinae ∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙91 Subfamily Amphipyrinae ∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙92 Subfamily Psaphidinae ∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙92 Subfamily Heliothinae ∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙93 Subfamily Condicinae ∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙95 Subfamily Eriopinae ∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙97 Subfamily Bryophilinae ∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙98 Subfamily Xyleninae ∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙100 Subfamily Hadeninae ∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙120 Subfamily Noctuinae ∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙128 References ∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙136 Plates ∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙143 Index ∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙∙449

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INTRODUCTION The Noctuidae (s. l.) is the largest, most diverse, worldwide-distributed family of the order Lepidoptera with over 35,000 species in the World, i.e. about 20~25% of the total number of Lepidoptera species. Well over 10,000 species are known from the Palaearctic region. The representatives of this family have great ecological as well as economic significance. It is known that larvae of the most Noctuidae are polyphagous, consuming mainly vascular plants. They are an important component of natural or agricultural systems due to their high specific diversity, the abundance of many species, high migratory ability of imagos, and phytophagous feeding habits of larvae. The identification of the Noctuidae is rather a difficult task, by the reasons of their great number of species, which are often represented by taxonomic complexes of sibling species, contradiction and controversial classification of the group and lacking adequate literature. The analysis of superficial morphological characters is often not enough for correct identification of species. The noctuids fauna of the Korean peninsula as a part of Manchurian subregion of Palaearctic is very rich, diverse, peculiar and saturated by Manchurian and Oriental faunal elements and considerably different from the other subregions of the Palaearctic, with 1,042 known species. In the comparison, with neighboring countries located in the same biogeographical regions, the Noctuidae fauna of Japan enumerated about 1,200 species and that of the South of Russian Far East is about 800 species. In result of human activity, developing and changing of agricultural system during last decades some species of noctuids became rare or vanishing in Korea, but some of the others are still serious pests of agricultural plants. Because of the transitional position of the Korean peninsula which is Mandschurian subregion of the Palaearctic and Oriental biogeographic region, many tropical migrating species occur regularly or occasionally here, and some of them have economic importance as serious quarantine pests. The inventory of the noctuids fauna of the Korean peninsula has been started from the end of XIX century by British entomologists J. H. Leech (1898a; 1898b, 1900). The first list of Korean Noctuidae was completed by the late Pak (1959), the significant part of Korean Noctuidae was illustrated by Nam (1982) and the faunal list has been compiled (Checklist of insects from Korea, 1994). A complete historical review of Lepidopterology in Korea has been done by Park (1976) and that concerning Noctuidae by Kononenko et al. (1998). The most comprehensive catalogue summarized the knowledge about noctuidae fauna of the Korean peninsula was published by Kononenko et al. in 1998, enumerating 961 species of the Noctuidae from Korea, without subfamily Nolidae (at present tribe Nolini in Noctuidae). During subsequent years numerous additions to Korean fauna have been reported by Korean and foreign specialists (Kim, 1999; Kim, Choi, 2000; Sohn, Ronkay, 2001; Sohn, Kim, 2003; Sohn, Han, 2005; Sohn, Ronkay and Choi, 2005; Sohn, Ronkay and Kononenko, 2004; Sohn, Choi and Cho, 2005; Sohn, Ronkay, Kim and Cho, 2005; Han, Kononenko, Park. 2005; Han, Li, Byun, 2007; Han, Byun, 2007; Han, Park and Lu, 2005). Moths of North Korea and the genus Catocala in Korea were 9

reviewed (Park et al., 2001; Park et al., 2006), and the subfamily (at present tribe) Nolidae in Korea was revised by Oh (1991, 2001). At the same time the taxonomy, classification and nomenclature of the Noctuidae have undergone dramatic changes. Some subfamilies in issues, Noctuidae Europaea (Ronkay et al, 2001, Hacker et al., 2002; Goater et al., 2003; Zilli et al., 2005), and Moths of Borneo (Holloway, 2005) and many genera of European and Asian fauna were revised (Hacker, 1998; 2004; Holloway, 1998; 2005; Holloway and Miller, 2003; Pogue, 2002; Owada et al, 2006); numerous changes introduced in the nomenclature (Hacker, 1998; Sugi, 2000; Owada, 2006), and numerous additions, taxonomic and nomenclature changes were published for neighboring Japanese fauna (Sugi, 2000; Sugi and Jinbo, 2004). The Keys for the Noctuidae of the Russian Far East were completed (Kljuchko, 2003, Kononenko, 2003, Sviridov, 2003). The checklists based on modern classifications were published for Europe (Nowacki and Fibiger, 1997; Fibiger and Hacker, 1998, 2005) and for the Asian part of Russia (Kononenko, 2005). The Nolidae was fixed as a subfamily of the Noctuidae (Holloway, 1998) and the nomenclature of the family group names of noctuioid moths was reviewed by Speidel and Naumann (2005). The Noctuidae, like many other families of Lepidoptera have been controversial on its classification of higher taxa. The former classifications of Noctuidae, based mainly on Hampson’s (1903-1913) classical work, used by researchers during the last over 100 years, has become outdated for the time being. During the last ten years, the higher classification of the Noctuoidea has been undergone with radical changes and modification due to usage of cladistic methodology, the discovery of new morphological characters, especially in genitalia, combination of comprehensive morphological study of adults and immature, and introducing the molecular study into classification of higher taxa of the Noctuidae. The most recent and most comprehensive morphological classification of the Noctuidae with account of imago and immature morphology as well as molecular data is those of Fibiger and Lafontaine (2005) and its latest modification (Lafontaine and Fibiger, 2006). In the present Checklist, we accepted the composition of subfamilies proposed by Fibiger and Lafontaine (2005, 2006), as the latest proposed classifications applicable to Holarctic fauna. However, because we did not treat the families Arctiidae and Lymantriidae in the Illustrated Catalogue of Korean Noctuidae (Kononenko et al., 1998), we do not include them to the present book. The modern subfamily Nolidae has been partly treated in the Catalogue (as subfamilies Sarrothripinae and Chloephorinae); therefore we include Nolidae (as Nolini) to the present study. Because of the genus Mimachrostia of current Micronoctuidae was included to the Catalogue, we treat the Micronoctuidae in this book. The order of subfamilies in recently proposed classification in relation to classification used by Kononenko et al. (1998) is presented in the [Table 1].

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MATERIALS AND METHODS The present Checklist is based on the examination of the materials collected by Korean researchers and students in the Center for Insect Systematic, Kangwon National University (CIS), Korean National Arboretum (KNA, Gwangneung) and the National Institute of Agriculture and Science Technology of the Rural Development Administration (NIAST, Suwon) and some other materials. Each species is provided with the reference to the Illustrated Catalogue of Korean Noctuidae with page number and catalogue number. The species reported for Korea after publication of the Illustrated Catalogue of Noctuidae in Korea or newly reported in the present checklist are marked by asterisk (*) and provided with the reference for the first report. For these taxa, we give full catalogue information as same as in the previous catalogue (i.e. original combination, citation of description, type locality and place of type(s) depository, full synonymy and distribution). The valid names are indicated by bold italic and the synonyms, subtribial and subgeneric names by regular italic characters. The paragraph “Notes” includes taxonomic, faunal or nomenclature notes and corrections for some species. The information for the generic names, its type species, authors and year of description were the catalogue “The generic names of moths of the world” (Nye, 1975). The sources of information on specific names, type localities, synonyms were “Noctuidae. Lepidopterorum Catalogues (New Series). Fasc. 118” by Poole (1989), taxonomic database loaned from Mr. Gottfried Behounek and “Lepindex” - the internet site of the Museum of Nature History, London (Becaloni et al. 2003). Most species of Korean Noctuidae are illustrated here with 995 species of the male and 960 species of the female genitalia, based on more than 3000 dissections. The genitalia slides were photographed using the Nikon Coolpix 950, 990 and 4500 digital cameras and LM-scope digital microscope adapter with various binocular microscopes. Many slides have been photographed with excellent results in CIS by Olympus C707D digital camera and Olympus BX50 microscope. Digital photographs have been processed and plates completed with Adobe Photoshop 7/CS graphic program. Each illustrated species in the systematic list is supplied with references to the plate and number of genitalia on the right side of the page. For each illustration on plates we provide following information: species name, country, and preparate number, acronym of preparatory name and acronym of the museum or collection depository. Most dissected specimens were originated from Korea, however for some routine species or in the case, if Korean specimens were unavailable for dissection, we used for the preparation of specimens from the other countries (Japan, China, Vietnam, Russia, Europe). Some species, especially those reported from Korea by early authors have not been found in the examined collections, or some materials, used for Illustrated Catalogue of Noctuidae in Korea, were unavailable for present study. In those case, we used the illustrations of genitalia from publications of the other Noctuidae researchers (Owada, 1987, 2006 [2007]; Lödl and Gaal, 1998; Lödl and Mayerl, 1998; Holloway, 2005; Choi, 2006). In the course of the identification and the revision of the noctuids from Korea, many type11

specimens of taxa described from Korea, Japan, the Russian Far East and neighboring territories and other material were examined from following museums: Zoological Museum of Humboldt University, Berlin, Germany (Zoologisches Museum für Naturkunde zu Humboldt Universit(t); Zoological Institute and Museum Alexander Koenig, Bonn, Germany (Zoologisches Forschungsinstitut und Museum Alexander Koenig); the Natural History Museum [British Museum (Natural History)], London, Great Britain; Swedish Museum of Natural History (Naturhistoriska Riksmuseet), Stockholm, Sweden; Entomological Institute Hokkaido University, Sapporo, Japan, National Science Museum, Tokyo, Japan; National Institute of Agro-Environmental Science, Japan, Tsukuba. The names of the museums are referred by acronyms (see list of museum and institutions).

Acronyms of persons who prepared genital preparations AM - Alexei Matov ASB. - Seung Bok Ahn AZ - Alberto Zilli BG - Barry Goater CB - Charl Boursin CY - Min-Young Chae DL - Don Lafontaine

ER - Erik Rukborn GB - Gottfried Behounek HL - Hui-Lin Han JK - Jakko Kullberg JN - Junus Nowacky JZ - Jannifer Zaspel KM - Kauri Mikkola

LR - Laszlo Ronkay MF - Michael Fibiger MH - Marton Hreblay NC - Nguyen Nhu Cuong SJC - Jae-Cheon Sohn SS - Shigero Sugi VK - Vladimir Kononenko

Acronyms of museums BMNH - The Nature History Museum (British Museum Nature History) (London, UK) CIS - Center for Insects Systematics (Chuncheon, Korea) HNHM - Hungarian Nature History Museum (Budapest, Hungary) IBP - Institute of Biology and Soil Science (Vladivostok, Russia) KNAE - Korean National Arboretum (Gwangneung, Korea) MNHU - Humbold University Nature History Museum [Museum für Naturkunde Humboldt Universitet] (Germany, Berlin) NHRM - Swedish Natural History Museum [Naturhistoriska Riksmuseet] (Stockholm, Sweden) NIAST - National Institute of Agriculture Science and technology (Korea, Suwon) ZFMK - Zoological Institute and Museum Alexander Koenig [Zoologisches Forschungsinstitut und Museum Alexander König] (Germany, Bonn) ZIN - Zoological Institute Russian Academy of Sciences (St.-Petersburg, Russia) ZMHU - Zoological Museum Helsinki (Helsinki, Finland).

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Techinque for the genitalia preparation The method of study for delicate morphological structures, i.e. everting of vesica in males and inflating of bursa copulatrix in female are widely used by researchers of the Noctuidae. This approach permits morphologically separate and identify species in difficult taxonomic complexes. The function of these characters, their significance for taxonomic research and methodology of preparation are described in many publications (Hardwick, 1950; McCabe, 1980; Lafontaine, 1981; Lafontaine, Mikkola, 1987; Fibiger, 1997). Here, we briefly outline main points of the method of preparation of genitalia slides. 1. The abdomen gently is removed from the body and immerse to the tube containing 10% KOH and then left in drying shell with temperature 50-60� C, or the tube could be heated to nearly boiling pointing the water bath for 10-15 minutes. 2. After maceration or boiling in KOH, the soften abdomen is washed by distillated water, cleaned from scales or hairs and placing to preparates glass with water or weak solution (10%) of alcohol. The genital capsule of male or bursa copulatrix of female should be removed from the abdomen with help of preparates needles and fine forceps and clean from hairs or rest of muscles. 3. Then, with fine watch forceps, the aedeagus should be removed from genital armature, its ductus ejaculatorius should be cuted on 1-2 mm before aedeagus wall. The armature of genitalia should be transfered to the solution of higher concentration alcohol (30-40%, then 75% and 96%). 4. The vesica could be everted by stronger alcohol (70-90%) or very weak solution of Chlorazole Black in alcohol (1-2 drops of water solution of Chlorazole Black for 50 ml of 70-90% alcohol. In this process, the membrane of vesica will be gently stained. 5. The alcohol or alcohol solution of Chlorazole is injected into tube of aedeagus with help of syringe and fine hypodermic needle. The tip of needle gently injects into the rest part of the ductus ejaculatorius. The diameter of needle should be corresponding to diameter of aedeagus. If the vesica is concentrated in caudal part of the aedeagus, it is possible gently to move it forward by hard bristle. Then the aedeagus is fixed by watch forceps (for large species it is possible to fix it by fingers) and gently pushing the syringe. In most cases, the vesica easy everts. After this action, the aedeagus with everted vesica is deposited to stronger solution of alcohol (75%) and after 10-15 minutes of exposition to 99% or absolute alcohol where it should stay as minimum as 20-30 minutes for hardening and dehydratation. Similarly, after cleaning, the female genitalia are removed from the abdomen. Then, gently inject needle of syringe to antrum or to ductus bursa, and flood the bursa by alcohol. The everted of vesica or flooded bursa copulatrix should not be placed in water or in weaker solution of alcohol. After exposition of the aedeagus or bursa copulatrix in 75% of alcohol they are transferred into absolute alcohol at least for 5-10 minutes for dehydratation and hardening sufficiently for mounting. The preparates are mounted with euparal or Canada Balsam. If they are mounting with Canada Balsam, they should be exposed 5-7 minutes in the xylene. Because the vesica or bursa copulatrix often have volumetric shape, the small pieces of plastic are placed under the cover glass. Ready preparates should be labeled and dried in drying shell during several days and at least one month they should be kept horizontally. It is not recommended to keep preparates with everted vesica as a temporary preparates in alcohol/glycerin solution in open tubes, because the vesica easy loose its shape and such preparates could be damage or became dirty easily, 13

or lost. The permanent preparates are very convenient for study, transportation, and sorting, photographing, making drawings and so on. However, it is not always necessary to make permanent genitalia slide. For the aims of routine identification, it is enough to prepare usual preparates without evertation of vesica and keep it in the mixture solution of alcohol and glycerin. Very often, in large species, it is possible to examine genitalia by gentle brushing of the type of abdomen by brush, needle and fine forceps. It is possible to advance the male genitalia in fresh specimens by pressing of 8th segment of abdomen by forceps.

SYNOPSIS The present checklist enumerated 1,042 species of Korean Noctuidae including of 27 species of tribe Nolinae which are not treated in the Illustrated Catalogue of Noctuidae in Korea (1998), 42 taxa described or reported from Korea during the last decade by various authors, 16 species newly reported for Korea and four possible new species. Three synonyms for generic names and two synonyms for species names are proposed. Nine new taxonomic changes are introduced. The corrections for many taxa are given in notes. New species Condica sp. Micronoctua sp. Paragona sp. Lophomilia sp. New synonymies Dysgonia coreana (Leech, 1889), syn. n. Nycteola costalis Sugi, 1959 syn. n. Heraema Staudinger, 1892, syn. n. Prolophota Hampson, 1896, syn. n. Atuntsea Berio, 1977, syn. n. Taxonomic changes Leucomelas Hampson, 1913, stat. n. Euclidia juvenilis (Bremer, 1861) comb. n. Stenoloba glauca ssp. kuytekparki Sohn & Han, 2005, stat. n. Trachea mandschurica (Bremer, 1861), comb. n. Litoligia fodinae (Oberthür, 1880), comb. n. Aventiola trigonifera Hampson, 1896, comb. n. Lophomilia kogii (Sugi, 1977), comb. n. Euclidia juvenilis (Bremer, 1861) comb. n. Blasticorhinus unduligera, comb. n., syn. rev.

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Species newly reported for Korea Hypenodes rectifascia Sugi, 1982 Araeopteron fragmenta. Zanclognatha tenuialis Rebel Lygephila lilacina (Butler, 1878) Meganola major (Hampson, 1891) Meganola mediofascia (Inoue, 1958) Earias roseiviridis Sugi, 1982 Condica cyclicoides (Draudt, 1950) Nikara castanea Moore, 1881 Heliothis cruentata (Moore, 1881) Apamea crenata (Hufnagel, 1766) Acrapex klapperichi Kononenko, 1999. Acrapex owadai Kononenko, 1999 Sesamia confusa (Sugi, 1982) Mythimna albiradiosa (Eersmann, 1852) Cerastis rubricosa ([Den. & Schiff.], 1775) Species added from the literature sources Hypenodes rectifascia Sugi, 1982 Araeopteron flaccida Inoue, 1958 Eublemma cochylioides (Guenée, 1852) Mataeomera esbiahni Sohn & Ronkay, 2001 Hepatica nakatanii Sugi, 1982

Oglasa bifidalis (Leech, 1889) Hadennia obliqua (Wileman, 1911) Hydrillodes lentalis Guenée, 1854 Adrapsa simplex (Butler, 1879) Zanclognatha lui Han & Park, 2005 Anoratha paritalis (Walker, ([1859] 1858) Grammodes geometrica (Fabricius, 1775) Hypersypnoides submarginata (Walker, 1865) Calesia dasypterus (Kollar, 1844) Fodina contigua Wileman, 1914 Ericeia pertendens (Walker, 1858) Ischyja manlia (Cramer, 1776) Premusia intrahens Walker, 1858 Lophoptera squammigera Guenée, 1852 Nola cristatula (Hübner, 1793) Nola taeniata Snellen, 1875 Nola aerugula (Hübner, 1793) Nola yoshinensis (Wileman & West, 1929) Nola japonibia Strand, 1920 Nola confusalis (Herrich-Schäffer, [1847] 1845) Nola longicosta Oh, 2001 Nola nami (Inoue, 1956) Nola ebatoi Inoue, 1970 Nola innocua Butler, 1880 Nola okanoi (Inoue, 1958) Nola trilinea Marumo, 1923 Meganola microphasma (Butler, 1885) Meganola banghaasi (West, 1925) Meganola gigantoides (Inoue, 1961) Meganola mediofascia (Inoue, 1958) Meganola satoi (Inoue, 1970) Meganola basifascia (Inoue, 1958)

Meganola costalis (Staudinger, 1887) Meganola fumosa (Butler, 1879) Meganola shimekii Inoue, 1970 Meganola subgigas Inoue, 1982 Meganola albula ([Den. & Schiff., 1775) Meganola triangulalis Leech, 1888 [1889] Meganola gigas (Butler, 1884) Evonima mandschuriana (Oberthür, 1880) Poecilonola pulchella (Leech, 1889) Koyaga virescens (Sugi, 1958) Maliattha khasanica Zolot. & Dubatolov, 1996 Neustrotia albicincta (Hampson, 1898) Trisuloides rotundipennis Sugi, 1976 Cucullia hostilis Boursin, 1934 Valeria tricristata Draudt, 1934 Heliothis cruentata (Moore, 1881) Condica cyclicoides (Draudt, 1950) Nikara castanea Moore, 1881 Stenoloba glauca Kononenko & Ronkay, 2001 ssp. kuytekparki Sohn & Han, 2005 Spodoptera mauritia (Boisduval, 1833) Orthogonia basimacula (Draudt, 1939) Feliniopsis indistans (Guenée, 1852) Apamea crenata (Hufnagel, 1766) Apamea sodalis (Butler, 1878) Xylena nihonica Höne, 1917 Eupsilia boursini Sugi, 1958 Rhynchaglaea scitula (Butler, 1879) Orthosia fausta Leech, 1889 Orthosia nigromaculata (Höne, 1917) Orthosia satoi Sugi, 1960 Anarta stigmosa (Christoph, 1889)

GENITALIA MORPHOLOGY AND TERMINOLOGY It is well known that the genital structures play a very important role in the taxonomy and systematics of Lepidoptera. A great number of papers have been devoted to morphology, anatomy of the Lepidoptera genitalia and Noctuidae particularly. For students we recommend comprehensive taxonomist’s glossary of genitalia in insects, edited by Tuxen (1970). The main features of the genitalia of Noctuidae are described in revisions sited in the reference. Although we do not describe the genitalia of each taxon in this book, we include here the brief general review of the morphology

15

and terminology of Noctuidae genitalia and the methodology of the genitalia preparation. Male genitalia. (Fig. 1). Male genitalia of the Noctuidae are mainly symmetrical, rarely asymmetrical. The sclerites of 9-th segment of abdomen together with part derived from 10th segment form a transverse sclerotized ring, which serves for attaching the other parts of genitalia. Its dorsal part derived from 9th tergite calls vinculum and its ventral part, derived ftom 9th stenite is tegumen. In its apex tegumen joined with uncus - unpaired end-appendage of the 10-th segment. In most case, it has simply shape, however it could be modified: widened, flattened, or bearing additional extensions. In some groups of the Noctuidae uncus partially or completely reduced. Ventrally from top of the tegument and under uncus runs anal tube. Dorsal surface of the anal tube in some groups of the Noctuidae has sclerotization formed as gutter like sclerite bifid on the base scaphium. It could be sclerotized or membranous. In some genera of the Noctuidae, the anal tube is sclerotized on the ventral surface. The ventral sclerotization of the anal tube called subscaphuim. In many groups of so-called trifinae noctuidae, tegumen often has large lateral lobes (peniculus) covered with hairs or hair-like scales. Tegumen and vinculum are connected between each other by pleural sclerites which have shape of narrow curved or retorted sclerotized bands or flat plates. In the quadrifinae subfamilies of noctuidae, tegumen and vinculum are joined directly, pleural sclerites are reduced and fused with tegumen. Vinculum is folk-shape; its base sometimes has saccus proximally directed extension. Aedeagus fixed by membranes between tegument and vinculum. The aedeagus represents sclerotized tube, which caudally continues by membranous inner tube (endophallus, or vesica) invaginated and packed inside the aedeagus. Size and degree of curving of the aedeagus is variable. The shape of vesica is variable, it often has additional diverticulums. Its surface is often armed by cornuti - spines or field of spines of variable shape and size. During copulation, the vesica invaginated into ductus bursa of the female. The shape and size of vesica and cornuti have great significance for identification of difficult taxonomic complexes of the Noctuidae. The tube of aedeagus supported by diaphragm which formed by intersegmental membrane of 9th and 10th segments of abdomen. The diaphragm, except membrane includes fultura superior from dorsal side and fultura interior or juxta from ventral side. Sclerites of fulturasuperior are continuation of the costa of valva called transtilla. In some cases, they formed compact channel, sometimes they have small spines. The lower fultura of juxta is represented by sclerotized plate of different shape and sclerotization, most often clypeate or cordate. The plates of valvae are fixed on both sides of vinculum. They are paired, usually plate-like sclerites, flexible joined with vinculum on the base, having dorsal (or costal) margin, ventral margin and apex. On its base, valva has sacculus - sclerotized introversion of ventral margin of valve holding of large intervalvae muscle. The proximal end of sacculus often has clavus - sclerotized finger-like extension covered by rare finely hairs. In some genera, the upper part of sacculus has sclerotized extension directed distally. The central part of valva has ark-like or oblique central sclerite - clasper, which is the place of fixation of intervalvae muscle (Forbes, 1939). In many cases, especially typical for trifinae noctidae, it keeps in apical part more or less curved extension, called harpe. The skeletal base of valva is costa - sclerotized swelling of dorsal margin of valva. Sometimes this sclerite is separated from membrane of valva in distal part, often it has extension, called as digitus. The apex of valva is more or less separated shape, called as cucullus. Usually it is weakly sclerotized and covered with hairs or bristles, its distal part armed with spines called corona. In some cases, the surface of 16

cucullus is covered with spines or spine like bristles. In many groups, cucullus is well separated from other part of valva by more or less narrow neck, in some groups the neck is not expressed. The shape of cucullus is variable from complete reduction for various modifications. The presence and modification of cucullus are typical for trifinae Noctuidae, in so-called quadrifinae, it as a rule not expressed. Musculature of the male genitalia of the Noctuidae generally is more or less uniform and could be characterize by the presence of muscles m2 and fixation of muscles m4 to vinculum, sometimes they moved to the base of tegument. Muscles m1, as a rule directed to the base of uncus (Tikhomirov, 1979). Males of many species of trifinae subfamilies often have androconial apparatus formed by hair brushes fixed on special “lever” (coremata) and pockets on the base of abdomen, connected with scent glands. They usually dispose paired or rarely unpaired laterally on ventral side in 1-2 segment of abdomen. Some species have unpaired brush of hairs on 7-8 sternites, in some species the androconial apparatus as special hairs disposed on outer surface of valva. The paired androconial brushes are most typical for subfamilies Cuculliinae, Xyleninae and Hadeninae. In the quadrifinae subfamilies, the androconial apparatus can be found on the legs of males or on outer surface of valva; in some genera it is large membranous structures covered by hairs and fixed on outer part of the base of valva. Female genitalia (Fig. 2). The terminal 8th-10th segments of female’s abdomen are formed ovipositor. In most noctuidae species, 8th sternite is narrow, membranous, and 8th tergite is more developed and comprise most part of the segment, especially in species with telescopic ovipositor. Two pairs of sclerotized bands - apophyses anteriores and apophyses posteriores are connected with 8th tergite and papillae annales. They are the places of fixation of apophyses muscles ensuring movement of ovipositor. Papillae anales in different group have different size, shape and sclerotization. Their structures often have adaptive dependence from the character of ovipositing. The females of species, which lay eggs free on leaves or disperse them during the flight usually, have weakly developed ovipositor with short apophyses posteriores. On the other hands, the species which lie eggs into holds of trunk or branches, under bud scales, in leaf axils of poaceous plants, in corolla of flowers have long or telescopic ovipositor with extended intersegmental membrane and very long apophises posterior (2-3 times logger then apophises anterior). The species, lying eggs inside plants steams, have very sclerotizeted papillae anales adopted for breaking of cauline tissues. The female genitalia open by copulative opening or ostium. The shape and position of ostium are variable depositing in the limits of 7th-8th sternites. Its opening lies on the surface of sternite or drowned into vaginal sinus. Postvaginal plate lies caudal of ostium and antevaginal plate lies in front of it. Ostuim opens to antrum - sclerotized part of duct of copulative bursa. In some cases, the antrum is weakly sclerotized, not expressed or has ribbed structure. Antrum transforms to ductus bursa, which could be often sclerotized, sometimes has ribbed structure. Copulative bursa (corpus bursae or bursae copulatrix) generally looks as a sack, often its surface armed with signums sclerotized bands or plaques. Often bursa has sclerotization in caudal part or extension or appendix which is sclerotized as a rule and sometimes significantly larger than bursa. Ductus seminalis falls into bursa copulatrix or appendix bursa.

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Fig. 1. Structure of male genitalia. (After Kononenko, 2003). 1, 3-genitalia armature; 2, 4-aedeagus (1, 2-Ochropleura plecta; 3, 4Catocala pirata); 5-tegument with peniculus; 6-10-modifications of uncus; 11-13-modifications of juxta; 14-17-modifications of valva.

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Fig. 2. Structure of female genitalia. (After Kononenko, 2003). 1-Pangrapta costaemacula; 3-Belciades niveola; 3-Cirrhia icteritia; 4-Agrotis trrifurca; 5-Nonagria typhae.

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CLASSIFICATION The earlier classifications of Noctuoidae, based mainly on Hampson’s (1903?1913) classical work, have become outdated and undergoing radical revisions and modifications during last decade. Most Hampsonian noctuid subfamilies are recognized as unnatural (Kitching, 1984). The classification of higher taxa of the family Noctuidae (s. l.) since Hampson has undergone with dramatic changes. Here we briefly outline the main changes of the classification of higher taxa of the Noctuidae in most recent publications. During the last 10 years, the classification of Noctuoidea, particularly Noctuidae, has been a subject for revisions based on adult morphology, study of genitalia musculature as well as morphology of immature stages. Various systems of the Noctuidae have been proposed for North American (Poole, 1995), Australian (Nielsen, 1996), European (Fibiger and Hacker, 1991; Beck, 1996, 1999?2000; Nowacki and Fibiger, 1998, after Rawlins and Kitching, 1999; Leraut, 1997), and World faunas of the whole family (Speidel et al., 1996; Rawlins and Kitching, 1999). Recently the molecular study for Noctuidae classification and the construction of the phylogeny of the family have been attempted (Weller et al., 1994; Mitchel et al., 1997, 2000, 2006). The systems proposed by Speidel et al. (1996) and those by Kitching and Rawlins (1999) were most comprehensive and well grounded among the recently published classifications of the higher taxa of the Noctuoidea at the end of the last century. The main features of above-mentioned classifications have been considered and analyzed by Lafontaine and Fibiger (2006). However, as the proposed classifications are based mainly on European, North American or Australian faunas, many taxa of East Asian origin and especially tropical taxa were not included there, by this reason the placement of some groups in these systems are still uncertain. As the immature stages of many taxa are still unknown, the application of the data on morphology of immature stages for classification of the Noctuidae is rather limited. The classification proposed by Beck (1996, 1999-2000), developed for European fauna and based mainly exclusively on larval characters has not been accepted by Noctuidae researchers. The using molecular study (Mitchel et al., 1997, 2000, 2006) is very progressive approach to developing of Noctuidae classification, but also is still limited by the reason of small number of examined taxa. Very important step for clarification of the nomenclature and the development classification of the high taxa of the Noctuoidae of Holarctic fauna has been undergone with survey of nomenclature of family-group names of the Noctuoidea (Speidel, Naumann, 2005), and a revision of the classification of higher taxa of the Noctuoidea (Fibiger and Lafontaine, 2005) and its latest modification (Lafontaine, Fibiger, 2006). The last two publications are grounded on syntheses of extensive analysis of imago and immature morphology as well as on molecular classification (Mitchel, et al., 2006). According to Fibiger and Lafontaine (2005), the family Noctuidae (s. l.) has been divided for three families: Erebidae, Micronoctuidae and Noctuidae. The Arctiid group of families (Arctiidae, Nolidae and Lymantriidae was placed in front of the upgraded family Erebidae, the quadrifinae subfamilies 20

(except Plusinae) were treated as subfamilies of Erebidae. The subfamilies Rivulinae, Boletobinae, Phytometrinae, Scoleocampinae, Erebinae, and Hypenodinae were reinstated. A new family Micronoctuidae was proposed. The generic and tribal compositions of many subfamilies were revised. The subfamily Araeopteronidae was proposed as a new subfamily in Erebidae. In contrast with Kitching and Rawlins (1999), the tribe Eublemini was removed from Eustrotiinae and raised for the subfamily of Erebidae. The Pantheinae was recognized as a subfamily of the Noctuidae. The classification of artificial assemblage Hadeninae (sensu Kitching and Rawlins 1999) has been revised, the classification of Noctuinae has been reviewed and many other taxonomic changes were proposed. Many genera of Amphipyrinae (sensu auctorum) or Hadeninae (sensu Kitching and Rawlins, 1999) were placed to the subfamily Xyleninae and divided for tribes and subtribes. A revised checklist of European Noctuoidea by Fibiger and Hacker (2005) has been completed based on a new classification of the Noctuoidea. The next step of developing of the Noctuidae classification was the publication by Lafontaine and Fibiger (2006), modifying the former classification (Fibiger and Lafontaine, 2005) as shown in [Table 1]. Micronoctuidae retained as a family of uncertain affinity in the Noctuoidea. Arctiidae and Lymantridae were proposed to treat as subfamilies of the sister group of erebine-calpine-cocytine lineage. Aganainae treated a sister group of Arctiinae. Nolidae placed into quadrifinae Noctuidae and treated as a subfamily, and Nolinae (s. str., i.e. genera Nola, Meganola, Evonima, ets.) as a tribe. Stictopterinae and Euteliinae are treated as a sister group of Erebinae - Lymantriinae lineage. Subtribe Arzamina was moved from Apameini and treated as s tribe of Xylenini. Upheinae was removed to Xylenini as a subtribe. The tribe Cosmiini (sensu Fibiger and Lafontaine, 2005) was treated as a subtribe Cosmiina in the Xylenini. The tribe Balsini was removed from Xylenini and treated as subfamily placed between Raphiinae and Acronictinae. Tribe Phosphilini was removed from the Psaphidinae to Xyleninae. The cladogram of classification and the list of higher taxa with some above-mentioned changes and additions of 11 names from Holloway’s revision of Borneo Catocalinae (Holloway, 2005) have been presented. This is the attempt to integrate the East Palaearctic (Manchurian) and some Oriental taxa to classification of the Holarctic taxa; therefore the taxonomic position of some genera is still provisional or tentative. For the sequence of genera within recently revised subfamilies or generic groups (i.e. Herminiinae, Plusiinae, part of “Acontiinae”, Acronictinae, part of Cuculliinae, Noctuinae), we follow selected revisions by various authors. Tribal subdivisions are followed according to Fibiger and Lafontaine (2005, 2006) and Fibiger and Hacker (2005). The short comments to position of some East Asian subfamilies and genera in a new classification of the Noctuidae (s. l.) are given below.

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Table 1. Recently proposed classifications of the Noctuidae (s. l.) (Only subfamily names listed. The subfamilies not represented in Korea are given in square brackets). Kononenko et al., 1998 (Korea)

Speidel et al., 1996 (World)

NOCTUIDAE HERMINIINAE RIVULINAE HYPENODINAE HYPENINAE CATOCALINAE PLUSIINAE EUTELIINAE SARROTHRIPINAE COLLOMELINAE NOLINAE ACONTIINAE PANTHEINAE ACRONICTINAE AGARISTINAE AMPHIPYRINAE CUCULLIINAE HADENINAE NOCTUINAE HELIOTHINAE

LYMANTRIIDAE AGANAIDAE ARCTIIDAE NOCTUIDAE HERMINIINAE RIVULINAE HYPENODINAE HYPENINAE EUTELIINAE STICTOPTERINAE CATOCALINAE PANTHEINAE CAMPTOLOMINAE CHLOEPHORINAE SARROTHRIPINAE NOLINAE EUSTROTIINAE BAGISARINAE PLUSIINAE ACONTIINAE [TYTINAE] AEDINAE [STIRIINAE] ACRONICTINAE BRYOPHILINAE SINOCHARINAE [LOPHONYCTINAE] PSEUDEUSTROTINAE AGARISTINAE HELIOTHINAE NOCTUINAE CUCULLIINAE HADENINAE AMPHIPYRINAE

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Kitching & Rawlins, 1999 (World)

Fibiger & Lafontaine, 2005 (Holarctic and World)

Lafontaine & Fibiger, 2006 (Holarctic and World)

NOCTUIDAE ACRONICTINAE BRYOPHILINAE RAPHIINAE HERMINIINAE [STREPSIMANINAE] CATOCALINAE CALPINAE [AGANAINAE] HYPENINAE STICTOPTERINAE EUTELIINAE PLUSIINAE AGARISTINAE [EUCOCYTIINAE] ACONTIINAE EUSTROTIINAE BAGISARINAE CUCULLIINAE AMPHIPYRINAE PSAPHIDINAE DILOBINAE CONDICINAE [STIRIINAE] HELIOTHINAE [GLOTTULINAE] HADENINAE [UFEINAE] NOCTUINAE NOLIDAE NOLINAE CHLOEPHORINAE EARIADINAE [WESTERMANIINAE] EARIDINAE BLENINAE RISOBINAE [COLLOMELINAE] [AFRIDINAE] ELIGMINAE ARCTIIDAE

NOLIDAE NOLINAE CHLOEPHORINAE WESTERMANNIINAE EARIADINAE BLENINAE RISOBINAE COLLOMENINAE AFRIDINAE ELIGMINAE ARCTIIDAE LYMANTRIIDAE EREBIDAE RIVULINAE [BOLETOBINAE] HYPENODINAE ARAEOPTERONINAE EUBLEMMINAE HERMINIINAE HYPENINAE PHYTOMETRINAE SCOLECOCAMPINAE AVENTIINAE EREBINAE CALPINAE CATOCALINAE [AGANAINAE ] COCYTIINAE STICTOPTERINAE EUTELIINAE MICRONOCTUIDAE NOCTUIDAE PLUSIINAE EUSTROTIINAE BAGISARINAE ACONTIINAE PANTHEINAE DIPHTHERINAE [DILOBINAE] ACRONICTINAE RAPHIINAE METOPONIINAE SINOCHARINAE [LOPHONYCTINAE] AGARISTINAE CUCULLIINAE ONCOCNEMIDINAE AMPHIPYRINAE PSAPHIDINAE [STIRIINAE] HELIOTHINAE CONDICINAE ERIOPINAE BRYOPHILINAE XYLENINAE HADENINAE NOCTUINAE

NOCTUIDAE RIVULINAE BOLETOBIINAE HYPENODINAE EUBLEMMINAE HERMINIINAE HYPENINAE PHYTOMETRINAE AVENTIINAE SCOLECOCAMPINAE EREBINAE CALPINAE CATOCALINAE COCYTIINAE [STREPSIMANINAE] NOLINAE [AGANAINAE] ARCTIINAE LYMANTRIINAE STICTOPTERINAE EUTELIINAE PLUSIINAE EUSTROTIINAE BAGISARINAE ACONTIINAE PANTHEINAE DILOBINAE RAPHIINAE ACRONICTINAE BALSINAE METOPONIINAE SINOCHARINAE [LOPHONYCTINAE] AGARISTINAE [EUCOCYTIINAE] CUCULLIINAE ONCOCNEMIDINAE AMPHIPYRINAE PSAPHIDINAE [STIRIINAE] HELIOTHINAE CONDICINAE ERIOPINAE BRYOPHILINAE XYLENINAE HADENINAE NOCTUINAE

Comments to the families and subfamilies Micronoctuidae. The Micronoctuidae as a family is reported for Korea for the first time. It contains two genera, Mimachrostia and Micronoctua, with one species in each. Both species are known from the South of the Russian Far East. The family is under revision now by Fibiger (pers. comm.); publications are pending in short time. Rivuliinae. Small, unrevised subfamily with some species and genera of uncertain position. We tentatively place the genus Hemipsestra here. Hypenodinae. The subfamily name is not synonymous with Strepsimaniinae, which is considered as a distinct family of Noctuoidea (Fibiger and Lafontaine, 2005). Except the genera Hypenodes and Schrankia, we placed the genera Micreremites and Anachrostis. Eublemminae The subfamily in Korean fauna includes two tribes Eublemmini and Pangraptini, the later is restricted by the genus Pangrapta, but we added the genus Naganoela here. The East Asian genera Enispa, Corgatha, Oruza, Hyposada, Lophoruza, Tamba, Corsa, Mataeomera, Paragona, Anatatha, Diomea and Maguda are placed to Eublemmini by affinity of the external and genitalia characters to Eublemma, the type genus of the subfamily. The genus Trisateles was correctly transferred to Eublemminae (Acontiinae auct.) by Sugi (1982), however in European publications it often treated in Herminiinae (Fibiger and Hacker, 2005). To the group of miscellaneous genera, we placed Ectogonia, Stenograpta, Amphitrogia, Lophomilia, Paragabara, Hepatica, Gonepatica, Hypostrotia, Scedopla, Leiostola, Oglasa, and Holocryptis. Their systematic positions are still uncertain and could be resolved in further revisions. Probably some genera will be removed to other subfamilies. The genus Naranga is transferred to Eustrotiinae. Herminiinae. The taxon is treated as a subfamily of the Noctuidae, while in earlier publications (Kitching, 1984; Nielsen, 1996) it was considered as a distinct family. In the treatment of the subfamily and sequence of genera, we follow to revisions of the Herminiinae by Owada (1987, 1992, 2006) as most comprehensive publications on Herminiinae of East Asian fauna. The order of genera is different from those accepted for the European Checklist of Noctuoidea (Fibiger and Hacker, 2005). The genus Gynaephila is considered here a member of Herminiinae following Lödl (1998). Hypeninae. Most genera and species of East Asian fauna are still unrevised. For the order of genera and species, we follow to Sugi (1982). The genus Stengbergmania is transferred from Catocalinae to Hypeninae following Poole (1989). The genera Anorata, Rhynchina and Zekelita are revised by Lödl and Gaal (1998), Lödl and Mayerl (1998), and Mayerl and Lödl (1999). The genus Itmacharela is placed to Hypeninae by Kononenko et al., 1989. Phytometrinae. The subfamily contains genera Phytometra, Rhesala, Raparna and Colobochyla. We tentatively placed Euwilemania here. Aventiinae. Except the genus Laspeyria, we placed here the genus Sophta by their affinity of the genitalia characters, especially in females. Erebidae. It was treated as a distinct family with subfamilies Rivulinae, Boletobinae, Araepteroninae, Hypenodinae, Eublemminae, Herminiinae, Hypeninae, Phytometrinae, Aventiinae, Erebinae, Calpinae and Catocalinae by Fibiger and Lafontaine (2005) and in subsequent faunal publications (Hacker and Fibiger, 2005; Kononenko, 2005). Presently the taxon is treated as a subfamily with tribes Erebini, Hypopyrini and Arcteini in the regional fauna. The position of the last tribe is uncertain. 23

Catocalinae. Following Fibiger and Lafontaine (2005), Lafontaine and Fibiger (2006), the subfamily includes tribes; Toxocampini, Arytrurini, Sypnini, Hypocalini, Euclidini, Ophiusini, Catocalini, Sypnini, Tinolini, Hulodini, Pandesmini, and Ercheini for the Korean fauna. The East Asian tribe Sypnini (with genera Hypersypnoides, Sypnoides, and Daddala in the regional fauna) was not listed by Fibiger and Lafontaine, its description and validation of “Phillum Sypna” sensu Berio for this generic group done by Holloway (2005). The genera of tropical origin Erygia, Erygia, Blasticorhinus, Hypospila, Premusia, Mecodina, and Hyposemansis are listed as miscallaneous (sensu Holloway, 2005). The generic name Bastilla instead Parallelia (sensu auctorum) and Dysgonia (part) (tribe Ophiusini) introduced according to revision by Holloway and Miller (2003). Stictopterinae. First reported as a new family for Korean fauna by Sohn, Ronkay and Choi (2005) Nolidae. The status and of this taxon has been debated for a long time. It was considered as a distinct family of the Noctuoidea or as a subfamily of the Noctuidae by previous authors. The Sarrothripinae, Chloephorinae and Nolinae are recognized as members of Nolidae by Holloway (1998). Kitching and Rawlins (1999) and Fibiger and Lafontaine (2005) treated Nolidae as a distinct family. In the latest publication (Lafontaine, Fibiger, 2006), Nolinae was treated as a subfamily of the Noctuidae. The subfamily contains tribes Nolini, Chloephorini, Eariadini, Blenini, Risobini and Eligmini in the regional fauna. The Nolinae (Nolini) was not included in the Illustrated Catalogue of Korean Noctuidae (Kononenko et al., 1998), while Sarrothripinae & Chloephorinae were included as a subfamilies. The Korean fauna of the tribe Nolini has been revised by Oh (1991, 2001). Arctiinae and Lymantriinae. Two these subfamilies were not treated in the Illustrated Catalogue of Korean Noctuidae, therefore they are not included in the present Checklist. Plusiinae. The Palaearctic and Nearctic fauna of the subfamily were revised by Kostrowicky (1961), Ichinose (1973), Eischling and Cunningham (1978), Kitching (1987), and Lafontaine and Poole (1991), the European fauna revised by .L. Ronkay (in Goater et al, 2003). In the treatment and sequence of genera in the subfamily, we follow the checklist of the Plusinnae of World fauna given by L. Ronkay (in Goater et al, 2003). The tribes Abrostolini, Argyrogrammatini, Plusiini are included. Eustrotiinae. The classification of the Deltote generic group, the core of Eustrotiinae, is accepted according to the revision by Ueda (1984, 1986). The genus Naranga is placed in the Eustrotiinae. The genus Anterastria was transferred to tribe Pseudeustrotiinae, subfamily Xyleninae. The group of miscellaneous genera includes Neustrotia, Bryophilina, Microxyla, Eulocastra, Ozarba, Amyna, and “Hyperstrotia”. Their systematic position, especially two last genera are uncertain, they are connected neither with Eustrotiinae nor Eublemminae or Acontiinae. Temporary, before further revision we retain these genera in Eustrotiinae. Bagisarinae. This small, recently recognized subfamily includes the genera Xanthodes, Brevipecten and Chasminia in the Oriental region. The genus Imosca was provisionally placed in Bagisarini by Holloway (1989) and confirmed by Sugi and Sasaki, (2001). The genus Sphragifera is considered here as a member of Bagisarinae. The Bagisarinae, has trifinae type of venation and semi-looper larvae, adults are characterized by specialized structure of male eight tergit and sternit (Sugi and Sasaki, 2001; S. Sugi, pers. comm.). Acontiinae. The subfamily contains two tribes, Acontiini and Aediini. The later was transferred from Catocalinae to Acontiinae by Fibiger and Lafontaine (2005). 24

Pantheinae. Despite the opinion of Kitching and Rawlins (1999), following Speidel et al., (1996) and Fibiger and Lafontaine (2005, 2006), the taxon is given in subfamily rank. Raphiinae. The taxon was upgraded to the subfamily rank by Kitching and Rawlins (1999). Balsinae. The taxon was reinstalled first as a tribe of Xyleninae and then as a subfamily by Fibiger and Lafontaine (2005) and Lafontaine and Fibiger (2006). Acronictinae. The order of genera is arranged according to Kozhantschikov (1950) and Sugi (1982) with some modifications. The composition of the genus Belciana is provisional. Two regional species of this genus are not congeneric with Belciana biformis, the type species of the genus . Agaristinae. Despite to the opinion of earlier authors who considered the taxon as a family rank, Agaristinae is well recognized as a subfamily of Noctuidae. In the treatment and sequence of genera of the subfamily, we follow to the revision of Palaearctic and Oriental faunas of Agaristidae by Kiriakoff (1977). Cuculliinae. The classification of the Cuculliinae and the list of the genus Cucullia particularly are accepted according to the revision by G. Ronkay and L. Ronkay (1995). Tribes Oncocnemidini, Psaphidini and Feralini formerly considered in Cuculliinae were upgraded to subfamilies Oncocnemidinae and Psaphidinae by Fibiger and Lafontaine (2005). Heliothinae. In treatment and sequence of genera of the subfamily, we follow the World revision of Heliothinae by Matthews (1991). Despite to the opinion of Matthews, the genus Protoschinia is considered as a distinct genus, not a synonym of Schinia. Condicinae. Except the genera listed in the European list (Fibiger and Hacker, 2005), we included to the Condicinae East Asian genera Prospalta, Chytonix, Niphonyx, Oligonyx, Nikara, Pyrrhidivalva, Dysmilichia, Iambia, Plusilia and Prometopus. Two Manchurian species “Hadjina” sinensis and “H.” biguttula are transferred to the genus Acosmetia. Bryophilinae. Following to Fibiger and Hacker (2005), we treated the Bryophilinae as a subfamily separating from Acronictinae. The subfamily, except genera recorded from Europe, includes the genera Bryomoia, Stenoloba and Athaumasta; the former has been transferred from Cuculliinae (sensu Hampson) to Acronictinae [+Bryophilinae] by Kononenko et al. (1998). The genus Cryphia with subgenera Cryphia and Bryoleuca (sensu Nowacky and Fibiger, 1998 and Fibiger and Hacker, 2005) is treated as two different genera: Cryphia and Bryoleuca. Xyleninae. The genus Anterastria is placed in the tribe Pseudeustrotiini; the sole species of the genus has trifinae type of hind wing venation and coremata, which are absent in other members of Eustrotiinae. Its male genitalia similar to those of Pseudeustrotia. The “Xanthograpta” basinigra is transferred from the Acontiinae (sensu auctorum) to the tribe Apameini according the genitalia structures. However, the affinity of basinigra to Xanthograpta should be revised. The genera Bambusiphila and Chandata are placed to Apameini, subtribe Apameina. The genus Virgo is placed in the Apameini, subtribe Sesamiina. The genus Heraema is recognized as a synonym of Trachea. The genera Mormo, Orthogonia, Olivenebula, Xenotrachea, Karana and Auchmis and Triphaenopsis are placed in the tribe Dypterygini; their systematic position will be clarified under further revision. The genera Pygopteryx is transferred to the tribe Xylenini. The genus Caradrina is treated according to revision by Hacker (2004). Hadeninae. The subfamily includes 4 tribes: Orthosiini, Tholerini, Hadenini, Leucanini and Eriopygini. The order of tribes and sequence of genera is followed according to Fibiger and Hacker 25

(2005) and Hacker et al. (2002) with some modifications. The genera Anarta (Hadula auct.) and Hadena are treated according to revisions by Hacker (1998b, 1999). The classification of Orthosiini is accepted according to Ronkay et al. (2001), Fibiger and Hacker (2005). In the classification of the genus Mythimna, we followed to Yoshimatsu (1994) and Hacker et al., 2002. Noctuinae. The subfamily includes two tribes, Noctuini and Agrotini. In treatment of the subfamily, we followed revisions of the European Noctuinae by Fibiger (1990; 1993; 1997), North American Noctuinae by Lafontaine (1987; 1998, 2004) and other recent publications. The sequence of genera are accepted according to Fibiger, Lafontaine (2005) and the Checklist of European Noctuoidae (Fibiger, Hacker, 2005). The genera Actebia and Dichagyris are accepted according to Lafontaine (2004) and Fibiger and Lafontaine (2005).

26

Table 2. Taxonomic composition and diversity Noctuoidea in Korea Taxon name Superfamily NOCTUOIDEA Latreille, 1809 Family NOTODONTIDAE Stephens, 1829 Family MICRONOCTUIDAE Fibiger, 2005 Family NOCTUIDAE Latreille, 1809 Subfamily RIVULINAE Grote, 1895 Subfamily HYPENODINAE Forbes, 1954 Subfamily ARAEOPTERONINAE Fibiger, 2005 Subfamily EUBLEMMINAE Forbes, 1954 Tribe EUBLEMMINI Forbes, 1954 Tribe PANGRAPTINI Grote, 1882 Miscellaneous genera Subfamily HERMINIINAE Leach, [1815] Subfamily HYPENINAE Herrich-Schäffer, [1851 Subfamily PHYTOMETRINAE Hampson, 1913 Subfamily AVENTIINAE Tutt, 1896 Subfamily EREBINAE Leach, [1815] Tribe EREBINI Leach, [1815] Tribe HYPOPYRINI Guenée, 1852 Tribe ARCTEINI Berio, 1992 Subfamily CALPINAE Boisduval, 1840 Tribe ANOMINI Grote, 1882 Tribe CALPINI Boisduval, 1840 Tribe SCOLIOPTERYGINI Herrich-Schäffer, [1852] Subfamily CATOCALINAE Boisduval, [1828] Tribe TOXOCAMPINI Guenée, 1852 Tribe ARYTRURINI Fibiger and Lafontaine, 2005 Tribe EUCLIDIINI Guenée, 1852 Tribe OPHIUSINI Guenée, 1837 Tribe CATOCALINI Boisduval, [1828] Tribe SYPNINI Holloway, 2005 Tribe HYPOCALINI Guenée, 1852 Tribe TINOLINI Moore, [1885] Tribe HULODINI Guenée, 1852 Tribe PANDESMINI Wiltshire, 1990 Tribe ERCHEINI Berio, 1992 Miscellaneous genera Subfamily STICTOPTERINAE Hampson, 1894 Subfamily EUTELIINAE Grote, 1882 Subfamily NOLINAE Bruand, 1846 Tribe NOLINI Bruand, 1846 Tribe CHLOEPHORINI Stainton, 1859 Subtribe CHLOEPHORINA Stainton, 1859 Subtribe SARROTHRIPINA Hampson, 1894 Subtribe CAMPTOLOMINA Mell, 1943 Subtribe ARIOLICINAE Mell, 1943 Tribe EARIADINI Hampson, 1912 Tribe BLENINI Mell, 1943 Tribe RISOBINI Mell, 1943 Tribe ELIGMINI Mell, 1943 Subfamily ARCTIINAE Leach, [1815] Subfamily LYMANTRIINAE Hampson, [1893] Subfamily PLUSIINAE Boisduval, [1828] Tribe ABROSTOLINI Eichlin and Cunningham, 1978 Tribe ARGYROGRAMMATINI Eichlin and Cunningham, 1978 Tribe PLUSIINI Boisduval, [1828] Subtribe AUTOPLUSIINA Kitching, 1987

Number of genera

Number of species

2

2 (1039) 7 9 4 (75) 37 16 22 60 33 5 3 (7) 5 1 1 (15) 5 9 1 (106) 16 2 7 19 36 5 4 2 1 1 1 12 1 5 (31)

2 4 1 (31) 15 2 14 18 10 5 2 (5) 3 1 1 (5) 1 3 1 (38) 5 1 5 9 2 3 1 2 1 1 1 7 1 3 (29) 4 5 9 2 1 3 1 1 2 1 ? ? (21) 1 7 13 5

22 14 4 1 3 4 1 3 1 ? ? (48) 5 10 34 16

27

Table 2. Contiuned. Taxon name Subtribe EUCHACIINA Chou and Lu, 1979 Subtribe PLUSIINA Boisduval, [1828] Subfamily EUSTROTIINAE Grote, 1882 Miscellaneous genera Subfamily BAGISARINAE Crumb, 1956 Subfamily ACONTIINAE Guenée, 1841 Tribe ACONTIINI Guenée, 1841 Tribe AEDIINI Beck, 1960 Subfamily PANTHEINAE Smith, 1898 Subfamily RAPHIINAE Beck, 1996 Subfamily BALSINAE Grote, 1896 Subfamily ACRONICTINAE Heinemann, 1859 Subfamily METOPONIINAE Herrich-Schaffer, [1851] Subfamily SINOCHARINAE Speidel, Fänger, and Naumann, 1996 Subfamily AGARISTINAE Herrich-Schäffer, [1858] Subfamily CUCULLIINAE Herrich-Schäffer, [1850] Subfamily ONCOCNEMIDINAE Forbes and Franclemont, 1954 Subfamily AMPHIPYRINAE Guenée, 1837 Subfamily PSAPHIDINAE Grote, 1896 Tribe PSAPHIDINI Grote, 1896 Tribe FERALIINI Poole, 1995 Subfamily HELIOTHINAE Boisduval, [1828] Subfamily CONDICINAE Poole, 1995 Subfamily ERIOPINAE Herrich-Schäffer, [1851] Subfamily BRYOPHILINAE Guenée, 1852 Subfamily XYLENINAE Guenée, 1837 Tribe PSEUDEUSTROTIINI Beck, 1996 Tribe PRODENIINI Forbes, 1954 Tribe ELAPHRIINI Beck, 1996 Tribe CARADRININI Boisduval, 1840 Subtribe CARADRININA Boisduval, 1840 Subtribe ATHETINA Fibiger and Lafontaine, 2005 Tribe DYPTERYGIINI Forbes, 1954 Tribe ACTINOTIINI Beck, 1996 Tribe PHLOGOPHORINI Hampson, 1918 Tribe APAMEINI Guenée, 1841 Subtribe OXYTRIPINA Goznamy, 1970 Subtribe APAMEINA Guenée, 1841 Subtribe SESAMIINA Fibiger and Goldstein, 2005 Tribe XYLENINI Guenée, 1837 Subtribe XYLENINA Guenée, 1837 Subtribe COSMIINA Guenée, 1852 Subtribe ANTITYPINA Forbes and Franclemont, 1954 Subfamily HADENINAE Guenée, 1837 Tribe ORTHOSIINI Guenée, 1837 Tribe HADENINI Guenée, 1837 Tribe LEUCANIINI Guenée, 1837 Tribe GLOTTULINI Guenée, 1852 Subfamily NOCTUINAE Rambur, 1848 Tribe AGROTINI Rambur, 1848 Subtribe AUSTRANDESIINA Angulo and Olivares, 1990 Subtribe AGROTINA Rambur, 1848 Tribe NOCTUINI Latreille, 1809 Subtribe AXYLIINA Fibiger and Lafontaine, 2005 Subtribe NOCTUINA Latreille, 1809

28

Number of genera

Number of species

5 3 9 7 4 (3) 2 1 7 1 1 14 1 1 3 1 3 1 (4) 3 1 6 13 1 5 (95) 3 1 1 (4) 3 1 11 2 5 (32) 1 29 2 (36) 22 9 5 (28) 8 15 4 1 (23) 5 1 4 18 2 16

7 11 28 13 6 (6) 5 1 9 1 1 46 1 1 4 15 3 8 (9) 8 1 11 25 9 18 (205 ) 3 6 1 (18) 6 12 20 3 7 (57) 1 49 5 (92) 54 29 9 (94) 27 33 32 1 (73) 20 1 19 53 2 51

SYSTEMATIC LIST Superfamily NOCTUOIDEA Latreille, 1809 Family MICRONOCTUIDAE Fibiger, 2005 Genus MIMACHROSTIA Sugi, 1982 1. Mimachrostia fasciata Sugi, 1982 � genit.: Pl. 1 fig. 1; ♀ genit.: Pl. 177, fig. 1 Ill. Cat. Noctuidae Korea, 1998: 114; No. 283.

Genus MICRONOCTUA Fibiger, 1997 (Type species: Micronoctua karsholti Fibiger, 1997 [Greece]). 2. Micronoctua sp.* � genit.: Pl. 1, fig. 2; ♀ genit.: Pl. 177, fig. 2 Ill. Cat. Noctuidae Korea, 1998 _____.

DISTRIBUTION. Korea, Japan, Taiwan, China, RFE (Primorye). NOTE. An undescribed species, distributed in Far Easter countries. The description of the species is pending (Fibiger & Kononenko, in press.).

Family NOCTUIDAE Latreille, 1809 Subfamily RIVULINAE Grote, 1859 Genus RIVULA Guenée, 1844 3. Rivula sericealis (Scopoli, 1763) � genit.: Pl. 1, fig. 3; ♀ genit.: Pl. 177, fig. 3 Ill. Cat. Noctuidae Korea, 1998: 41; No. 57.

NOTE. The name Rivula sericealis ssp. tanitalis Rebel, 1912 should be exclude from the synonymy to R. sericealis. According to Hacker (pers. comm.), Rivula tanitalis is a distinct species. 4. Rivula unctalis Staudinger, 1892

� genit.: Pl. 1, fig. 4

Ill. Cat. Noctuidae Korea, 1998: 42; No. 59.

NOTE. The male genitalia of the holotype is illustrated here. The female was unavailable for dissection. 5. Rivula curvifera Walker, 1862

� genit.: Pl. 1, fig. 5; ♀ genit.: Pl. 177, fig. 4


6. Rivula inconspicua (Butler, 1881)



7. Rivula leucanoides (Walker, [1863] 1864) Ill. Cat. Noctuidae Korea, 1998: 42; No. 60.

29

NOTE. The specimens were unavailable for dissection during preparation of the book. ♀ genit.: Pl. 177, fig. 6

8. Rivula errabunda Wileman, 1911 Ill. Cat. Noctuidae Korea, 1998: 43; No. 62.

NOTE. The male specimen was unavailable for dissection during preparation of the book. Genus HEMIPSECTRA Hampson, 1891 9. Hemipsectra fallax (Butler, 1789) � genit.: Pl. 2, fig. 1; ♀ genit.: Pl. 177, fig. 7 Ill. Cat. Noctuidae Korea, 1998: 113; No. 278.

NOTE. The genus is tentatively placed to Rivilinae.

Subfamily HYPENODINAE FORBES, 1954 Genus HYPENODES Doubleday, 1850 10. Hypenodes humidalis Doubleday, 1850 � genit.: Pl. 2, fig. 2; ♀ genit.: Pl. 177, fig. 8 Ill. Cat. Noctuidae Korea, 1998: 44; No. 66.

11. Hypenodes curvilinea Sugi, 1982



12. Hypenodes rectifascia Sugi, 1982*


Moths of Japan, 1: 899, Pl. 220: 21, (TL: Japan, Honshu, Saitama Pref., Iruma, Bushi [NIAES]) Ill. Cat. Noctuidae Korea, 1998. See No. 67 (Hypenodes curvilinea)

DISTRIBUTION. S Korea, Japan (Honshu). NOTE. The species was not included in the Catalogue of Korean Noctuidae, althouth it was dissected in 1995 by Kononenko. Hereby we report it for Korea for the first time. Material examined: 1 male, South Korea, genitalia preparate 30/95 VK (NIAST). Genus MICREREMITES Warren, 1891 13. Micreremites pyraloides Sugi, 1982 � genit.: Pl. 2, fig. 5; ♀ genit.: Pl. 178, fig. 1 Ill. Cat. Noctuidae Korea, 1998: 113; No. 280.

NOTE. The genus is placed to Hypenodinae. Genus SCHRANKIA Hübner, [1825] 1816 14. Schrankia costaestrigalis (Stephens, 1834) � genit.: Pl. 2, fig. 6; ♀ genit.: Pl. 178, fig. 2 Ill. Cat. Noctuidae Korea, 1998: 43; No. 63.

15. Schrankia separatalis (Herz, 1904)



16. Schrankia kogii Inoue, 1979 Ill. Cat. Noctuidae Korea, 1998: 44; No. 65.

30


Genus ANACHROSTIS Hampson, 1893 17. Anachrostis nigripunctalis (Wileman, 1911) � genit.: Pl. 3, fig. 1; ♀ genit.: Pl. 178, fig. 5 Ill. Cat. Noctuidae Korea, 1998: 114; No. 282.

NOTE. The genus is placed here to Hypenodinae. � genit.: Pl. 3, fig. 4

18. Anachrostis minutissima Sugi, 1982 Ill. Cat. Noctuidae Korea, 1998: 114; No. 281.

NOTE. The female specimen was unavailable for dissection.

Subfamily ARAEOPTERONINAE Fibiger, 2004 Genus ARAEOPTERON Hampson, 1893 19. Araeopteron amoena Inoue, 1958 � genit.: Pl. 3, fig. 5; ♀ genit.: Pl. 178, fig. 6 Ill. Cat. Noctuidae Korea, 1998: 152; No. 385.

20. Araeopteron nebulosa Inoue, 1965



21. Araeopteron fragmenta Inoue, 1965*


Araeopteron fragmenta Inoue, 1965, Tinea, 7: 81, Pl., 15: 5A, 5B (TL: Japan, Kanagawa Pref., Fujisawa [BMNH, London]). Ill. Cat. Noctuidae Korea, 1998 _____.

DISTRIBUTION. S Korea, RFE (Primorye terr.), Japan (Honshu, Shikoku). NOTE. The species was refered in the Catalogue Korean Noctuidae as a possible species occurring for Korea. Hereby we confirm its occurrence in Korea. Material examined: 1 female, Korea GG, Hanam-shi, Mt. Geomdah 1. VIII 200 (Lee, Kom, Kim and Yu). 22. Araeopteron flaccida Inoue, 1958*


Araeopteron flaccisa Inoue, 1958, Tinea 4, 229, Pl. 32: 1 (TL: Japan, Chigasaki, Kanagawa Pref. [BMNH, London]). Ill. Cat. Noctuidae Korea, 1998 _____. Sohn, Ronkay, Kim and Cho, 2005.

DISTRIBUTION. S Korea, Japan (Honshu, Shikoku). NOTE. The female unknown.

Subfamily EUBLEMMINAE Forbes, 1954 Tribe EUBLEMMINI Forbes, 1954 Genus ODICE Hübner, [1823] (Type species: Noctua inamoena Hübner, [1803] [Europe]). 23. Odice arcuinna (Hübner, 1790)) � genit.: Pl. 4, fig. 2; ♀ genit.: Pl. 179, fig. 1 Ill. Cat. Noctuidae Korea, 1998: 155; No. 390 (Eublemma arcuina).

31

Genus EUBLEMMA Hübner, [1821] 1816 24. Eublemma amasina (Eversmann, 1842) � genit.: Pl. 4, fig. 3; ♀ genit.: Pl. 179, fig. 2 Ill. Cat. Noctuidae Korea, 1998: 155; No 389.

25. Eublemma cochylioides (Guenée, 1852)* Micra cochylioides Guenée, 1852, in Boisduval and Guenée, Hist. nat. Insectes (Lépid.) 2: 245 (TL: Réunion). Micra rosita Guenée, 1852, in Boisduval and Guenée, Hist. nat. Insectes (Lépid.) 2: 245 (TL: Australia). Thalpochares phoenissa Lederer, 1855, Verh Verh. zool. -bot. Ges. Wien, 5: 207, Pl. 2: 13 (TL: Cyprus [MNHU, Berlin]). Micra derogata Walker, [1858] 1857, List Specimens lepid. Insects Colln. Br. Mus. 12: 825 (TL: Australia [BMNH, London]). Thalpochares calida Rebel, 1894, Ann. K. K. Nat. Hist. Hofmus. Wien, 9: 63. Thalpochares calida Staudinger, [1895] 1894) Dt. ent. Z. Iris, 7: 283 (TL: Lebanon [MNHU, Berlin]). Porphyrinia confuscata Warren, 1913 in Seitz; 11: 221, Pl. 24b (TL: Australia, Queensland [BMNH, London]). Ill. Cat. Noctuidae Korea, 1998 _____. Park et al., 2001.

DISTRIBUTION. Korea (North), Japan, Taiwan, Philippines, Indonesia, Vietnam, India, Sri Lanka, Andamanes, Pakistan, Near East, S Europe, N Africa, Australia. NOTE. A single female was reported from Hajeu, North Korea (Park et al., 2001), but the specimen was unavailable for dissection. Genus ENISPA Walker, [1866] 1865 26. Enispa lutefascialis (Leech, 1889) � genit.: Pl. 4, fig. 4; ♀ genit.: Pl. 179, fig. 3 Ill. Cat. Noctuidae Korea, 1998: 149; No. 372.

27. Enispa albosignata (Staudinger, 1892)



28. Enispa bimaculata (Staudinger, 1892)



Genus CORGATHA Walker, 1858 29. Corgatha nitens (Butler, 1879) � genit.: Pl. 5, fig. 1; ♀ genit.: Pl. 179, fig. 6 Ill. Cat. Noctuidae Korea, 1998: 150; No. 376.

30. Corgatha argillacea (Butler, 1879)



31. Corgatha costimacula (Staudinger, 1892)



32. Corgatha dictaria (Walker, 1861)



33. Corgatha pygmaea Wileman, 1911 Ill. Cat. Noctuidae Korea, 1998: 151; No. 380.

32



34. Corgatha marumoi Sugi, 1982 Ill. Cat. Noctuidae Korea, 1998: 151; No. 381.

NOTE. The female specimen was unavailable for dissection. 35. Corgatha obsoleta Marumo, 1932



Genus ORUZA Walker, 1861 � genit.: Pl. 6, fig. 2

36. Oruza divisa (Walker, 1862) Ill. Cat. Noctuidae Korea, 1998: 157; No. 394.

NOTE. The female specimen was unavailable for dissection. 37. Oruza glaucotorna Hampson, 1910



NOTE. The female specimen was unavailable for dissection during preparation of the book. 38. Oruza mira (Butler, 1879)



39. Oruza submira Sugi, 1982



NOTE. The female specimen was unavailable for dissection during preparation of the book. 40. Oruza obliquaria Marumo, 1932 Ill. Cat. Noctuidae Korea, 1998: 157; No. 398.

NOTE. The species was previously reported for Korea by Pak (1959: 19), but no futher documentated records from known, and its distribution in Korea requires confirmation. 41. Oruza kunsuki Ahn, 1998



42. Oruza submirella Ahn, 1998



NOTE. The female specimen was unavailable for dissection during preparation of the book. Genus HYPOSADA Hampson, 1910 43. Hyposada brunnea (Leech, 1900) � genit.: Pl. 7, fig. 2; ♀ genit.: Pl. 180, fig. 6 Ill. Cat. Noctuidae Korea, 1998: 159; No. 402.

Genus LOPHORUZA Hampson, 1910 44. Lophoruza pulcherrima (Butler, 1879) � genit.: Pl. 7, fig. 3; ♀ genit.: Pl. 180, fig. 7 Ill. Cat. Noctuidae Korea, 1998: 156; No. 393.

33

Genus TAMBA Walker, 1869 � genit.: Pl. 7, fig. 4

45. Tamba corealis (Leech, 1889) Ill. Cat. Noctuidae Korea, 1998: 99; No. 229.

NOTE. The distribution of the species in the North Korea was confirmed by Park, et al. (2004) and also in the South Korea hereby. Material examined. 3 males, Korea, Wando, 8 IX 2002, 24, 25. IX 2002 (Gab-Jae Weon). 46. Tamba gensanalis (Leech, 1889) Ill. Cat. Noctuidae Korea, 1998: 99; No. 230.

NOTE. After description of the species from N Korea (Wonsan [Gensan]), no subsequent records are known. The specimens were not available for examination. Genus CORSA Walker, [1858] 1857 47. Corsa petrina (Butler, 1879) � genit.: Pl. 8, fig. 6; ♀ genit.: Pl. 182, fig. 2 Ill. Cat. Noctuidae Korea, 1998: 110; No. 270.

MATAEOMERA Butler, 1886 (Type species: Mataeomera dubia Butler, 1886 [Australia]). 48. Mataeomera esbiahni Sohn & Ronkay, 2001* � genit.: Pl. 7, fig. 4; ♀ genit.: Pl. 181, fig. 1 Sohn and Ronkay, 2001, Ins. Koreana 18(3): 220, figs 3, 4, 10, 12 (� genit.), 14 (♀ genit.) (TL: Korea, Gwangneung, Pocheon, Gyunggi Prov. [HNHM, Budapest]). Ill. Cat. Noctuidae Korea, 1998 _____. Sohn and Ronkay, 2001.

DISTRIBUTION. S Korea. Genus TRISATELES Tams, 1939 49. Trisateles emortualis ([Den. & Schiff.], 1775) � genit.: Pl. 7, fig. 5; ♀ genit.: Pl. 181, fig. 2 Ill. Cat. Noctuidae Korea, 1998: 158; No. 401.

Genus AVENTIOLA Staudinger, 1892 (PROLOPHOTA Hampson, 1896, syn. n.) NOTE. The examination of the external morphology and the genitalia of Aventiola pusilla and Prolophota trigonifera revealed their congenerity. Hereby the generic name Prolophota is synonimized with Aventiola. 50. Aventiola pusilla (Butler, 1879)



51. Aventiola trigonifera Hampson, 1896, comb. n. � genit.: Pl. 8, fig. 2; ♀ genit.: Pl. 181, fig. 4 Ill. Cat. Noctuidae Korea, 1998: 113; No. 279 (Prolophota trigonifera).

Genus PARAGONA Staudinger, 1892 52. Paragona multisignata (Christoph, 1881) � genit.: Pl. 8, fig. 3; ♀ genit.: Pl. 181, fig. 5 Ill. Cat. Noctuidae Korea, 1998: 112; No. 276.

34

53. Paragona inchoata (Wileman, 1911)




54. Paragona sp.* Ill. Cat. Noctuidae Korea, 1998 _____.

DISTRIBUTION. S Korea, RFE (Primorye terr.). NOTE. The description of the species is pending. Material examined. 1 female, Korea, Gwangleung (GG), 25. VI 1999 (T. S. Kwon and B. K. Byun); 8 specimens, Korea, Gwangleung, 8. VIII 2005 (H. L. Han). Genus ANATATHA Hampson, 1926 55. Anatatha lignea (Butler, 1879) � genit.: Pl. 9, fig. 1; ♀ genit.: Pl. 182, fig. 3 Ill. Cat. Noctuidae Korea, 1998: 106; No. 256.


56. Anatatha misae Sugi, 1982 Ill. Cat. Noctuidae Korea, 1998: 106; No. 257.

NOTE. The female specimen was unavailable for dissection during preparation of the book. 57. Anatatha basinigra Kononenko & Ahn, 1998 � genit.: Pl. 9, fig. 30; ♀ genit.: Pl. 182, fig. 4 Ill. Cat. Noctuidae Korea, 1998: 107; No. 258.

NOTE. The female genitalia was described and illustrated by Sohn, Ronkay and Kononenko (2004). Genus DIOMEA Walker, [1858] 1857 58. Diomea cremata (Butler, 1878) � genit.: Pl. 9, fig. 4; ♀ genit.: Pl. 182, fig. 5 Ill. Cat. Noctuidae Korea, 1998: 107; No. 259.

59. Diomea discisigna Sugi, 1963



60. Diomea jankowskii (Oberthür, 1880)



Genus MAGUDA Walker, [1866] 1865 61. Maguda suffusa (Walker, 1863)



NOTE. The female specimen was unavailable for study. Tribe PANGRAPTINI Grote, 1882 Genus PANGRAPTA Hübner, 1818 62. Pangrapta perturbans (Walker, 1858) � genit.: Pl. 10, fig. 1; ♀ genit.: Pl. 183, fig. 1 Ill. Cat. Noctuidae Korea, 1998: 100; No. 232.

35

63. Pangrapta costaemacula Staudinger, 1888



64. Pangrapta suaveola Staudinger, 1888



65. Pangrapta curtalis (Walker, [1866] 1865)



66. Pangrapta vasava (Butler, 1881)



67. Pangrapta flavomacula Staudinger, 1888



68. Pangrapta trilineata (Leech, 1900)



69. Pangrapta disruptalis (Walker, [1866] 1865)



70. Pangrapta lunulata (Sterz, 1915)



71. Pangrapta costinotata (Butler, 1881)



NOTE. The systematic position of the species in the genus Pangrapta is uncertain. By the male genitalia structures, the species show affinity with Naganoella timandra and Condate angulina (Guenée, 1852), illustrated by Holloway (2005). 72. Pangrapta marmorata Staudinger, 1888



73. Pangrapta obscurata (Butler, 1879)



74. Pangrapta textilis (Leech, 1889)



75. Pangrapta griseola Staudinger, 1892



76. Pangrapta parvula (Leech, 1900) Ill. Cat. Noctuidae Korea, 1998: 104; No. 246.

36


Genus NAGANOELLA Sugi, 1982 77. Naganoella timandra (Alphéraky, 1897) � genit.: Pl. 12, fig. 4; ♀ genit.: Pl. 185, fig. 2 Ill. Cat. Noctuidae Korea, 1998: 109; No. 265.

Miscellaneous genera Genus ECTOGONIA Hampson, 1896 78. Ectogonia butleri (Leech, 1900) � genit.: Pl. 12, fig. 5; ♀ genit.: Pl. 185, fig. 3 Ill. Cat. Noctuidae Korea, 1998: 114; No. 284.

NOTE. By the male genitalia structures, the species show affinity with the genus Olulis Walker, 1863. Genus STENOGRAPTA Sugi, 1959 79. Stenograpta stenoptera Sugi, 1959

♀ genit.: Pl. 185, fig. 5


NOTE. The male specimen was not available for examination. Genus AMPHITROGIA Hampson, 1926 80. Amphitrogia amphidecta (Butler, 1879) � genit.: Pl. 12, fig. 6; ♀ genit.: Pl. 185, fig. 4 Ill. Cat. Noctuidae Korea, 1998: 104; No. 248.

Genus LOPHOMILIA Warren, 1913 (ATUNTSEA Berio, 1977, syn. n.) NOTE. The examination of the genitalia structures of the members of genera Lophomilia and Athuntsea shows their congenerity. The generic name Athuntsea is synonimised with Lophomilia. 81. Lophomilia flaviplaga (Warren, 1912)



82. Lophomilia takao Sugi, 1982



83. Lophomilia polybapta (Butler, 1879)



84. Lophomilia sp.*


Ill. Cat. Noctuidae Korea, 1998 _____.

DISTRIBUTION. S. Korea, China, RFE (Primorye terr.). NOTE. An undescribed species; distributed in Korea, China and the Russian Far East. The description is pending. 85. Lophomilia kogii (Sugi, 1977), comb. n.


Ill. Cat. Noctuidae Korea, 1998: 99; No. 231 (Atuntsea kogii).

37

Genus PARAGABARA Hampson, 1926 86. Paragabara flavomacula (Oberthür, 1880) � genit.: Pl. 13, fig. 6; ♀ genit.: Pl. 186, fig. 3 Ill. Cat. Noctuidae Korea, 1998: 112; No. 273.

87. Paragabara ochreipennis Sugi, 1962



88. “Paragabara” secunda Remm, 1983



NOTE. The species is not congeneric with P. flavomacula, the type species of the genus Paragabara. The generic position of the species is required clarification. Genus HEPATICA Staudinger, 1892 89. Hepatica anceps Staudinger, 1892 � genit.: Pl. 14, fig. 3; ♀ genit.: Pl. 186, fig. 6 Ill. Cat. Noctuidae Korea, 1998: 106; No. 106.

90. Hepatica nakatanii Sugi, 1982*


Moths of Japan, 1: 881, 2: Pl.. 218: 34, 35, pl. 378: 5, 6 (male genitalia) (Japan, Honshu, Wakayama Pref., Oto Mts., Shimokawa Osugidani [NIAES]). Ill. Cat. Noctuidae Korea, 1998 ______. Han and Byun, 2007.

DISTRIBUTION. S. Korea (Wando I.), Japan (Honshu, Shikoku, Kyushu, Tsushima, Yakushima, Fukushima I.). Genus GONEPATICA Sugi, 1982 91. Gonepatica opalina (Butler, 1879) � genit.: Pl. 14, fig. 5; ♀ genit.: Pl. 187, fig. 1 Ill. Cat. Noctuidae Korea, 1998: 105; No. 254.

Genus HYPOSTROTIA Hampson, 1926 92. Hypostrotia cinerea (Butler, 1878) � genit.: Pl. 14, fig. 6; ♀ genit.: Pl. 187, fig. 2 Ill. Cat. Noctuidae Korea, 1998: 108; No. 262.

NOTE. The systematic position of the genus in the subfamily Eublemminae is uncertain.

93. Scedopla diffusa Sugi, 1959

Genus SCEDOPLA Butler, 1878 � genit.: Pl. 15, fig. 1; ♀ genit.: Pl. 187, fig. 3


Genus LEIOSTOLA Hampson, 1926 94. Leiostola mollis (Butler, 1879) � genit.: Pl. 15, fig. 2; ♀ genit.: Pl. 187, fig. 4 Ill. Cat. Noctuidae Korea, 1998: 108; No. 264.

Genus OGLASA Walker, [1859] 1858 (Type species: Oglasa lagusalis Walker, [1859] 1858 [Borneo]). 95. Oglasa bifidalis (Leech, 1889)* Harmatelia bifidalis Leech, 1889, Entomologist, 22: 64, Pl. 2: 11 (TL: Japan, Hakodate [BMNH, London]).

38

Ill. Cat. Noctuidae Korea, 1998 _____. Kim, 1999.

DISTRIBUTION. Korea, Japan. NOTE. Reported previously for Korea from the Ulrung Isl. (Kim, 1999). The specimen was unavailable for this study. Genus HOLOCRYPTIS Lucas, 1892 NOTE. The systematic position of the genus Holocryptis in the subfamily Eublemminae is uncertain. 96. Holocryptis ussuriensis (Rebel, 1901)



97. Holocryptis nymphula (Rebel, 1909)



Subfamily HERMINIINAE Herrich-Schüffer, 1845 Genus EDESSENA Walker, [1859] 1858 98. Edessena gentiusalis Walker, [1859] 1858 � genit.: Pl. 15, fig. 4; ♀ genit.: Pl. 187, fig. 7 Ill. Cat. Noctuidae Korea, 1998: 24; No. 1.

99. Edessena hamada (Felder & Rogenhofer, 1874) � genit.: Pl. 15, fig. 6; ♀ genit.: Pl. 187, fig. 8 Ill. Cat. Noctuidae Korea, 1998: 24; No. 2.

Genus HADENNIA Moore, 1887 100. Hadennia incongruens (Butler, 1879) � genit.: Pl. 16, fig. 1; ♀ genit.: Pl. 188, fig. 1 Ill. Cat. Noctuidae Korea, 1998: 25; No. 3.

101. Hadennia obliqua (Wileman, 1911)*


Trans. ent. Soc. Lond. 1911: 255, Pl. 31: 15 (Nodaria) [TL: Japan, Kyushu]). Ill. Cat. Noctuidae Korea, 1998 _____. Han, Sohn and Weon, 2006.

DISTRIBUTION. Korea, Japan, Taiwan. NOTE. The species was first reported for Korea by Pak (1959: 37) and later the name was included to Check list of Insects from Korea (1994: 381). However it was not included in the Catalogue of Korean Noctuidae by the reason of uncertain identification. Its occurrence in Korea was reconfirmed by Han, Sohn and Weon (2006). Genus MOSOPIA Walker, [1866] 102. Mosopia sordidum (Butler, 1879) � genit.: Pl. 16, fig. 3; ♀ genit.: Pl. 188, fig. 2 Ill. Cat. Noctuidae Korea, 1998: 25; No. 4.

Genus CIDARIPLURA Butler, 1879 103. Cidariplura gladiata Butler, 1879 � genit.: Pl. 16, fig. 4; ♀ genit.: Pl. 188, fig. 3 39


104. Cidariplura bilineata (Wileman & South, 1919) � genit.: Pl. 296, fig. 2; ♀ genit.: Pl. 188, fig. 4 Ill. Cat. Noctuidae Korea, 1998: 25; No. 6.

105. Cidariplura signata (Butler, 1879)



Genus PARACOLAX Hübner, [1825] 1816 106. Paracolax contigua (Leech, 1900) � genit.: Pl. 16, fig. 6; ♀ genit.: Pl. 188, fig. 5 Ill. Cat. Noctuidae Korea, 1998: 26; No. 9.

107. Paracolax sugii Owada, 1992



108. Paracolax tristalis (Fabricius, 1794)



109. Paracolax trilinealis (Bremer, 1864)



110. Paracolax pryeri (Butler, 1879)



111. Paracolax fascialis (Leech, 1889)

genit.: Pl. 17, fig. 4; ♀ genit.: Pl. 189, fig. 1


112. Paracolax fentoni (Butler, 1879)



Genus IDIA Hübner, [1813] 113. Idia quadra (Graeser, 1888 [1889]) � genit.: Pl. 17, fig. 6; ♀ genit.: Pl. 189, fig. 3 Ill. Cat. Noctuidae Korea, 1998: 28; No. 15.

114. Idia curvipalpis (Butler, 1879)



Genus GYNAEPHILA Staudinger, 1892 115. Gynaephila maculifera Staudinger, 1892 � genit.: Pl. 18, fig. 2; ♀ genit.: Pl. 189, fig. 5 Ill. Cat. Noctuidae Korea, 1998: 114; No. 285.

NOTE. The genus Gynaephila is transferred from Ophiderinae (sensu auctorum) to Herminiinae by Lödl (1998). Genus HYDRILLODES Guenée, 1854 116. Hydrillodes morosa (Butler, 1879) � genit.: Pl. 18, fig. 3; ♀ genit.: Pl. 189, fig. 6

40


117. Hydrillodes pacificus Owada, 1982



118. Hydrillodes lentalis Guenée, 1854*


Hydrillodes lentalis Guenée, 1854,, in Boisduval and Guenée, Hist. nat. Insectes (Lépid.) 8: 66, Pl. 5: 3 (TL: India [BMNH, London]). Hypochalcia repugnalis Walker, 1863, List Specimens lepid. Insects Colln. Br. Mus. 27: 47 (TL: Borneo [BMNH, London]). Bocana basalis Moore, 1867, Proc. zool. Soc. Lond. 1867: 88 (TL: [India] Bengalia [BMNH, London]). Hydrillodes submorosa Strand, 1920, Arch. Naturgesh. 84A(12): 165 (TL: Taiwan, Kosempo [EI, Eberswalde]). Ill. Cat. Noctuidae Korea, 1998 _____ (See p. 29). Sohn, Ronkay and Choi, 2005.

DISTRIBUTION. Korea, Japan, China, Taiwan, Vietnam, Laos, Burma, India, Bangladesh. SriLanka, Andamanes, Indonesia, S. Afrika. NOTE. The species was reported for Korea as H. lentalis by Leech (1889a: 565) and some subsequent Korean authors, and also as H. repugnalis by Pak (1964: 128). It was refered by Kononenko et al. (1998), but not included in the Catalogue of Korean Noctuidae by the reason of its high similarity with H. morosa and uncertain identity of the previous records. The residence of the species in Korea was reconfirmed by Sohn, Ronkay and Choi (2005). Genus BERTULA Walker, 1858 119. Bertula bistrigata (Staudinger, 1888) � genit.: Pl. 18, fig. 6; ♀ genit.: Pl. 189, fig. 8 Ill. Cat. Noctuidae Korea, 1998: 30; No. 19.

120. Bertula spacoalis (Walker, 1859)



121. Adrapsa simplex (Butler, 1879)*


Egnasia simplex Butler, 1879, Illustr. typical Specimens Lepid. Heterocera Colln Br. Mus. Pl. 57: 5 (TL: Japan, Yokohama [BMNH, London]). Ill. Cat. Noctuidae Korea, 1998 _____. Sohn, Ronkay and Choi, 2005.

DISTRIBUTION. Korea, China, Japan, Taiwan. Genus NODARIA Guenée, 1854 122. Nodaria tristis (Butler, 1879) � genit.: Pl. 19, fig. 3; ♀ genit.: Pl. 190, fig. 3 Ill. Cat. Noctuidae Korea, 1998: 30; No. 21.

Genus SIMPLICIA Guenée, 1854 123. Simplicia rectalis (Eversmann, 1842) � genit.: Pl. 19, fig. 4; ♀ genit.: Pl. 190, fig. 4 Ill. Cat. Noctuidae Korea, 1998: 30; No. 22.

124. Simplicia niphona (Butler, 1878)



41

125. Simplicia xanthoma Prout, 1928



126. Simplicia caeneusalis (Walker, [1859] 1858) � genit.: Pl. 20, fig. 1; ♀ genit.: Pl. 190, fig. 7 Ill. Cat. Noctuidae Korea, 1998: 31; No. 25.

Genus ZANCLOGNATHA Lederer, 1857 127. Zanclognatha griselda (Butler, 1879) � genit.: Pl. 20, fig. 2; ♀ genit.: Pl. 190, fig. 8 Ill. Cat. Noctuidae Korea, 1998: 32; No. 26.

128. Zanclognatha lilacina (Butler, 1879)



129. Zanclognatha lunalis (Scopoli, 1763)



130. Zanclognatha lui Han & Park, 2005*


Zanclognatha lui Han and Park, 2005, Korean J. Syst. Zool. 21(1): 2, figs 1A-D, 3A-B (� genit.) (N China, Changboshan [CIS, Kangwon]). Ill. Cat. Noctuidae Korea, 1998 _____. Han, Sohn and Weon, 2006.

DISTRIBUTION. N Korea, N China. 131. Zanclognatha curvilinea (Wileman & South, 1917)



132. Zanclognatha fumosa (Butler, 1879)



133. Zanclognatha tarsipennalis (Treitschke, 1835) � genit.: Pl. 21, fig. 2; ♀ genit.: Pl. 191, fig. 6 Ill. Cat. Noctuidae Korea, 1998: 33; No. 31.

134. Zanclognatha triplex (Leech, 1900)



135. Zanclognatha helva (Butler, 1879)



136. Zanclognatha umbrosalis Staudinger, 1892



137. Zanclognatha tenuialis Rebel, 1899

genit.: Pl. 21, fig. 6; ♀ genit.: Pl. 192, fig. 2

Zanclognatha tenuialis Rebel, 1899, Verh. zool.-bot. Ges. Wien, 49: 169 (TL: Italy). Zanclignatha triplex koreognatha Bryk, 1948, Ark. Zool. 41(A) 1: 147, pl. 5: 13 (TL: North Korea: Hamgyeong bugdo [Motojondo] [NRM, Stockholm]). Reference: Kononenko, 1996: 14 (Zanclognata umbrosalis, incorrect synonymy); Kononenko et al., 1996 (Zanclognatha umbrosalis, part nec Staudinger, 1892, misidentification); Owada, 2006 (Zanclognatha tenuialis).

42

Ill. Cat. Noctuidae Korea, 1998: 34; No. 36 (Zanclognatha umbrosalis, part nec Staudinger, 1892, misidentification).

NOTE. Kononenko (1996) incorrectly synonimized Zanclignatha triplex koreognatha Bryk with Zanclognatha umbrosalis, this synonymy was cited in the Catalogue of the Noctuidae of Korea (Kononenko et al., 1998). Owada (2006) re-examined the holotype of Zanclignatha triplex koreognatha, and revealed incorrect synonymy and synonymized the taxon with Zanclognatha tenuialis. DISTRIBUTION. Korea, RFE (Primorye terr., South of Khabarovsk terr. and Amur reg.), South Transbaikalia, West Siberia, Central and South Europe. The species is reported for Korea for the first time. 138. Zanclognatha perfractalis Bryk, 1948


Zanclognatha sviridovi Owada, 1992, Tinea 13(18): 190 (TL: Japan: Aomori Pref., Kiuzukuri, Miyagi Pref., Gunma Pref.; Korea: Suwon, Yangdok, Pyongan-do [NSM, Tokyo]). Ill. Cat. Noctuidae Korea, 1998: 34; No. 35 (Zanclognatha perfractalis = southi, wrong synonymy; Zanclognatha sviridovi).

NOTE. After examination of the holotype of Zanclognatha perfractalis, Owada (2006) stated that Zanclognatha perfractalis is not identical with Zanclognatha southi Owada, 1982, but with Zanclognatha sviridovi Owada, 1992. Zanclognatha southi, cited in the Catalogue of Korean Noctuidae (Kononenko et al., 1998) as a synonym of Zanclognatha perfractalis in fact is a distinct species, known for the present only from Japan. 139. Zanclognatha violacealis Staudinger, 1892



Genus PECHIPOGO Hübner, [1825] 1816 140. Pechipogo strigilata (Linnaeus, 1758) � genit.: Pl. 22, fig. 3; ♀ genit.: Pl. 192, fig. 5 Ill. Cat. Noctuidae Korea, 1998: 35; No. 38.

Genus POLYPOGON Schrank, 1802 141. Polypogon tentacularia (Linnaeus, 1758) � genit.: Pl. 22, fig. 4; ♀ genit.: Pl. 192, fig. 6 Ill. Cat. Noctuidae Korea, 1998: 36; No. 39.

142. Polypogon tarsicrinata (Bryk, 1948)


Zanclognatha tarsicrinata (Bryk, 1948), Ark. Zool. 41(A)1: 146, Pl. 5: 5 (TL: Korea, Shuotsu, RM, Stockholm). Reference: Poole, 1989: 831 (Polypogon tarsicrinata); Sviridov, 1990: 99 (Polypogon gryphalis, incorrect synonymy, misidentification); Kononenko et al., 1989 (Polypogon gryphalis, misidentification); Owada, 2006 (Polypogon tarsicrinata, correction). Ill. Cat. Noctuidae Korea, 1998: 36; No. 40 (Polypogon gryphalis, misidentification).

NOTE. Sviridov (1990) synonymised this taxon with Polypogon gryphalis Herrich-Schüffer, 1851), however Owada (2006) restored the name which is applicable to the distinct species P. tarsicrinata and described the morphological differences between two taxa, i. e. P. gryphalia and P. tarsicrinata. Here we follow the result by Owada (2006). DISTRIBUTION. Korea, Japan (Hokkaido, Honshu, RFE (Primorye terr., South of Khabarovsk terr., South of Amur region, Sakhalin and Southern Kuriles. It is not clear which taxon is occurs in 43

the southern Siberia. Genus HERMINIA Latreille, 1802 143. Herminia grisealis ([Den. & Schiff.], 1775) � genit.: Pl. 23, fig. 1; ♀ genit.: Pl. 192, fig. 8 Ill. Cat. Noctuidae Korea, 1998: 37; No. 41.

144. Herminia robiginosa (Staudinger, 1888)



145. Herminia tarsicrinalis (Knoch, 1782)



146. Herminia arenosa Butler, 1878



147. Herminia stramentacealis Bremer, 1864



148. Herminia dolosa Butler, 1879



149. Herminia innocens Butler, 1878



Genus STENHYPENA Hampson, 1895 150. Stenhypena nigripuncta (Wileman, 1911) � genit.: Pl. 24, fig. 2; ♀ genit.: Pl. 193, fig. 6 Ill. Cat. Noctuidae Korea, 1998: 38; No. 48.

Genus HIPOEPA Walker, [1859] 1858 151. Hipoepa fractalis (Guenée, 1854) � genit.: Pl. 24, fig. 3; ♀ genit.: Pl. 193, fig. 7 Ill. Cat. Noctuidae Korea, 1998: 39; No. 49.

Genus SINARELLA Bryk, 1948 152. Sinarella aegrota (Butler, 1879) � genit.: Pl. 24, fig. 4; ♀ genit.: Pl. 193, fig. 7 Ill. Cat. Noctuidae Korea, 1998: 39; No. 50.

153. Sinarella cristulalis Staudinger, 1892



154. Sinarella nigrisigna (Leech, 1900)



155. Sinarella punctalis (Herz, 1904)



156. Sinarella japonica (Butler, 1881) 44



157. Sinarella rotundipennis Owada, 1982



Subfamily HYPENINAE Herrich-Schüffer, 1845 Genus RHYNCHINA Guenée, 1854 158. Rhynchina cramboides (Butler, 1879) � genit.: Pl. 25, fig. 4; ♀ genit.: Pl. 194, fig. 6 Ill. Cat. Noctuidae Korea, 1998: 52; No. 92.

159. Rhynchina sagittata (Butler, 1889) Ill. Cat. Noctuidae Korea, 1998: 52; No. 94.

NOTE. The only known Korean record of this species was by Leech (1900: 651) from Wonsan [Gensan], but no subsequent records after Leech (1900). The specimen was unavailable for study. Quite possible Leech misidentified is probably one of below mentioned species (i.e. Rhynchina changyangis or Zekelita plusioides) as Rhynchina sagittata. 160. Rhynchina changyangis Mayerl & Lödl, 1999

� genit.: Pl. 25, figs 5, 6; ♀ genit.: Pl. 194, fig. 7

Rhynchina changyangis Mayerl & Lödl, 1999, Quadrifina, 2: 44, figs 183-184 (TL: China, Prov. Hubei, Changyang [BMNH, London]). Ill. Cat. Noctuidae Korea, 1998: 53; No. 95. (Zekelita sp. unidentified).

DISTRIBUTION. Korea, Japan. NOTE. The species was included in the Catalogue of Korean Noctuidae as Zekelita sp. (unidentified). Hereby it identified as Rhynchina changyangis by original description (Mayerl and Lödl, 1999). We assume that the same species is occurring in Japan. It was reported as an unidentified species (Rhynchodontodes sp.) in Ophiderinae (sensu auctorum) by Sugi (1982, Pl. 20, fig. 30) . Genus ZEKELITA Walker, 1863 161. Zekelita plusioides (Butler, 1889) � genit.: Pl. 26, fig. 1; ♀ genit.: Pl. 194, fig. 8 Ill. Cat. Noctuidae Korea, 1998: 52; No. 93.

NOTE. For identification of this species see Lödl and Mayerl (1998). Genus LATIROSTRUM Hampson, 1895 162. Latirostrum bisacutum Hampson, 1895 � genit.: Pl. 26, fig. 2; ♀ genit.: Pl. 195, fig. 1 Ill. Cat. Noctuidae Korea, 1998: 52; No. 91.

Genus ANORATHA Moore, 1867 (Type species: Anoratha costalis Moore, 1867 [India]). 163. Anoratha paritalis (Walker, ([1859] 1858)* � genit.: Pl. 26, fig. 3; ♀ genit.: Pl. 195, fig. 2 Hypena paritalis Walker, ([1859] 1858), List Specimens lepid. Insects Colln. Br. Mus. (TL: Sri Lanka [BMNH, London]). Ill. Cat. Noctuidae Korea, 1998 _____. Sohn and Kim, 2003.

45

DISTRIBUTION. Korea, Vietnam, India, Sri Lanka, Indonesia, New Guinea, New Caledonia. Genus HYPENOMORPHA Sugi, 1977 164. Hypenomorpha calamina (Butler, 1879)



Genus CHIBIDOKUGA Matsumura, 1933 165. Chibidokuga hypenodes Inoue, 1979 � genit.: Pl. 26, fig. 5; ♀ genit.: Pl. 195, fig. 3 Ill. Cat. Noctuidae Korea, 1998: 115; No. 286.

NOTE. The genus Chibidokuga placed to Hypeninae following to Inoue (1979). Genus ITMAHARELA Nye, 1975 166. Itmaharela basalis (Moore, 1882)



167. Harita belinda (Butler, 1879)

Genus HARITA Moore, 1882 � genit.: Pl. 27, fig. 1; ♀ genit.: Pl. 195, fig. 4


Genus HYPENA Schrank, 1802 Subgenus BOMOLOCHA Hübner, [1825] 1816 168. Hypena stygiana Butler, 1878 � genit.: Pl. 27, fig. 2; ♀ genit.: Pl. 195, fig. 5 Ill. Cat. Noctuidae Korea, 1998: 45; No. 69.

169. Hypena zilla Butler, 1879



170. Hypena albopunctalis Leech, 1889 Ill. Cat. Noctuidae Korea, 1998: 46; No. 71.

NOTE. The species was described (Leech, 1889) fron N Korea (Wonsan [Gensan]), its distribution in N Korea was reconfirmed (Park, et al., 2001). The specimen was not available for examination. 171. Hypena bicoloralis Graeser, 1888 [1889]



172. Hypena melanica Sugi, 1959



173. Hypena nigrobasalis (Herz, 1904)



174. Hypena rivuligera (Butler, 1881) Ill. Cat. Noctuidae Korea, 1998: 46; No. 75.

46


175. Hypena squalida (Butler, 1878)



Subgenus HYPENA Schrank, 1802 176. Hypena subcyanea Butler, 1881 Ill. Cat. Noctuidae Korea, 1998: 47; No. 77.

NOTE. Reported for Korea by Leech (1889a: 559; 1900: 656) from Wonsan [Gensan], no other authentic records since Leech is available. The specimen was not available for examination. 177. Hypena tristalis Lederer, 1853




178. Hypena abducalis (Walker, 1859) Ill. Cat. Noctuidae Korea, 1998: 48; No. 79.

NOTE. The male specimen was not available for dissection. 179. Hypena kengkalis Bremer, 1864



180. Hypena strigatus (Fabricius, 1798) (Synonymy: commixtalis Zeller, 1852; fumidalis Zeller, 1852; jussalis Walker, [1859] 1858; rectivittalis Moore, 1867; beatalis Felder & Rogenhofer, 1874; minna Butler, 1879; speculalis Swinhoe, 1885; vecordialis Swinhoe, 1885; haplochrosta Hulstaert, 1924). Ill. Cat. Noctuidae Korea, 1998: 48; No. 81.

NOTE. Reported from Korea (Leech, 1989, 1900, Nam, 1982: 515, pl. 70, fig, 580) under synonymic names as Hypena minna and also as H. rectivittalis. The species was included in the Check List of Insects from Korea (1994: 381) under its synonymic name H. rectivittalis. We did not find this species in examined collections. 181. Hypena proboscidalis (Linnaeus, 1758)



182. Hypena furva Vileman, 1911*


Trans. ent. Soc. Lond. 1911: 262, Pl. 31: 17 (TL: Japan, Satsuma, Kagoshima, [BMNH, London]) Ill. Cat. Noctuidae Korea, 1998 _____.

NOTE. A female of this species is newly found in Korea. The species was identified according to illustration by Sugi (1982a, 1: 909, 2, pl. 222, 33). Material examined: 1 female, Korea, Ok-chon, CB, 14. VII 2002 (Han Hui Lin). DISTRIBUTION. Korea, Japan. 183. Hypena occata Moore, 1882 Ill. Cat. Noctuidae Korea, 1998: 49; No. 83.

47

NOTE. Reported for Korea from Wonsan [Gensan] by Leech (1900: 654). Its distribution in Korea was reconfirmed by Park et al. (2001). The specimen was not unavailable for examination. 184. Hypena indicatalis Walker, 1859 Ill. Cat. Noctuidae Korea, 1998: 50; No. 84.

NOTE. Reported for Korea from Wonsan [Gensan] by Leech (1889a: 556; 1900: 654). Its occurence in N and S Korea was reconfirmed (Park et al., 2001) and (Sohn, pers. comm.) [1 female, Cheju, 22. IV 1999 V.K. Byun] is confirmed. The specimens were unavailable for examination. 185. Hypena conspersalis Staudinger, 1888



186. Hypena sagitta (Fabricius, 1775)



NOTE. The male specimen was not available for dissection. 187. Hypena trigonalis (Guenée, 1854)



188. Hypena claripennis (Butler, 1878)



189. Hypena amica (Butler, 1878)




190. Hypena sp. 1 Ill. Cat. Noctuidae Korea, 1998: 51; No. 90 (Hypena sp. unidentified).

NOTE. This unidentified species of Hypena was reported and illustrated from Japan by Sugi (1982a, 1: 909, 2, pl. 222, 44-45) as Hypena sp. 8, and the same species was recorded in Korea. We can not identify the species for the time being. DISTRIBUTION. Korea, Japan. Genus STENBERGMANIA Bryk, 1948 191. Stenbergmania albomaculalis (Bremer, 1864) � genit.: Pl. 29, fig. 5; ♀ genit.: Pl. 197, fig. 3 Ill. Cat. Noctuidae Korea, 1998: 105; No. 253.

Subfamily PHYTOMETRINAE Hampson, 1913 Genus PHYTOMETRA Haw., 1809 192. Phytometra amata (Butler, 1879) � genit.: Pl. 29, fig. 6; ♀ genit.: Pl. 197, fig. 4 Ill. Cat. Noctuidae Korea, 1998: 109; No. 266.

48

Genus RHESALA Walker, 1858 193. Rhesala moestalis (Walker, [1866] 1865)


Magulaba moestalis Walker, [1866] 1865, List Specimens lepid. Insects Colln. Br. Mus. 34: 1127 (TL: Sierra Leone [BMNH, London]). Hingula figurata Moore, [1885] 1884-1887. Lepid. Ceylon, 3: 199, Pl. 174: 10 (TL.: Sri Lanka [BMNH, London]). Egnasia erebina Butler, 1879, Ann. Mag. nat. Hist. (5)4: 450 (TL: Japan, Yokohama [BMNH, London]). Rimulia malgassica Heyden, 1891, in Saalmüller; Lep. Madagascar: 483, fig. 173 (TL: Madagascar). Rhesala imparata, auct. Ill. Cat. Noctuidae Korea, 1998: 111; No. 271 (Rhesala imparata, misidentification).

DISTRIBUTION. Korea, Japan, Vietnam, ?Sri Lanka, Sierra Leone, ?Madagascar. NOTE. Holloway (2005) stated that “Rhesala imparata” (sensu auctorum) is a complex of species. Among numerous names refered in literature (Poole, 1982; Nielsen et al., 1996) as synonyms of Rhesala imparata Walker, 1858, he revealed that Egnasia erebina Butler, 1879, was described from Japan and cited in literature as a synonym of imparata is a junior synonym of Magulaba moestalis Walker, [1866] 1865, and some other names: R. albolunata Moore, 1882, and R. cervina Moore, 1882, are applicable to distinct species. Holloway (2005) illustrated the male genitalia of imparata which is distinct from our specimen from Korea. We assume that the same species as in Japan occurring in Korea. Consequently for this taxon we use the name Rhesala moestalis. The distribution of the species is not clear due to unrevised synonymy, however the same species is occurring in Vietnam (male genitalia examined). Two names are cited in literature as a synonyms of Rh. moestalis: figurata Moore, [1885] 1884-1887 (TL: Sri Lanka) and malgassica Heyden, 1891(TL: Madagascar). Genus RAPARNA Moore, 1882 194. Raparna roseata Wileman & South, 1917 Ill. Cat. Noctuidae Korea, 1998: 109; No. 267.

NOTE. Raparna transversa Moore, 1882, reported in the Check List of Insects from Korea (1994: 380) was excluded from the Catalogue of the Korean Noctuidae. This record is based on Leech (1900: 11) report and repeated by other authors. No other documentated records of R. transversa from Korea since Leech are available. Mr. S. Sugi (pers. comm.) informed us that although Raparna transversa has been retained in the catalogue “Moths of Japan”, the only R. roseata is known from Japan. Apparently, data of R. transversa given by earlier authors from Japan and Korea may refer to misidentifications of R. roseata. The description of R. roseata is based on Leech’s specimens from Kyushu. The latter name actually replaces the earlier record of transversa from Japan. Consequently, herewith we use the name roseata instead of transversa. The distribution of the species was reconfirmed by Park et al. (2001): one male was collected in Kaesong, N Korea. The specimen was unavailable for dissection. Genus COLOBOCHYLA Hübner, [1825] 1816 195. Colobochyla salicalis ([Den. & Schiff.], 1775) � genit.: Pl. 30, fig. 2; ♀ genit.: Pl. 197, fig. 5 Ill. Cat. Noctuidae Korea, 1998: 41; No. 56.

Genus EUWILEMANIA 196. Euwilemania angulata (Wileman, 1911) � genit.: Pl. 30, fig. 3; ♀ genit.: Pl. 197, fig. 6 Ill. Cat. Noctuidae Korea, 1998: 105; No. 252.

49

NOTE. We tentatively place the genus Euwilemania in the subfamily Phytometrinae.

Subfamily AVENTIINAE Tutt, 1896 Genus LASPEYRIA Germ., 1810 197. Laspeyria flexula ([Den. & Schiff.], 1775) � genit.: Pl. 30, fig. 4; ♀ genit.: Pl. 197, fig. 7 Ill. Cat. Noctuidae Korea, 1998: 111; No. 272.

Genus SOPHTA Walker, [1863] 1862 198. Sophta subrosea (Butler, 1881) � genit.: Pl. 30, fig. 5; ♀ genit.: Pl. 197, fig. 8 Ill. Cat. Noctuidae Korea, 1998: 155; No. 391.

NOTE. The genus Sophta is placed here to the subfamily Aventiinae. 199. Sophta ruficeps (Walker, 1864)



Subfamily EREBINAE Leech, [1815] Tribe EREBINI Leech, [1815] Genus EREBUS Latreille, 1810 200. Erebus macrops (Linnaeus, 1768) � genit.: Pl. 31, fig. 1; ♀ genit.: Pl. 198, fig. 1 Ill. Cat. Noctuidae Korea, 1998: 74; No. 161.

201. Erebus ephesperis (Hübner, [1823])



Genus METOPTA Swinhoe, 1900 202. Metopta rectifasciata (Ménétriés, 1863) � genit.: Pl. 31, fig. 3; ♀ genit.: Pl. 198, fig. 3 Ill. Cat. Noctuidae Korea, 1998: 75; No. 165.

Tribe HYPOPYRINI Guenée, 1852 Genus SPIRAMA Guenée, 1852 203. Spirama helicina (Hübner, 1831) � genit.: Pl. 32, fig. 1; ♀ genit.: Pl. 198, fig. 4 Ill. Cat. Noctuidae Korea, 1998: 74; No. 163.

204. Spirama retorta (Clerck, 1759)



Genus HYPOPYRA Guenée, 1852 205. Hypopyra vespertilio (Fabricius, 1775) � genit.: Pl. 32, fig. 3; ♀ genit.: Pl. 198, fig. 6 Ill. Cat. Noctuidae Korea, 1998: 73; No. 160.

50

Tribe ARCTEINI Berio, 1992 Genus ARCTE Kollar, [1844] 206. Arcte coerula (Guenée, 1852) � genit.: Pl. 32, fig. 4; ♀ genit.: Pl. 198, fig. 7 Ill. Cat. Noctuidae Korea, 1998: 72; No. 159.

NOTE. The esystematic position of the genus Arcte in Erebinae is dubious.

Subfamily CALPINAE Boisduval, 1840 Tribe ANOMINI Grote, 1882 Genus ANOMIS Hübner, [1821] 1816 207. Anomis flava (Fabricius, 1775) � genit.: Pl. 33, fig. 1; ♀ genit.: Pl. 190, fig. 1 Ill. Cat. Noctuidae Korea, 1998: 81; No. 180.


208. Anomis mesogona (Walker, 1858) Ill. Cat. Noctuidae Korea, 1998: 81; No. 181.


209. Anomis involuta (Walker, [1858] 1857) Ill. Cat. Noctuidae Korea, 1998: 81; No. 182.


210. Anomis privata (Walker, 1865)

Ill. Cat. Noctuidae Korea, 1998: 82; No. 183 . (Synonyms: commoda Butler, 1878; subfulvida Warren, 1913; griseolineata Warren, 1913)

NOTE. For the synonymy cited see Sugi (1992a). 211. Anomis leucolopha Prout, 1928

� genit.: Pl. 33, fig. 5, 6; ♀ genit.: Pl. 199, fig. 5

(Synonyms: longipennis Sugi, 1982; maxima Berio, 1956) Ill. Cat. Noctuidae Korea, 1998: 82; No. 184 (Anomis longipennis).

NOTE. For the synonymy cited see Galsworthy (1997), Sugi and Junbo (2004) and Kononenko, (2005). Tribe CALPINI Boisduval, 1840 Genus CALYPTRA Ochsenheimer, 1816 212. Calyptra thalictri (Borkhausen, 1790) � genit.: Pl. 34, fig. 1; ♀ genit.: Pl. 199, fig. 6 Ill. Cat. Noctuidae Korea, 1998: 83; No. 186.

213. Calyptra hokkaida (Wileman, 1922)



214. Calyptra lata (Butler, 1881)



51

215. Calyptra gruesa (Draudt, 1950)



216. Calyptra fletcheri (Berio, 1956)



Genus ORAESIA Guenée, 1852 217. Oraesia excavata (Butler, 1878) � genit.: Pl. 34, fig. 6; ♀ genit.: Pl. 200, fig. 4 Ill. Cat. Noctuidae Korea, 1998: 84; No. 192.

218. Oraesia emarginata (Fabricius, 1794)



Genus PLUSIODONTA Guenée, 1852 219. Plusiodonta casta (Butler, 1878) � genit.: Pl. 35, fig. 2; ♀ genit.: Pl. 200, fig. 6 Ill. Cat. Noctuidae Korea, 1998: 85; No. 194.

220. Plusiodonta coelonota (Kollar, [1844])



NOTE. male specimen was unavailable for dissection. Genus EUDOCIMA Billberg, 1820 221. Eudocima salaminia (Cramer, 1777) � genit.: Pl. 35, fig. 4; ♀ genit.: Pl. 200, fig. 7 Ill. Cat. Noctuidae Korea, 1998: 86; No. 195.

222. Eudocima falonia (Linnaeus, 1763)


Ill. Cat. Noctuidae Korea, 1998: 86; No. 196. (Othreis fullonia).

NOTE. For the nomenclature, synonymy and priority the name falonia over fullonia see Zilli and Hogenes (2002). 223. Eudocima tyrannus (Guenée, 1852)



Tribe SCOLIOPTERIGINI Spuler, 1908 Genus SCOLIOPTERYX Germ., 1810 224. Scoliopteryx libatrix (Linnaeus, 1758) � genit.: Pl. 36, fig. 2; ♀ genit.: Pl. 201, fig. 3 Ill. Cat. Noctuidae Korea, 1998: 83; No. 185.

52

Subfamily CATOCALINAE Boisduval, [1828] Tribe TOXOCAMPINI Guenée, 1852 Genus CHRYSORITHRUM Butler, 1878 225. Chrysorithrum amata (Bremer & Grey, 1853) � genit.: Pl. 36, fig. 3; ♀ genit.: Pl. 201, fig. 4 Ill. Cat. Noctuidae Korea, 1998: 94; No. 217.

226. Chrysorithrum flavomaculata (Bremer, 1861)



Genus LYGEPHILA Billberg, 1820 227. Lygephila maxima (Bremer, 1861) � genit.: Pl. 36, fig. 5; ♀ genit.: Pl. 201, fig. 6 Ill. Cat. Noctuidae Korea, 1998: 77; No. 169.

228. Lygephila ludicra Hübner, 1790



229. Lygephila vulcanea (Butler, 1881)



230. Lygephila mirabilis (Bryk, 1948)



NOTE. We did not find the substantive differences in male genitalia between L. vulcanea and L. mirabilis, while the female genitalia have different structure of antevaginal plate and ductus bursae. Further investigation of this pair of the species is needed. 231. Lygephila moellendorfi (Herz, 1904)



232. Lygephila craccae ([Den. & Schiff.], 1775)



233. Lygephila viciae (Hübner, [1822])



234. Lygephila pastinum (Treitschke, 1826)



235. Lygephila subrecta Sugi, 1982



236. Lygephila recta (Bremer, 1864)



53

� genit.: Pl. 38, fig. 4.

237. Lygephila lilacina (Butler, 1878)*

Toxocampa lilacina Butler, 1878, Ann. Mag. nat. Hist., (5)1: 289 (TL: Japan, Yokohama [BMNH, London]). Lygephila ogatai Kinoshita & Sasaki, 1986, Tyo Ga, 38: 209, figs 1, 2, 5, 6, 9, 11, 15, 18, 20 (TL: Japan, Fukui [MNHO, Osaka]). Ill. Cat. Noctuidae Korea, 1998 _____.

DISTRIBUTION. Korea, Japan, Honshu). NOTE. The species was identified by comparison of the male genitalia preparation with photograph of genitalia lectotype (Kishida and Yoshimoo, 1988) and authentical specimens of L. recta complex (Qwada, 1988). First record for Korea. Material examined: 2 males, Korea, Chuncheon, 4. IX 1984 (K. T. Park). 238. Lygephila nigricostata (Graeser, 1890)



Genus AUTOPHILA Hübner, [1823] 1816 239. Autophila inconspicua (Butler, 1881) � genit.: Pl. 38, fig. 6; ♀ genit.: Pl. 203, fig. 4 Ill. Cat. Noctuidae Korea, 1998: 76; No. 168.

Genus APOPESTES Hübner, [1823] 1816 240. Apopestes indica Moore, 1883 � genit.: Pl. 39, fig. 1; ♀ genit.: Pl. 203, fig. 5 Ill. Cat. Noctuidae Korea, 1998: 76; No. 167.

Genus PERINAENIA Butler, 1878 241. Perinaenia accipiter (Felder & Rogenhofer, 1874)



NOTE. The female specimen was unavailable for dissection. Tribe ARYTRURINI Fibiger & Lafontaine, 2005 Genus ARYTRURA John, 1912 242. Arytrura musculus (Ménétriés, 1859) � genit.: Pl. 39, fig. 2; ♀ genit.: Pl. 203, fig. 6 Ill. Cat. Noctuidae Korea, 1998: 98; No. 226.

243. Arytrura subfalcata (Ménétriés, 1859)



Tribe EUCLIDINI Guenée, 1852 Genus EUCLIDIA Ochsenheimer, 1816 244. Euclidia dentata Staudinger, 1871 � genit.: Pl. 39, fig. 5; ♀ genit.: Pl. 204, fig. 2 Ill. Cat. Noctuidae Korea, 1998: 68; No. 146.

Subgenus LEUCOMELAS Hampson, 1913, stat. n. 245. Euclidia juvenilis (Bremer, 1861) comb. n. � genit.: Pl. 40, fig. 1; ♀ genit.: Pl. 204, fig. 3 Ill. Cat. Noctuidae Korea, 1998: 69; No. 148.

54

Subenus CALLISTEGE Hübner, 1816 246. Euclidia mi (Clerck, 1759) � genit.: Pl. 40, fig. 2; ♀ genit.: Pl. 204, fig. 4 Ill. Cat. Noctuidae Korea, 1998: 68; No. 147.

Genus MELAPIA Sugi, 1968 247. Melapia electaria (Bremer, 1864) � genit.: Pl. 40, fig. 3; ♀ genit.: Pl. 204, fig. 5 Ill. Cat. Noctuidae Korea, 1998: 70; No. 152.

Genus MOCIS Hübner, [1823] 1816 248. Mocis undata (Fabricius, 1775) � genit.: Pl. 40, fig. 4; ♀ genit.: Pl. 205, fig. 1 Ill. Cat. Noctuidae Korea, 1998: 69; No. 149.


249. Mocis annetta (Butler, 1878) Ill. Cat. Noctuidae Korea, 1998: 70; No. 150.

250. Mocis ancilla (Warren, 1913)



Tribe OPHIUSINI Guenée, 1852 (OMOPTERINI Boisduval, 1833, suppr. senior syn.) Genus SERRODES Guenée, 1852 251. Serrodes campana Guenée, 1852 � genit.: Pl. 41, fig. 1; ♀ genit.: Pl. 205, fig. 2 Ill. Cat. Noctuidae Korea, 1998: 89; No. 89.

Genus ARTENA Fabricius, 1794 252. Artena dotata (Fabricius, 1794) � genit.: Pl. 41, fig. 2; ♀ genit.: Pl. 205, fig. 3 Ill. Cat. Noctuidae Korea, 1998: 72; No. 158.

253. Thyas juno (Dalman, 1823)

Genus THYAS Hübner, [1824] � genit.: Pl. 41, fig. 3; ♀ genit.: Pl. 205, fig. 4


Genus OPHIUSA Ochsenheimer, 1816 254. Ophiusa disjungens (Walker, 1858)



255. Ophiusa tirhaca (Cramer, 1777)



256. Ophiusa olista (Swinhoe, 1893)



257. Ophiusa triphaenoides (Walker, 1858)



55

NOTE. The species was previously reported for Korea (Jeong, 1997: 51-54). Due to the external similarity of O. triphaenoides with O. olista, it could be confused with the former. We had no opportunity to check the reported specimen. The appearance of migrant specimens of O. triphaenoides in Korea is impossible, but required confirmation. The male specimen from Borneo (after Holloway, 2005) and female from Vietnam are illustrated here. PARALLELIA Hübner, 1818 (Type-species: Parallelia bistriarts Hübner, 1818) NOTE. In treatment of Parallelia-Dysgonia generic group we follow to the revision of Parallelia generic complex by Holloway and Miller (2003). � genit.: Pl. 42, fig. 3; ♀ genit.: Pl. 206, fig. 1

258. “Parallelia” arctotaenia (Guenée, 1852)

Ill. Cat. Noctuidae Korea, 1998: 67; No. 142 (Dysgonia arctotaenia).

Genus BASTILLA Swinhoe, 1918 (Type-species: Ophiusa redunca Swinhoe, 1900 [Australia]) 259. Bastilla stuposa (Fabricius, 1794) � genit.: Pl. 42, fig. 4; ♀ genit.: Pl. 206, fig. 2 Ill. Cat. Noctuidae Korea, 1998: 65; No. 137 (Dysgonia stuposa).


260. Bastilla maturata (Walker, 1858)

Ill. Cat. Noctuidae Korea, 1998: 67; No. 140 (Dysgonia maturata).

261. Bastilla arcuata (Moore, 1887)


Ill. Cat. Noctuidae Korea, 1998: 66; No. 139 (Dysgonia arcuata).

262. Bastilla absentimacula (Guenée, 1852)


Ill. Cat. Noctuidae Korea, 1998: 67; No. 141 (Dysgonia absentimacula).

NOTE. First reported for Korea by Kononenko et al. (1998). No subsequent records after the first report of the species from Korea known. The residence of the species in Korea require confirmation. 263. Dysgonia mandschuriana (Staudinger, 1892)



Genus DYSGONIA Hübner, [1823] 1816 264. Dysgonia dulcis (Butler, 1878) � genit.: Pl. 43, fig. 3; ♀ genit.: Pl. 207, fig. 1 Ill. Cat. Noctuidae Korea, 1998: 67; No. 143.

265. Dysgonia obscura (Bremer & Grey, 1853)


Dysgonia coreana (Leech, 1889), syn. n. (Ill. Cat. Noctuidae Korea, 1998: 68; No. 145). Ill. Cat. Noctuidae Korea, 1998: 67; No. 144 (Dysgonia obscura ); 68; No. 145 (Dysgonia coreana).

NOTE. The examinations of numerous material from different parts of distribution of Dysgonia obscura and Dysgonia coreana have shown no differences in the male genitalia structures and revealed their conspecifity. The coreana is the form of Dysgonia obscura with uncontrast wings pattern. Both forms occur sympatrically and synchronously. 56

Genus GRAMMODES Guenée, 1852 (Type species: Noctua geometrica Fabricius, 1775 [India]). 266. Grammodes geometrica (Fabricius, 1775)* � genit.: Pl. 43, fig. 5; ♀ genit.: Pl. 298, fig. 1 Noctua geometrica Fabricius, 1775, Ent. Syst.: 599 (Noctua) (TL: India oriental). Phalaena ammonia Cramer, 1870, Uitl. Kapellen 3: 98, Pl. 250: D (Phalaena) (TL: India, “Coté de Coromandel”), nec Esper, [1794]). Grammodes orientalis Warren, 1913, in Seitz, Macrolepid. World 3: 331, Pl. 61d (TL: China; India [BMNH, London]). Ill. Cat. Noctuidae Korea, 1998 _____. Sohn and Kim, 2003.

DISTRIBUTION. Korea, China, Taiwan, Japan, Philippines, Indonesia (Sumatra, Borneo, Sulawesi), Thailand, Laos, Cambodia, Vietnam, Myanmar, India, Nepal, Pakistan, Australia, Afrika south of Sahara, Ghana, Madagascar. Genus ACHAEA Hübner [1823] 267. Achaea janata (Linnaeus, 1758) � genit.: Pl. 44, fig. 1; ♀ genit.: Pl. 207, fig. 3 Ill. Cat. Noctuidae Korea, 1998: 65; No. 136.

Genus LACERA Guenée, 1852 268. Lacera procellosa Butler, 1879 � genit.: Pl. 44, fig. 2; ♀ genit.: Pl. 298, fig. 2 Ill. Cat. Noctuidae Korea, 1998: 88; No. 202.

NOTE. The female specimen was not available for dissection. Genus OXYODES Guenée, 1852 269. Oxyodes scrobiculata (Fabricius, 1775) � genit.: Pl. 44, fig. 3; ♀ genit.: Pl. 207, fig. 4 Ill. Cat. Noctuidae Korea, 1998: 93; No. 215.

NOTE. The record of O. scrobiculata in Korea is based on a specimen preserved in the Staudinger collection (MNHU, Berlin), labelled as “Korea”, without exact data. The occurring of the species in Korea is required for the confirmation. Tribe CATOCALINI Boisduval, [1828] Genus CATOCALA Schrank, 1802 270. Catocala lara Bremer, 1861 � genit.: Pl. 44, fig. 4; ♀ genit.: Pl. 207, fig. 5 Ill. Cat. Noctuidae Korea, 1998: 54; No. 97.

271. Catocala nivea Butler, 1877



272. Catocala fraxini (Linnaeus, 1758)



273. Catocala nupta (Linnaeus, 1767)



274. Catocala adultera Ménétriés, 1856

� genit.: Pl. 45, fig. 3; ♀ genit.: Pl. 208, fig. 2 57


275. Catocala electa (Vieveg, 1790)



276. Catocala dula Bremer, 1861



277. Catocala patala Felder & Rogenhofer, 1874 � genit.: Pl. 46, fig. 2; ♀ genit.: Pl. 208, fig. 5 Ill. Cat. Noctuidae Korea, 1998: 57; No. 104.

278. Catocala obscena Alphéraky, 1879



279. Catocala paki Kishida, 1981



280. Catocala jonasii Butler, 1877



281. Catocala doerriesi Staudinger, 1888



282. Catocala musmi (Hampson, 1913) Ill. Cat. Noctuidae Korea, 1998: 58; No. 109.

NOTE. Described by Hampson from Wonsan [Gensan], North Korea. No subsequent records from Korea are available after original description. 283. Catocala abamita Bremer & Grey, 1853



284. Catocala separans Leech, 1889



285. Catocala helena Eversmann, 1856



286. Catocala puella Leech, 1889



NOTE. Female unknown. 287. Catocala deuteronympha Staudinger, 1861



288. Catocala fulminea (Scopoli, 1763) 58



289. Catocala eminens Staudinger, 1892



290. Catocala koreana Staudinger, 1892



291. Catocala proxeneta Alphéraky, 1895



292. Catocala danilovi (Bang-Haas, 1927)



293. Catocala praegnax Walker, 1858



294. Catocala streckeri Staudinger, 1888



295. Catocala agitatrix Graeser, 1888 [1889]



296. Catocala bella Butler, 1877



297. Catocala ella Butler, 1877



298. Catocala duplicata Butler, 1885



299. Catocala nubila Butler, 1881



300. Catocala dissimilis Bremer, 1861



301. Catocala nagioides Wileman, 1924



302. Catocala actaea Felder & Rogenhofer, 1874 � genit.: Pl. 51, fig. 4; ♀ genit.: Pl. 211, fig. 3 Ill. Cat. Noctuidae Korea, 1998: 63; No. 129.

303. Catocala pirata (Herz, 1904)



59

304. Catocala bokhaica (Kononenko, 1979)



Genus CORTYTA Walker, [1858] 1857 305. Cortyta grisea (Leech, 1900) � genit.: Pl. 52, fig. 1; ♀ genit.: Pl. 211, fig. 6 Ill. Cat. Noctuidae Korea, 1998: 71; No. 156.

NOTE. The generic relegation of “Cortyta” grisea, involved into an eremial African-west Palaearctic genus Cortyta seems to be doubtful. Tribe SYPNINI Holloway, 2005 Genus HYPERSYPNOIDES Berio, 1958 306. Hypersypnoides astrigera (Butler, 1885) � genit.: Pl. 52, fig. 2; ♀ genit.: Pl. 211, fig. 7 Ill. Cat. Noctuidae Korea, 1998: 91; No. 209.

307. Hypersypnoides submarginata (Walker, 1865)* � genit.: Pl. 52, fig. 3; ♀ genit.: Pl. 298, fig. 3 Tavia submarginata Walker, 1865, List Specimens lepid. Insects Colln. Br. Mus. 33: 941 (TL: India, Hindustan [MNHU, Berlin]). Sypna infrapicta Strand, 1917, Arch. Naturgesh. 83(A) 10: 140 (TL: Taiwan, Suisharyo [DEI, Eberswalde]). Ill. Cat. Noctuidae Korea, 1998 _____. Sohn, Ronkay and Kononenko, 2004.

DISTRIBUTION. Korea, Japan, China, Taiwan, India, Indonesia. Genus SYPNOIDES Hampson, 1913 Subgenus SUPERSYPNOIDES Berio, 1958 308. Sypnoides picta (Butler, 1877) � genit.: Pl. 52, fig. 4; ♀ genit.: Pl. 211, fig. 8 Ill. Cat. Noctuidae Korea, 1998: 90; No. 205.

309. Sypnoides amplifascia (Warren, 1913)



NOTE. Male specimen was unavailable for examination. The distribution of this species in Korea is required with further material including the male. 310. Sypnoides hercules (Butler, 1881)



311. Sypnoides fumosa (Butler, 1877)



Genus DADDALA Walker, 1865 312. Daddala lucilla (Butler, 1881) � genit.: Pl. 53, fig. 1; ♀ genit.: Pl. 212, fig. 4 Ill. Cat. Noctuidae Korea, 1998: 91; No. 210.

60

Tribe HYPOCALINI Guenée, 1852 Genus HYPOCALA Guenée, 1852 313. Hypocala deflorata (Fabricius, 1794) � genit.: Pl. 53, fig. 2; ♀ genit.: Pl. 212, fig. 5 Ill. Cat. Noctuidae Korea, 1998: 87; No. 198.

314. Hypocala violacea Butler, 1879



315. Hypocala subsatura Guenée, 1852



316. Hypocala rostrata (Fabricius, 1794)



Tribe TINOLINI Moore, [1885] Genus CALESIA Guenée, 1852 (Type species: Calesia comosa Guenée, 1852 [Bangladesh: Silhet]). a junior subjective synonym of Erebus dasypterus Kollar, 1844). PASIPEDA Walker, 1858 (Type species: Pasipeda rufipalpis Walker, 1858 [Ceylon]). MECYRA Walker, 1865 (Type species: Mecyra invaria Walker, 1865 [Indonesia, Java], a junior subjective synonym of Calesia gastropachoides Guenée, 1852). ZIRONA Walker, 1869 (Type species: Zirona marginata Walker, 1869 [Congo]). ASYMBATA Gerstaecker, 1871 (Type species: Asymbata roseiventris Gerstaecker, 1871 [Kenya]), a junior subjective synonym of Calesia zambesita Walker, 1865). LECASIA Fawcett, 1916 (Type species: Lecasia othello Fawcett, 1916 [Kenya: Kedai, South Africa: Durban]).

317. Calesia dasypterus (Kollar, 1844)*


Erebus dasypterus Kollar, 1844, In Hügel, Kaschmir und das Reich Siek, 4: 476 (TL: “Himalaya”, [Nepal?]). Erebus leucostigma Kollar, [1844], in Hügel, Kaschmir und das Reich Siek, 4: 476 (TL: “Himalaya” [BMNH, London]). Noctua comosa Guenée, 1852, in Boisduval and Guenée, Hist. nat. Insectes (Lépid.) 7: 258, Pl. 21: 7 (TL: Bangkadesh, Silhet). Noctua stigmaleuca Guenée, 1852, in Boisduval and Guenée, Hist. nat. Insectes (Lépid.) 7: 258 (TL: Central India). dasyptera; auct., nec Kollar, 1844. Ill. Cat. Noctuidae Korea, 1998 _____. Sohn and Ronkay, 2001.

DISTRIBUTION. Korea, South China, Taiwan, Thailand, Laos, Vietnam, Nepal, India, Bangladesh. NOTE. Judging from the general distribution of the species, its distribution in Korea seems to be doubtful and could be resulting of mislabelling. The occuring of the species in Korea is required with a further study. Genus FODINA Guenée, 1852 (Type species: Fodina oriolus Guenée, 1852 [Bangladesh: Silhet]). ANOCALA Scott, 1891, unavail. publ. syn.

318. Fodina contigua Wileman, 1914*

♀ genit.: Pl. 213, fig. 1 61

Fodina contigua Wileman, 1914, Entomologist, 47: 221 (TL: Taiwan, Kanshirei [BMNH, London]). Fodina stola f. formosenis Strand, 1914, Arch. Naturgesh. 80A(2): 58 (TL: Taiwan [DEI, Eberswalde]). Fodina contigua ab. fusca Wileman, 1914, Entomologist, 47: 221 (TL: Taiwan, Kanshirei [BMNH, London]). Fodina kosemponis Strand, 1919, Arch. Naturgesh. 83A(10): 144 (TL: Taiwan, Kosempo, Shisa [DEI, Eberswalde]). Fodina antemedia Strand, 1919, Arch. Naturgesh. 83A(10): 144 (TL: Taiwan, Kosempo [DEI, Eberswalde]). Ill. Cat. Noctuidae Korea, 1998 _____. Sohn and Ronkay, 2001.

DISTRIBUTION. Korea, South China, Taiwan, Thailand, Laos, Vietnam, Buthan, India, Sri Lanka. NOTE. Once reported for Korea from Gangwong province. Judging from general distribution of the species its distribution in Korea seems to be doubtful and could be resulting of mislabelling. The occuring of the species in Korea is required with a further study. Tribe HULODINI Guenée, 1852 (SPEIREDONIINAE Swinhoe, 1900) Genus ERICEIA Walker, [1858] 1857 (Type species: Ericeia sobria Walker, [1858] 1857 [Australia: Quisland, Moreton Bay]). VILLOSA Koch, 1865 (Type species: Villosa leichardtii Koch, 1865 [Australia]). GIRPA Walker, 1858 (Type species: Girpa aliena Walker, 1858 [Australia: Quisland, Moreton Bay]). EREIA; Turner, 1933, emend.

319. Ericeia pertendens (Walker, 1858)*


Remigia pertendens Walker, 1858, List Specimens lepid. Insects Colln. Br. Mus. 14: 1512 (TL: Sri Lanka [BMNH, London]). Ericeia gonioptila Prout, 1922, Bull. HIll. Mus. 1: 233, Pl. 21: 2 (TL: Indonesia, Central Ceram, Manusela [BMNH, London]). Ericeia eurytaenia Prout, 1929, Bull. HIll. Mus. 3: 113 (TL: Sri Lanka, Haldumulla [BMNH, London]). Ericeia occidua Prout, 1929, Bull. HIll. Mus. 3: 115 (TL: West China, Teng-yueh-Ting [BMNH, London]). Ericeia iopolia Fletcher, 1957, Nat. Hist. Renell I., Br. Solomon Isl. 2: 50, fig. 65, 66 (TL: Solomon Isl., Rennell Isl., Kasipa Hill [BMNH, London]). Ill. Cat. Noctuidae Korea, 1998 _____. Sohn, Ronkay, Kim and Cho, 2005.

DISTRIBUTION. Korea, China, Taiwan, Japan, Philippines, Vietnam, Laos, Thailand, Malaysia, Nepal, India, Sri Lanka, Indonesia, Pacific Islands. Tribe PANDESMINI Wiltshire, 1990 Genus POLYDESMA Boisduval, 1833 320. Polydesma boarmoides Guenée, 1852 � genit.: Pl. 54, fig. 2; ♀ genit.: Pl. 213, fig. 2 Ill. Cat. Noctuidae Korea, 1998: 93; No. 214.

Tribe ERCHEINI Berio, 1992 Genus ERCHEIA Walker, [1858] 321. Ercheia umbrosa Butler, 1881 � genit.: Pl. 54, fig. 3; ♀ genit.: Pl. 213, fig. 3 Ill. Cat. Noctuidae Korea, 1998: 70; No. 153.

62

322. Ercheia niveostrigata Warren, 1913



323. Ercheia dubia (Butler, 1874) Ill. Cat. Noctuidae Korea, 1998: 71; No. 155.

NOTE. The specimens were unavailable for genitalia dissection. Miscellaneous genera Genus DINUMMA Walker, 1858 324. Dinumma deponens Walker, 1858 � genit.: Pl. 54, fig. 6; ♀ genit.: Pl. 212, fig. 9 Ill. Cat. Noctuidae Korea, 1998: 95; No. 219.

Genus ERYGIA Guenée, 1852 325. Erygia apicalis Guenée, 1852 � genit.: Pl. 54, fig. 5; ♀ genit.: Pl. 213, fig. 5 Ill. Cat. Noctuidae Korea, 1998: 89; No. 203.

Genus ISCHYJA Hübner, [1823] 1816 (Type species: Phalaena manlia Cramer, 1776 [India: Coromandel Coast]). POTAMORPHA Guenée, 1852 (Type species: Phalaena manlia Cramer, 1776 [India]). ISCHYIA; Agassiz, [1846], emend.

326. Ischyja manlia (Cramer, 1776)*


Phalaena, Noctua manlia Cramer, 1776, Uitl. Kapellen 1: 144, Pl. 92: A (TL: India, Cote de Coromandel). Noctua squalida Fabricius, 1787, Mant. Ins.: 136 (TL: India, Coromandel). Potamophora amboinensis Felder, 1862, Sitzungsb. kaisl. Akad. wiss. 43(1): 42 (TL: Indonesia, Amboina). Ill. Cat. Noctuidae Korea, 1998 _____. Sohn, Choi and Cho, 2005.

DISTRIBUTION Korea, Japan, China, Taiwan, Philippines, Indonesia, Malaysia, Thailand, Cambodia, Vietnam, Nepal, India, Sri Lanka. Genus BLASTICORHINUS Butler, 1893 327. Blasticorhinus hoenei Berio, 1956 � genit.: Pl. 55, fig. 2; ♀ genit.: Pl. 214, fig. 1 Ill. Cat. Noctuidae Korea, 1998: 92; No. 212.

328. Blasticorhinus ussuriensis (Bremer, 1861)


Ill. Cat. Noctuidae Korea, 1998: 92; No. 211 (Blasticorhinus ussuriensis, misidentification of B. unduligera).

DISTRIBUTION Korea, Japan, China, RFE (Primorye and Khabarovsk terr., Amur reg.). NOTE. The examination of the types of ussuriensis [ZIN, St.-Petersburg] and unduligera [BMNH, London] revealed their non-conspecifity. The male specimen, illustrated in the Catalogue of Korean Noctuidae as 211. Blasticorhinus ussuriensis in fact is the male of B. unduligera and the female, illustrated as 213. Blasticorhinus rivulosa is the female of B. unduligera. These species are represented sibling species which hardly distinguishable by external characters. Boths species are sympatrically distributed in Korea, North China and Russian Far East. Here we illustrate authentical male and female genitalia figurs. 63

329. Blasticorhinus unduligera (Butler, 1878), syn. rev. � genit.: Pl. 55, fig. 3; ♀ genit.: Pl. 213, fig. 8 Azazia unduligera Butler, 1878. Ann. Mag. nat. Hist. (5)1: 293 (TL: Japan, Yokohama [BMNH, London]). Ill. Cat. Noctuidae Korea, 1998: 92; No. 211 (Blasticorhinus ussuriensis, misidentification); 92; No. 213 (Blasticorhinus rivulosa, misidentification).

NDISTRIBUTION Korea, Japan, China, RFE (Primorye terr.). NOTE. The name unduligera is not a synonym of Blasticorhinus ussuriensis, but applicable to a distinct species. See also note for preceding species. Genus HYPOSPILA Guenée 330. Hypospila bolinoides Gunenée, 1852 � genit.: Pl. 55, fig. 5; ♀ genit.: Pl. 214, fig. 2 Ill. Cat. Noctuidae Korea, 1998: 96; No. 221.

Genus PREMUSIA Walker, 1858 (Type species: Premusia intrahens Walker, 1858 [Borneo: Sarawak]) DYSEDIA Rogenhofer, 1874 (Type species: Dysedia zibellina Felder & Rogenhofer, 1874 [Borneo: Sarawak], a junior subjective synonym of Premusia intrahens Walker, 1858). HERCHUNDA Swinhoe, 1900 (Type species: Premusia hirtissima Walker, [1863] 1864 [Borneo: Sarawak]). REMUSIA; Walker, [1863] 1864, misspelling.


331. Premusia intrahens Walker, 1858*

Premusia intrahens Walker, 1858, List Specimens lepid. Insects Colln. Br. Mus. 15: 1780 (TL: Borneo, Sarawak). Dysedia zibellina Felder & Rogenhofer, 1874, Reise öst. Fregatte Novara (Zool.) 2: Pl. 112: 8 (TL: Borneo, Sarawak [BMNH, London]). Premusia smaragdina Walker, 1858, List Specimens lepid. Insects Colln. Br. Mus. 15: 1811 (TL: Borneo, Sarawak [BMNH, London]). Ill. Cat. Noctuidae Korea, 1998 _____ Han, Sohn and Weon, 2006.

DISTRIBUTION. Thailand, Vietnam, Indonesia (Sumatra, Borneo). NOTE. The distribution of the species in Korea is required a reconfirmation. Genus MECODINA Guenée, 1852 332. Mecodina subviolacea (Butler, 1881)



NOTE. The female specimen was not available for dissection. � genit.: Pl. 56, fig. 1

333. Mecodina subcostalis (Walker, 1865) Ill. Cat. Noctuidae Korea, 1998: 97; No. 223.

NOTE. The female specimen was not available for dissection. 334. Mecodina nubiferalis (Leech, 1889)



335. Mecodina cineracea (Butler, 1879) Ill. Cat. Noctuidae Korea, 1998: 97; No. 225.

64


NOTE. The female specimen was not available for dissection. Genus HYPOSEMANSIS Hampson, 1895 336. Hyposemansis albipuncta (Wileman, 1914) � genit.: Pl. 56, fig. 4; ♀ genit.: Pl. 214, fig. 5 Ill. Cat. Noctuidae Korea, 1998: 96; No. 220.

Subfamily STICTOPTERINAE Genus LOPHOPTERA Guenée, 1852 (Type species: Lophoptera squammigera Guenée, 1852 [Australia]) EVIA Walker, 1863 (Type species: Evia ferrinalis Walker, 1863 [Borneo: Sarawak]). CIASA Walker, 1864 (Type species: Ciasa pustulifera Walker, [1863] 1864 [Borneo: Sarawak]). SADARSA Moore, 1882 (Type species: Sadarsa longtpennis Moore, 1882 [India: Darjiling]).


337. Lophoptera squammigera Guenée, 1852*

Lophoptera squammigera Guenée, 1852, In Boisduval and Guenée, Hist. nat. Insectes (Lépid.) 7: 55, Pl. 14: 13 (TL: Australia, New Holland). Lophoptera costata Moore, [1885], Lepid. Ceylon, 3: 123 (TL: Sri Lanka [BMNH, London]). Ill. Cat. Noctuidae Korea, 1998 _____. Sohn, Ronkay and Choi, 2005.

DISTRIBUTION. Korea, China, Taiwan, Japan, Philippines, Thailand, Myanmar, Laos, Vietnam, Nepal, Pakistan, India, Sri Lanka, Indonesia, Pacific Islands, Australia.

Subfamily EUTELIINAE Grote, 1882 Genus EUTELIA Hübner, [1823] 1816 (Type species: Noctua adulatrix Hübner, [1813] [Europe]). 338. Eutelia geyeri (Felder & Rogenhofer, 1874) � genit.: Pl. 56, fig. 5; ♀ genit.: Pl. 218, fig. 4 Ill. Cat. Noctuidae Korea, 1998: 134; No. 336.

339. Eutelia adulatricoides (Mell, 1943)



340. “Eutelia” hamulatrix (Draudt, 1950)



NOTE. The species is not congeneric with Eutelia adulatrix Hübner, [1813], the type species if the genus Eutelia, nor with E. geyeri and E. adulatricoides, its systematic position is uncertain. Genus ATACIRA Swinhoe, 1900 341. Atacira grabczewskii (Püngeler, 1903) � genit.: Pl. 57, fig. 2; ♀ genit.: Pl. 218, fig. 7 Ill. Cat. Noctuidae Korea, 1998: 135; No. 339.

Genus ANUGA Guenée, 1852 342. Anuga multiplicans (Walker, 1858) � genit.: Pl. 57, fig. 3; ♀ genit.: Pl. 218, fig. 8 65


Subfamily NOLINAE Bruard, 1846 Tribe NOLINI Bruand, 1846 (By H. S. Oh and V. S. Kononenko) Genus NOLA Leach, [1815] 1830 (Type-species: Noctua palliola [Denis & Schiffermüller], 1775 [Austria]) ROESELIA Hübner, [1825] 1816 (Type-species: Phalaena cucullatella Linnaeus, 1758 [Europe]). CELAMA Walker, 1865 (Type-species: Celama liparisalis Walker, 1865 [Borneo, Saravak]. LIRA Billberg, 1820, Proposed, unnecessarily as an objective replacement name for Nola Leach, [1815]. CHLAMIFERA Hübner, 1822 (Type-species: Noctua palliola [Denis & Schiffermüller], 1775 [Europe]). DIMONA Walker, 1858 (Type-species: Dimona porrigens Walker, 1858 [Australia]). SELCA Walker, [1866] 1865 (Type-species: Seica latifascialis Walker, [1866] 1865 [Borneo: Sarawak]). LEBENA Walker, 1866 (Type-species: Lebena trinotata Walker, 1866 [North America]). MINNAGARA Walker, 1866 (Type-species: Minnagara fasciata Walker, 1866 [Moluccas]). SOROCOSTIA Rosenstock, 1885 (Type-species: Eromene vetustella Walker, 1866 [Australia]). EPIZEUCTIS Meyrick, 1889 (Type-species: Nola innocua Butler, 1880 [Taiwan]). STENOLA Möschler, 1890 (Type-species: Stenola bistriga Möschler, 1890 [Puerto Rico]). POLIOTHRIPA Hampson, 1902 Type-species: Poliothripa niphostena Hampson, 1902.

NOTE. Choi (2006) incorrectly applied the generic name Roeselia to Meganola and the generic name Celama to Nola. 343. Nola cristatula (Hübner, 1793)*


Phalaena cristatula Hübner, 1793, Samml. auserlesener Vögel Schmett. Pl. 34 (TL: Europe). Nola minualis Leech, 1888, Proc. zool. Soc. London: 608, Pl. 30: 17 (TL: Japan [BMNH]). Ill. Cat. Noctuidae Korea, 1998 _____. Oh, 1991; 2001.

DISTRIBUTION. Korea, Japan (Honshu, Shikoku, Kyushu), European part of Russia, Central and Southern Europe. 344. Nola taeniata Snellen, 1875*


Nola taeniata Snellen, 1875, Tijdschr. ent. 17: 65, Pl. 6: 1 (TL: Celebes [RNH, Leiden]). Nola candida Butler, 1879, Illustr. typical Specimens Lepid. Heterocera Colln Br. Mus. 3: 9, Pl. 30: 17 (TL: Japan [BMNH, London]). Roeselia fragilis Swinhoe, 1890. Sorocostia mesozona Lucas, 1890. Celama anpingicola Strand, 1917. Ill. Cat. Noctuidae Korea, 1998 _____. Oh, 1991; 2001.

DISTRIBUTION. Korea, Japan, China, South of RFE (Primorye terr.), Taiwan, Indonesia, Sri Lanka, Malaysia, India, Nepal, Myanmar, Australia. 345. Nola aerugula (Hübner, 1793)*


Phalaena aerugula Hübner, 1793, Samml. auserlesener Vögel Schmett.: 11, Pl. 61 (TL: Europe).

66

Pyralis centonalis Hübner, 1796, Samml. eur. Schmett., Pry., fig. 15. Glaphyra atomosa Bremer, 1861, Bull. Acad. imp. Sci. St.-P(tersb. 3: 491 (TL: RFE, Primorye terr. [ZIN, St.Petersbourg]). Nola candidalis Staudinger, 1861, in Romanoff, Mém. Lépid. 6: 258 (TL: “Amur” [MNHU, Berlin]). Ill. Cat. Noctuidae Korea, 1998 _____. Oh, 1991, 2001.

DISTRIBUTION. Korea, Japan (Hokkaido, Honshu), NE China, RFE (Amur reg, Khabarovsk and Primorie territory, Sakhalin, South Kuriles), Transbaikalia, South Siberia, Ural, European part of Russia, Europe (from England and Spain to South Fennoscandia). 346. Nola yoshinensis (Wileman & West, 1929)*


Celama yoshinensis Wileman & West, 1929, Ann. Mag. nat. Hist. (10) 3: 187 Ill. Cat. Noctuidae Korea, 1998 _____. Oh, 2001, 1991 _____.

DISTRIBUTION. Korea, Japan (Honshu, Shikoku, Kyushu). 347. Nola japonibia Strand, 1920*


Celama innocua var. japonibia Strand, 1920, Lep. Cat. 24: 454 (TL: Japan [DEI. Eberswalde]). Ill. Cat. Noctuidae Korea, 1998 _____. Oh, 2001.

DISTRIBUTION. Korea (Central, South), RFE (Primorye terr.), Japan (Hokkaido, Honshu, Shikoku, Kyushu). 348. Nola confusalis (Herrich-Schüffer, [1847] 1845)* � genit.: Pl. 299, fig. 3; ♀ genit.: Pl. 302, fig. 7 Roeselia confusalis Herrich-Schüffer, [1847] 1845, Syst. Bearb. Schmett. Eur. 2: 164 (TL: [Europe] not stated). Celama confusalis ssp. fumosensis Daniel, 1957. Ill. Cat. Noctuidae Korea, 1998 _____. Oh, 1991; 2001.

DISTRIBUTION. Korea, Japan (Hokkaido, Honshu), RFE (Amur, Khabarovsk and Primorye terr., Sakhalin, S Kuriles), Transbaikalia, Ural, European part of Russia, Europe. 349. Nola longicosta Oh, 2001*


Nola longicosta Oh, 2001, Ins. Koreana, 18(2): 131, figs 7, 19, 31, 40 [CIS. Kangwon]. Ill. Cat. Noctuidae Korea, 1998 _____. Oh, 1991; 2001.

DISTRIBUTION. Korea (Central). 350. Nola nami (Inoue, 1956)*


Celama nami Inoue, 1956, Kontyû 24: 158, Pl. 14: 2, 5. Ill. Cat. Noctuidae Korea, 1998 _____. Oh, 1991; 2001.

DISTRIBUTION. Korea, Japan (Hokkaido, Honshu, Shikoku, Kyushu), RFE (S Kuriles, Kunashir I.). 351. Nola ebatoi Inoue, 1970*


Celama ebatoi Inoue, 1970, Bull. Japan ent. Acad. 6: 2, Pl. 1: 22-24 (TL: Japan, Tokyo [BMNH, London]). Ill. Cat. Noctuidae Korea, 1998 _____. Oh, 1991; 2001.

DISTRIBUTION. Korea, Japan (Honshu, Shikoku, Kyushu). 67

352. Nola innocua Butler, 1880*


Nola innocua Butler, 1880, Proc. zool. Soc. Lond. 1880: 671 (TL: [Taiwan] Formosa [BMNH]). Ill. Cat. Noctuidae Korea, 1998 _____. Oh, 1991; 2001.

DISTRIBUTION. Korea (South), RFE (Primorye, Sakhalin, S Kuriles: Kunashir I.), Japan (Honshu, Shikoku, Kyushu, Tsushima, Okinawa), Taiwan 353. Nola okanoi (Inoue, 1958)*


Celama okanoi Inoue, 1958, Kontyû 26: 233, figs 1, 6 (TL: Japan, Morioka [BMNH, London]). Ill. Cat. Noctuidae Korea, 1998 _____. Oh, 1991; 2001.

DISTRIBUTION. Korea (Central), Japan (Honshu). 354. Nola trilinea Marumo, 1923*


Nola trilinea Marumo, 1923, J. Coll. Agric. Tokyo imp. Univ. 8: 137, pl. 3: 1. Ill. Cat. Noctuidae Korea, 1998 _____. Oh, 1991; 2001.

DISTRIBUTION. Korea (Central), Japan (Honshu, Shikoku, Kyushu, Okinawa). POECILONOLA Hampson, 1900 (Type-species: Seica plagiola Hampson, 1898 [Sri Lanka]) 355. Poecilonola pulchella (Leech, 1889)* � genit.: Pl. 300, fig. 5; ♀ genit.: Pl. 304, fig. 4 Acontia pulchella Leech, 1889, Proc. zool. Soc. Lond. 1889: 524, Pl. 53: 10 (TL: Japan, Ohoyama [BMNH, London]). Ill. Cat. Noctuidae Korea, 1998 _____. Oh, 1991.

DISTRIBUTION. Korea, Japan (Honshu, Kyushu, Tsushima), China, RFE (Primorye terr.). Genus MEGANOLA Dyar, 1898 (Type-species: Meganola conspicua Dyar, 1898 [USA, Texas; Colorado; Arizona]). RHYNCHOPALPUS Hampson, 1893 (Type-species: Aglossa argentalis Moore, 1867 [India]).

NOTE. The present list of Meganola species is based on the thesis of Oh (1991). Except below listed species, Oh (1991) refered three species as a new for the science. Because of the material was unavailable for our study, the descripton of a new species is not placed here, they will be describe separately. Choi (2006) incorrectly apply the name Roeselia to this group. The type species of Roeselia is Nola cucullatela, which belongs to the genus Nola, but not congeneric with the members of Meganola. 356. Meganola microphasma (Butler, 1885)*


Meganola microphasma Butler, 1885, Cistula ent. 3: 117 (TL, Japan, Yokohama [BMNH, London]). Ill. Cat. Noctuidae Korea, 1998 _____. Oh, 1991.

DISTRIBUTION. Korea, Japan (Honshu, Shikoku, Kyushu). 357. Meganola banghaasi (West, 1925)*


Nola banghaasi West, 1925, Ann. Mag. nat. Hist. (10) 3: 194 (TL: RFE, Primorye [Ussuri] [BMNH, London]). Nola banghaasi sumi Inoue, 1956, Kontyû (24) 3: 158, Pl. 14: 3, 6 (TL: Japan [BMNH, London]). Ill. Cat. Noctuidae Korea, 1998 _____. Oh, 1991.

68

DISTRIBUTION. Primorye. Korea, Japan (Yaku). 358. Meganola major (Hampson, 1891)*


Nola major Hampson, 1891, Ill. typ. Spec. lep. Het. B.M. 8: 48, Pl. 139: 13 (TL: India, Nilgiris [BMNH, London). Ill. Cat. Noctuidae Korea, 1998 _____. Oh, 1991 (as fulvusana, manuscript name).

DISTRIBUTION. S Korea, Japan, India, Nepal. First report for Korea. 359. Meganola gigantoides (Inoue, 1961)*


Roeselia gigantoides Inoue, 1961, Check List Lepid. Japan 6: 682 (TL: Japan, Gumma [BMNH, London]). Ill. Cat. Noctuidae Korea, 1998 _____.

DISTRIBUTION. Korea (Central), Japan (Honshu). � genit.: Pl. 305, fig. 7

360. Meganola mediofascia (Inoue, 1958)*

Roeselia mediofascia Inoue, 1958, Kontyû 26: 236, 3, 8 (TL: Japan, Yamanshi [BMNH, London]). Ill. Cat. Noctuidae Korea, 1998 _____. Oh, 1991 (as fuscofasciata, manuscript name).

DISTRIBUTION. Korea, Japan. First report for Korea. 361. Meganola strigulosa (Staudinger, 1887)*


Nola strigulosa Staudinger, 1887, in Romanoff, Mém. Lépid. 3: 180, Pl. 10: 4 [MNHU, Berlin]. Roeselia satoi Inoue, 1970, Bull. Jap. ent. Acad. 6: 4, Pl. 1: 19-21 (TL: Japan, Mt. Akiba [BMNH, London]). Ill. Cat. Noctuidae Korea, 1998 _____. Oh, 1991.

DISTRIBUTION. Korea, Japan (Hokkaido, Honshu), RFE (Primorye terr.). � genit.: Pl. 301, fig. 2

362. Meganola bryophilalis (Staudinger, 1887)*

Nola bryophilalis Staudinger, 1887, in Romanoff, Mém. Lépid. 3: 181, Pl. 10: 5 [MNHU, Berlin]. Ill. Cat. Noctuidae Korea, 1998 _____. Oh, 1991.

DISTRIBUTION. Korea, RFE, (Primorye terr.), Japan (Hokkaido, Honshu). 363. Meganola costalis (Staudinger, 1887)*


Nola costalis Staudinger, 1887, in Romanoff, Mém. Lépid. 3: 179, Pl. 10: 3 (TL: RFE, Primorye terr, Askold I. [MNHU, Berlin]). Roeselia melanocosta Inoue, 1961, Check List Lepid. Japan 6: 683 (TL: Japan [BMNH, London]). Ill. Cat. Noctuidae Korea, 1998 _____. Oh, 1991; Choi, 2006.

DISTRIBUTION. Primorye. Japan (Hokkaido, Honshu). 364. Meganola fumosa (Butler, 1879)*


Nola fumosa Butler, 1879, Illustr. typical Specimens Lepid. Heterocera Colln Br. Mus. 3: 9, Pl. 43: 2 (TL: Japan, Yokohama [BMNH, London]). Ill. Cat. Noctuidae Korea, 1998 _____. Oh, 1991.

DISTRIBUTION. Korea, China (Heilongjiang), Japan (Hokkaido, Honshu, Shikoku, Kyushu, Tsushima), RFE (Amur reg., Primorye terr., S Kuriles), Transbaikalia.

69

365. Meganola shimekii Inoue, 1970*


Roeselia shimekii Inoue, 1970, Bull. Japan ent. Acad. 6: 5, Pl. 1: 15-18 (TL: Japan, Mt. Akiba [BMNH, London]). Ill. Cat. Noctuidae Korea, 1998 _____. Oh, 1991.

DISTRIBUTION. Korea, Japan (Honshu, Shikoku, Tsushima), RFE (S Primorye terr.). 366. Meganola subgigas Inoue, 1982*


Meganola subgigas Inoue, 1982, Moths of Japan 1: 665, 2: 343, Pl. 229 45, 351: 10, 353: 8 (TL: Japan, Hokkaido, Oshima, Nakayama Pass [BMNH, London]) Ill. Cat. Noctuidae Korea, 1998 _____. Oh, 1991.

DISTRIBUTION. Korea, China (Heilongjiang, Jiangxi), Japan (Hokkaido), RFE (S Primorye terr.). 367. Meganola albula ([Den. & Schiff., 1775)*


Noctua albula [Denis & Schiffermüller, 1775], Ankündung syst. Werkes Schmett. Wienergegend: 69 (TL: Austria: Vienna). Ill. Cat. Noctuidae Korea, 1998 _____. Oh, 1991.

DISTRIBUTION. Korea, China, Japan, Taiwan, RFE (Amur, Khabarovsk and Primorye terr.), South Siberia, European part of Russia, Europe. 368. Meganola triangulalis Leech, 1888 [1889]*


Melanographia triangulalis Leech, 1888 [1889], Proc. zool. Soc. London 1888: 608, Pl. 31: 12 (TL: Japan [BMNH]). Ill. Cat. Noctuidae Korea, 1998 _____. Oh, 1991.

DISTRIBUTION. Korea (Jeju I.), Japan (Honshu, Shikoku, Kyushu, Tsushima), China, taiwan, Thailand, North India. 369. Meganola gigas (Butler, 1884)*


Nola gigas Butler, 1884, Ann. Mag. nat. Hist. (5) 5: 247 (TL: Japan: Hokkaido [BMNH, London]). Nola maculata Staudinger, 1887, in Romanoff, Mém. Lépid. 3: 180 (TL: RFE: Primorye terr. [MNHU, Berlin]). Roeselia nigromaculata Nagano, 1918, Insect World 22: 414, Pl. 12: 5 (TL: Japan). Ill. Cat. Noctuidae Korea, 1998 _____. Oh, 1991.

DISTRIBUTION. Korea, Japan (Hokkaido, Honshu), China (Heilongjiang, Guangxi), RFE (Midd Amur, Primorye, S Kuriles: Kunashir I.). 370. Meganola protogigas (Inoue, 1979)


Roeselia protogigas Inoue, 1970, Bull. Jap. Ent. Acad. 6: 3, pl. 1: 1 (TL: Japan, Ichibata, Shimane Pref. [BMNH, London]). Ill. Cat. Noctuidae Korea, 1998 _____. Choi, 2006.

DISTRIBUTION Korea, Japan. Genus EVONIMA Walker, 1865 (Type-species: Evonima aperta Walker, 1865 [Java]) MIMERASTRIA Butler, 1881 (Type-species: Erastria mandschuriana Oberthür, 1880 [RFE, Primorye]).

70

371. Evonima mandschuriana (Oberthür, 1880)*


Erastria mandschuriana Oberthur, 1880, Etudes Ent. 5: 83, pl. 2, fig. 9 (TL: RFE, Primorye terr., Askold I. [BMNH, London]). Ill. Cat. Noctuidae Korea, 1998 _____. Oh, 1991.

DISTRIBUTION. Korea, Japan (Hokkaido, Honshu, Shikoku, Kyushu, Tsushima), China (Heilongjiang, Beijing, Henan, Jiangxi, Sichuan), RFE (Amur reg., Khabarovsk terr., Primorye, Sakhalin), South Siberia, North Mongolia. Genus NOLATHRIPA Inoue, 1970 372. Nolathripa lactaria (Graeser, 1892) � genit.: Pl. 57, fig. 5; ♀ genit.: Pl. 214, fig. 6 Ill. Cat. Noctuidae Korea, 1998: 136; No. 341.

Tribe CHLOEPHORINI Staiton, 1859 Subtribe CHLOEPHORINA Staiton, 1859 Genus NEGRITOTHRIPA Inoue, 1970 373. Negritothripa hampsoni (Wileman, 1911) � genit.: Pl. 57, fig. 6; ♀ genit.: Pl. 214, fig. 7 Ill. Cat. Noctuidae Korea, 1998: 137; No. 342.

Genus IRAGAODES Matsumura, 1931 374. Iragaodes nobilis (Staudinger, 1887) � genit.: Pl. 58, fig. 1; ♀ genit.: Pl. 215, fig. 7 Ill. Cat. Noctuidae Korea, 1998: 141; No. 352.

Genus KERALA Moore, 1881 375. Kerala decipiens (Butler, 1878) � genit.: Pl. 58, fig. 2; ♀ genit.: Pl. 215, fig. 8 Ill. Cat. Noctuidae Korea, 1998: 142; No. 356.

Genus GELASTOCERA Butler, 1877 376. Gelastocera exusta Butler, 1877 � genit.: Pl. 58, fig. 3; ♀ genit.: Pl. 216, fig. 1 Ill. Cat. Noctuidae Korea, 1998: 143; No. 357.

377. Gelastocera ochroleucana Staudinger, 1887 � genit.: Pl. 58, fig. 4; ♀ genit.: Pl. 216, fig. 2 Ill. Cat. Noctuidae Korea, 1998:143; No. 358.

378. Gelastocera hallasana Ronkay, 1998



379. Gelastocera eminentissima Bryk, 1948

�genit.: Pl. 58, fig. 6; ♀ genit.: Pl. 216, fig. 3


Genus MACROCHTHONIA Butler, 1881 380. Macrochthonia fervens Butler, 1881 � genit.: Pl. 58, fig. 1; ♀ genit.: Pl. 216, fig. 5 Ill. Cat. Noctuidae Korea, 1998: 144; No. 361.

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Genus PSEUDOIPS Hübner, 1822 381. Pseudoips fagana (Fabricius, 1781) � genit.: Pl. 59, fig. 3; ♀ genit.: Pl. 216, fig. 6 Ill. Cat. Noctuidae Korea, 1998: 145; No. 364.

382. Pseudoips sylpha (Butler, 1879)



Genus PARACRAMA Moore, 1884 383. Paracrama angulata Sugi, 1985



Genus CLETHROPHORA Hampson, 1894 384. Clethrophora distincta (Leech, 1889) � genit.: Pl. 59, fig. 2; ♀ genit.: Pl. 217, fig. 2 Ill. Cat. Noctuidae Korea, 1998: 145; No. 363.

Genus SIGLOPHORA Butler, 1892 385. Siglophora ferreilutea Hampson, 1895



386. Siglophora sanguinolenta (Moore, 1888)



Subtribe SARROTHIPINA Hampson, 1854 Genus CHARACOMA Walker, 1863 387. Characoma ruficirra (Hampson, 1905) � genit.: Pl. 60, fig. 2; ♀ genit.: Pl. 215, fig. 3 Ill. Cat. Noctuidae Korea, 1998: 139; No. 348.

Genus NYCTEOLA Hübner, 1822 388. Nycteola degenerana (Hübner, [1799]) � genit.: Pl. 60, fig. 3; ♀ genit.: Pl. 214, fig. 8 Ill. Cat. Noctuidae Korea, 1998: 140; No. 349.

389. Nycteola asiatica (Krulikowski, 1904)



390. Nycteola coreana (Leech, 1900)


Sarrothripa coreana Leech, 1900, Trans. Ent. Soc. Lond. 1900: 518 (TL: Korea: Wonsan [Gensan] [BMNH]). Nycteola costalis Sugi, 1959, Tinea 5: 279, pl. 38: 13, 14 (TL: Mt. Takao-san, Japan [NIAES, Tsukuba]), syn. n. Ill. Cat. Noctuidae Korea, 1998: 141; No. 351; Park et al., 2001 (Nycteola corena); Sohn, Ronkay, Kim and Cho, 2005 (Nycteola costalis).

DISTRIBUTION. Korea (North, South), Japan (Honshu). NOTE. The taxon has been known only by the type specimen. Park et al. (2001) reported and illustrated one female from N Korea, identified as N. coreana. Judging from the photo of imago of the holotype of N. coreana illustrated by Sugi (1982) and Kononenko et al. (1998), illustration by Park et al. (2001), and the illustrations of the holotype of Nycteola costalis Sugi, 1959 (Sugi, 1959,

72

Sohn, Ronkay, Kim and Cho, 2005), we assume their conspecifity and consider Nycteola costalis Sugi, 1959 as junior synonym (syn. n.) of Nycteola coreana (Leech, 1900). Recently the species was reported for Korea as Nycteola costalis with illustration of male and female genitalia (Sohn et al., 2005). The genitala of the type specimens of N. coreana were unavailable for the revision and illustration. We illustrate here the male and female genitalia citing from Sohn et al. (2005). Subtribe CAMPTOLOMINA Mell, 1943 Genus CAMPTOLOMA Felder, 1874 391. Camptoloma interiorata Walker, [1865] � genit.: Pl. 60, fig. 6; ♀ genit.: Pl. 217, fig. 4 Ill. Cat. Noctuidae Korea, 1998: 148; No. 371.

Subtribe ARIOLICINA Mell, 1943 Genus GABALA Walker, [1866] 1865 392. Gabala argentata Butler, 1878 � genit.: Pl. 60, fig. 1; ♀ genit.: Pl. 217, fig. 6 Ill. Cat. Noctuidae Korea, 1998: 147; No. 369.

Genus ARIOLICA Walker, [1863] 1864 393. Ariolica argentea (Butler, 1881) � genit.: Pl. 61, fig. 1; ♀ genit.: Pl. 217, fig. 5 Ill. Cat. Noctuidae Korea, 1998: 147; No. 368.

394. Sinna extrema (Walker, 1854)

Genus SINNA Walker, 1865 � genit.: Pl. 61, fig. 2; ♀ genit.: Pl. 217, fig. 7


Tribe EARIADINI Hampson, 1912 Genus EARIAS Hübner, [1825] 1816 395. Earias pudicana Staudinger, 1887 � genit.: Pl. 61, fig. 3; ♀ genit.: Pl. 217, fig. 8 Ill. Cat. Noctuidae Korea, 1998: 141; No. 353.

396. Earias roseifera Butler, 1881



397. Earias roseoviridis Sugi, 1982


Earias roseoviridis Sugi, 1982, Moths of Japan 1: 797, 2: 378, pl. 194: 39, 371: 3 (TL: Japan: Amami-Oshima Isl., Sataikuma [NIAES, Tsukuba]). Ill. Cat. Noctuidae Korea, 1998: 141; No. 354, fig. 354b (as synonym of Earias roseifera).

NOTE. The taxon was incorrectly referred in the Catalogue of Lorean Noctuidae (Kononenko et al., 1998) as a synonym of Earias roseifera. Althouth the taxa of the genus Earias are very uniform by the external characters as well as by genitalia, we belive that E. roseiviridis is a distinct species, distributed in South of Japan, Korea and South of the Primorye territory, RFE. The species differs from its allied E. roseifera by larger wingspan, more bright green colouration of forewings and large, well expressed red-pinky patch in the middle part of forewings. In genital structures, E. roseiviridis 73

differs from E. roseifera by larger habitus and proportions of male and female genitalia. In Russian Far East, the moths appear in late summer of beginning of autumn, while E. roseifera flying in the beginning of summer. DISTRIBUTION. Korea, South Japan, South of the Primorie terr., RFE. 398. Earias cupreoviridis (Walker, 1862) Ill. Cat. Noctuidae Korea, 1998: 142; No. 355.

NOTE. The distribution of the species in Korea was confirmed by Park et al. (2001). A series of 18 specimens were collected in Mt. Myohyang-san, N Korea, Pyungyang, Kumgan-san, Mt. Ryongaksan. The specimens were unavailable for dissection. Tribe BLENINI Genus BLENINA Walker, [1858] 1857 399. Blenina senex (Butler, 1878) � genit.: Pl. 61, fig. 6; ♀ genit.: Pl. 215, fig. 6 Ill. Cat. Noctuidae Korea, 1998: 139; No. 347.

Tribe RISOBINI Genus RISOBA Moore, 1881 400. Risoba prominens Moore, 1881



NOTE. The male specimen was unavailable for dissection during preparation of the book. Genus ISCADIA Hampson, 1912 401. Iscadia uniformis (Sugi, 1982) � genit.: Pl. 62, fig. 1; ♀ genit.: Pl. 215, fig. 5 Ill. Cat. Noctuidae Korea, 1998: 138; No. 344.


402. Iscadia pulchra (Butler, 1886) Ill. Cat. Noctuidae Korea, 1998: 138; No. 345.

NOTE. The female specimen was unavailable for dissection during preparation of the book. Tribe ELIGMINI Genus ELIGMA Hübner, [1819] 1816 403. Eligma narcissus (Cramer, 1775) � genit.: Pl. 62, fig. 3; ♀ genit.: Pl. 218, fig. 3 Ill. Cat. Noctuidae Korea, 1998: 137; No. 343.

Subfamily ARCTIINAE Leach, [1815] NOTE. The Arctiidae is downgraded to the subfamily level of the Noctuidae by Lafontaine and Fibiger (2006). The subfamily Arctiinae is not considered in the persent book.

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Subfamily LYMANTRIINAE Hampson, [1893] NOTE. The Lymantriidae is also downgraded to the subfamily level of the Noctuidae by Lafontaine and Fibiger (2006). The subfamily Lymantriinae is not considered in the persent book.

Subfamily PLUSIINAE Boisduval, [1828] Tribe ABROSTOLINI Eichlin & Cunningham, 1978 Genus ABROSTOLA Ochsenheimer, 1816 404. Abrostola triplasia (Linnaeus, 1758) � genit.: Pl. 62, fig. 4; ♀ genit.: Pl. 219, fig. 1 Ill. Cat. Noctuidae Korea, 1998: 115; No. 287.

405. Abrostola ussuriensis Dufay, 1958




406. Abrostola major Dufay, 1957 Ill. Cat. Noctuidae Korea, 1998: 116; No. 289.

NOTE. The female specimen was unavailable for dissection. 407. Abrostola pacifica Dufay, 1960

� genit.: Pl. 63, fig. 2♀ genit.: Pl. 219, fig. 3


408. Abrostola abrostolina (Butler, 1879)



Tribe ARGYROGRAMMATINI Eichlin & Cunningham, 1978 Genus TRICHOPLUSIA McDunnough, 1944 409. Trichoplusia ni (Hübner [1803]) � genit.: Pl. 63, fig. 4; ♀ genit.: Pl. 219, fig. 5 Ill. Cat. Noctuidae Korea, 1998: 118; No. 296.

Genus THYSANOPLUSIA Ichin., 1973 410. Thysanoplusia intermixta (Warren, 1913) � genit.: Pl. 63, fig. 5; ♀ genit.: Pl. 219, fig. 5 Ill. Cat. Noctuidae Korea, 1998: 118; No. 297.

Genus CTENOPLUSIA Duf., 1970 Subgenus CTENOPLUSIA Duf., 1970 411. Ctenoplusia albostriata (Bremer & Grey, 1853) � genit.: Pl. 63, fig. 6; ♀ genit.: Pl. 219, fig. 7 Ill. Cat. Noctuidae Korea, 1998: 119; No. 298.

Subgenus ACANTHOPLUSIA Duf., 1970 412. Ctenoplusia agnata (Staudinger, 1892) � genit.: Pl. 64, fig. 1; ♀ genit.: Pl. 219, fig. 8 Ill. Cat. Noctuidae Korea, 1998: 119; No. 299.

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413. Ctenoplusia ichinosei (Dufay, 1965) Ill. Cat. Noctuidae Korea, 1998: 120; No. 300.

NOTE. The male specimen was unavailable for dissection. Genus PLUSIOPALPA Holland, 1894 414. Plusiopalpa adrasta (Felder & Rogenhofer, 1874)



NOTE. The female specimen was unavailable for dissection. Genus ZONOPLUSIA Chou & Lu, 1979 415. Zonoplusia ochreata (Walker, 1865) Ill. Cat. Noctuidae Korea, 1998: 118; No. 295.

NOTE. The specimens were unavailable for dissection. Genus CHRYSODEIXIS Hübner, [1821] 1816 416. Chrysodeixis eriosoma (Doubleday, 1843)



NOTE. The female specimen was unavailable for dissection. Genus ANADEVIDIA Kostrowicky, 1961 417. Anadevidia peponis (Fabricius, 1775) � genit.: Pl. 64, fig. 4; ♀ genit.: Pl. 220, fig. 2 Ill. Cat. Noctuidae Korea, 1998: 117; No. 292.

418. Anadevidia hebetata (Butler, 1889)



Tribe PLUSIINI Boisduval, [1828] 1829 Subtribe AUTOPLUSIINA Kitching, 1987 Genus ERYTHROPLUSIA Ichinose, 1962 419. Erythroplusia rutilifrons (Walker, 1858) � genit.: Pl. 64, fig. 6; ♀ genit.: Pl. 220, fig. 4 Ill. Cat. Noctuidae Korea, 1998: 125; No. 317.

420. Erythroplusia pyropia (Butler, 1879)



Genus MACDUNNOUGHIA Kostrowicky, 1961 Subgenus MACDUNNOUGHIA Kostrowicky, 1961 421. Macdunnoughia confusa (Stephens, 1850) � genit.: Pl. 65, fig. 2; ♀ genit.: Pl. 220, fig. 6 Ill. Cat. Noctuidae Korea, 1998: 127; No. 321.

422. Macdunnoughia hybrida Ronkay, 1986 Ill. Cat. Noctuidae Korea, 1998: 127; No. 322.

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423. Macdunnoughia crassisigna (Warren, 1913) � genit.: Pl. 65, fig. 4; ♀ genit.: Pl. 220, fig. 8 Ill. Cat. Noctuidae Korea, 1998: 127; No. 323.

Subgenus PURIPLUSIA Chou & Lu, 1974 424. Macdunnoughia purissima (Butler, 1878) � genit.: Pl. 65, fig. 5; ♀ genit.: Pl. 220, fig. 9 Ill. Cat. Noctuidae Korea, 1998: 128; No. 324.

Genus SCLEROGENIA Ichinose, 1973 425. Sclerogenia jessica (Butler, 1878) � genit.: Pl. 65, fig. 6; ♀ genit.: Pl. 221, fig. 1 Ill. Cat. Noctuidae Korea, 1998: 126; No. 319.

Genus ANTOCULEORA Ichinose., 1973 426. Anthoculeora locuples (Oberthür, 1881) � genit.: Pl. 66, fig. 1; ♀ genit.: Pl. 221, fig. 2 Ill. Cat. Noctuidae Korea, 1998: 126; No. 320.

Genus DIACHRYSIA Hübner, [1821] 1816 427. Diachrysia chryson (Esper, 1789) � genit.: Pl. 66, fig. 2; ♀ genit.: Pl. 221, fig. 3 Ill. Cat. Noctuidae Korea, 1998: 123; No. 309.

428. Diachrysia coreae (Inoue & Sugi, 1958)



429. Diachrysia leonina (Oberthür, 1884)



430. Diachrysia bieti (Oberthür, 1884)



431. Diachrysia zosimi (Hübner, [1822])



432. Diachrysia nadeja (Oberthür, 1880)



433. Diachrysia chrysitis (Linnaeus, 1758)



434. Diachrysia stenochrysis (Warren, 1913)


(= tutti Kostrowicky, 1961). Ill. Cat. Noctuidae Korea, 1998: 125; No. 316.

NOTE. For the taxonomic treatment of this species and synonymy, see Goater et al. (2003). Subtribe EUCHALCIINA Choi & Lu, 1979 Genus EUCHALCIA Hübner, [1821] 1816 435. Euchalcia sergia (Oberthür, 1884) � genit.: Pl. 67, fig. 4; ♀ genit.: Pl. 222, fig. 3 77


436. Euchalcia modestoides Poole, 1989


Euchalcia modestoides Poole, 1989, Lepid. Cat. 118(1): 396, replacement name for Euchalcia modesta Hübner, 1786. Phalaena Noctua modesta Hübner, 1786, Beitr. Gesch. Schmett. 1(1): 3, pl. (1), 1: A (TL: Germany: ?Augsburg), nec Cramer, 1777. Ill. Cat. Noctuidae Korea, 1998: 122; No. 306 (Euchalcia cuprea).

NOTE. Koçak (1980) proposed the name Euchalcia cuprea (Esp. [1787]) for E. modesta (Hbn., 1766) as a replacement name, but later Poole (1989) synonymised E. cuprea with E. variabilis (Piller, 1783). In the original description of E. cuprea, the description and the colour picture of the male, are identical with E. modesta. Consequently, Kocak was right, E. cuprea can be considered the first subsequent replacement name of the species, it is not a synonym of E. variabilis as stated by Poole (1989). On the other hand, the specimen described by Esper as the female of E. cuprea in the second part of the section undoubtedly refers to E. variabilis. Hacker (1998a) in his revision of the types of Noctuidae described by Esper designated the lectotype of E. cuprea as the female specimen (i.e. E. variabili). Therefore, contrary to our previous opinion (Kononenko et al., 1998), we accept the latest revision and revalidated synonymy. Genus POLYCHRYSIA Hübner, [1823] 1816 437. Polychrysia esmeralda (Oberthür, 1880) � genit.: Pl. 67, fig. 6; ♀ genit.: Pl. 222, fig. 5 Ill. Cat. Noctuidae Korea, 1998: 121; No. 302.

438. Polychrysia splendida (Butler, 1878)



Genus PANCHRYSIA Hübner, [1821] 1816 439. Panchrysia ornata (Bremer, 1864) � genit.: Pl. 68, fig. 2; ♀ genit.: Pl. 222, fig. 7 Ill. Cat. Noctuidae Korea, 1998: 121; No. 304.

Genus LAMPROTES R. L., 1817 440. Lamprotes mikadina (Butler, 1878) � genit.: Pl. 68, fig. 3; ♀ genit.: Pl. 222, fig. 8 Ill. Cat. Noctuidae Korea, 1998: 123; No. 308.

Genus PLUSIDIA Butler, 1879 441. Plusidia cheiranthi (Tauscher, 1809) � genit.: Pl. 68, fig. 4; ♀ genit.: Pl. 223, fig. 1 Ill. Cat. Noctuidae Korea, 1998: 123; No. 307.

Subtribe PLUSIINA Boisduval, [1828] 1829 Genus AUTOGRAPHA Hübner, [1821] 1816 442. Autographa gamma (Linnaeus, 1758) � genit.: Pl. 68, fig. 5; ♀ genit.: Pl. 223, fig. 2 Ill. Cat. Noctuidae Korea, 1998: 128; No. 325.

443. Autographa nigrisigna (Walker, 1858) Ill. Cat. Noctuidae Korea, 1998: 128; No. 326.

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444. Autographa amurica (Staudinger, 1892)



445. Autographa v-minus (Oberthür, 1884)



NOTE. We consider A. v-minus as a good species distinct from A. amurica. 446. Autographa buraetica (Staudinger, 1892)



447. Autographa mandarina (Freyer, 1846)



448. Autographa excelsa (Kretschmar, 1862)



Genus SYNGRAPHA Hübner, [1821] 1816 449. Syngrapha interrogationis (Linnaeus, 1758) � genit.: Pl. 69, fig. 6; ♀ genit.: Pl. 224, fig. 1 Ill. Cat. Noctuidae Korea, 1998: 132; No. 334.

450. Syngrapha ain (Hochenwarth, 1785)



Genus PLUSIA Ochsenheimer, 1816 451. Plusia festucae (Linnaeus, 1758) � genit.: Pl. 70, fig. 2; ♀ genit.: Pl. 224, fig. 3 Ill. Cat. Noctuidae Korea, 1998: 131; No. 332.

452. Plusia putnami Grote, 1873



Subfamily EUSTROTIINAE Grote, 1882 Genus NARANGA Moore, 1881 453. Naranga aenescens Moore, 1881 � genit.: Pl. 70, fig. 4; ♀ genit.: Pl. 224, fig. 5 Ill. Cat. Noctuidae Korea, 1998: 170; No. 442.

Genus PROTODELTOTE Ueda, 1984 454. Protodeltote pygarga (Hufnagel, 1766) � genit.: Pl. 70, fig. 5; ♀ genit.: Pl. 224, fig. 6 Ill. Cat. Noctuidae Korea, 1998: 159; No. 403.

455. Protodeltote distinguenda (Staudinger, 1888)



456. Protodeltote inexpectata Ueda, 1987



79

457. Protodeltote atricolor Ahn, 1898




458. Protodeltote maculana Ahn, 1898 Ill. Cat. Noctuidae Korea, 1998: 160; No. 407.

NOTE. Female unknown. 459. Protodeltote minima Ahn, 1898



460. Koyaga falsa (Butler, 1885)

Genus KOYAGA Ueda, 1984 � genit.: Pl. 71, fig. 5; ♀ genit.: Pl. 225, fig. 1


461. Koyaga numisma (Staudinger, 1888)




462. Koyaga magnumisma Ahn, 1998 Ill. Cat. Noctuidae Korea, 1998: 161; No. 411.

NOTE. Female unknown. 463. Koyaga virescens (Sugi, 1958)*


Jaspidia virescens Sugi, 1958, Tinea, 4: 225, fig. 7 (TL: Japan, Gumma Pref., Kumanotaira [NIAES, Tsukuba]). Ill. Cat. Noctuidae Korea, 1998 _____. Sohn and Ronkay, 2001.

DISTRIBUTION. Korea, Japan, Taiwan.

464. Sugia stygia (Butler, 1878)

Genus SUGIA Ueda, 1984 � genit.: Pl. 72, fig. 3; ♀ genit.: Pl. 225, fig. 5


465. Sugia idiostygia (Sugi, 1958)



� genit.: Pl. 72

466. Sugia erastroides (Draudt, 1950) Ill. Cat. Noctuidae Korea, 1998: 162; No. 414.

NOTE. The female specimen was unavailable for dissection. Genus DELTOTE R. L., 1817 467. Deltote bankiana (Fabricius, 1775) � genit.: Pl. 72, fig. 6; ♀ genit.: Pl. 225, fig. 7 Ill. Cat. Noctuidae Korea, 1998: 162; No. 415.

468. Deltote uncula (Clerck, 1759) Ill. Cat. Noctuidae Korea, 1998: 163; No. 416.

80


469. Deltote nemorum (Oberthür, 1880)



Genus MICARDIA Butler, 1878 470. Micardia pulchra Butler, 1878 � genit.: Pl. 73, fig. 3; ♀ genit.: Pl. 226, fig. 2 Ill. Cat. Noctuidae Korea, 1998: 164; No. 418.

NOTE. The species was reported from N Korea, Wonsan [Gensan] by Leech (1889a: 535, 1900a: 148). No subsequent records is available since Leech and the occurence of this species in Korea is not reconfirmed yet. � genit.: Pl. 73

471. Micardia argentata Butler, 1878 Ill. Cat. Noctuidae Korea, 1998: 164; No. 419.

NOTE. The species was recorded from North Korea, Wonsan [Gensan] by Leech (1889a: 535, 1900a: 148), neither subsequent data are known, nor specimens from Korea were found in the collections examined. The occurence of this species in Korea, similarly to the preceding species, is not reconfirmed yet. Genus MALIATTHA Walker, 1863 472. Maliattha rosacea (Leech, 1889) � genit.: Pl. 73, fig. 4; ♀ genit.: Pl. 226, fig. 3 Ill. Cat. Noctuidae Korea, 1998: 166; No. 426.

473. Maliattha subrosacea Ahn., 1998



474. Maliattha signifera (Walker, 1857)



475. Maliattha arefacta (Butler, 1879)



476. Maliattha chalcogramma (Bryk, 1948)



477. Maliattha bella (Staudinger, 1888)



478. Maliattha khasanica Zolotarenko & Dubatolov, 1996*


Maliattha khasanica Zolotarenko and Dubatolov, 1996, Atalanta, 26(1-2): 299 (TL: Russia, Primorye, Khasan disrtrict, Furugel’m I.). Ill. Cat. Noctuidae Korea, 1998 _____. Sohn, Ronkay and Kononenko, 2004.

DISTRIBUTION. Korea, RFE (South Primorye).

479. Inabaia culta (Butler, 1879)

Genus INABAIA Sugi, 1982 � genit.: Pl. 74, fig. 6; ♀ genit.: Pl. 227, fig. 1 81


Genus ERASTROIDES Hampson, 1893 480. Erastroides fentoni (Butler, 1881) � genit.: Pl. 75, fig. 1; ♀ genit.: Pl. 227, fig. 2 Ill. Cat. Noctuidae Korea, 1998: 166; No. 433.

Miscellaneous genera Genus NEUSTROTIA Sugi, 1982 481. Neustrotia costimacula (Oberthür, 1880) � genit.: Pl. 75, fig. 2; ♀ genit.: Pl. 227, fig. 3 Ill. Cat. Noctuidae Korea, 1998: 164; No. 420.

482. Neustrotia japonica (Warren, 1912)



483. Neustrotia noloides (Butler, 1879)



484. Neustrotia rectilineata Ueda, 1987



485. Neustrotia albicincta (Hampson, 1898)*


Naranga albicincta Hampson, 1898, Jl. Bombay nat. Hist. Soc. 11: 447 (TL: India, Meghalaya, Khasis [BMNH, London]). Rivula curvilinea Wileman, 1911, Trans. ent. Soc. London 1911: 269, Pl. 31: 18 (TL: Japan, Yamato, Yoshino [BMNH, London]). Ill. Cat. Noctuidae Korea, 1998 _____. Sohn, Ronkay, Kim and Cho, 2005.

DISTRIBUTION. Korea, China, Japan, India, Assam. NOTE. The female specimen was unavailable for dissection. Genus BRYOPHILINA Staudinger, 1892 486. Bryophilina mollicula (Graeser, 1888 [1889]) � genit.: Pl. 76, fig. 1; ♀ genit.: Pl. 227, fig. 7 Ill. Cat. Noctuidae Korea, 1998: 168; No. 436.

Genus MICROXYLA Sugi, 1982 487. Microxyla confusa (Wileman, 1911) � genit.: Pl. 76, fig. 2; ♀ genit.: Pl. 228, fig. 2 Ill. Cat. Noctuidae Korea, 1998: 165; No. 425.

NOTE. The systematic position of the genus is uncertain. The genus Microxyla was transferred to Acontiinae (sensu auctorum) by Sugi (1992: 190). Holloway (2005) considered it to be a member of Catocalinae. Genus EULOCASTRA Butler, 1866 488. Eulocastra excisa (Swinhoe, 1885) Ill. Cat. Noctuidae Korea, 1998: 169; No. 438.

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NOTE. The specimen was unavailable for examination. Genus OZARBA Walker, 1856 489. Ozarba punctigera Walker, 1865 � genit.: Pl. 76, fig. 3; ♀ genit.: Pl. 225, fig. 4 Ill. Cat. Noctuidae Korea, 1998: 170; No. 441.

Genus AMYNA Guenée, 1852 NOTE. The species Amyna glaucopera (Hampson, 1910) [India] is not included in the Catalogue. It was reported from North Korea by Leech (1900: 115) from Gensan [Wonsan] under synonymic name Ilattia apicalis (Moore, 1882). No subsequent records from Korea are known, the data are probably based on misidentification of the other Amyna species. 490. Amyna punctum (Fabricius, 1794)



491. Amyna axis (Guenée, 1852)



492. Amyna stellata Butler, 1878 Ill. Cat. Noctuidae Korea, 1998: 174; No. 449.

NOTE. The residence of the species in Korea requires reconfirmation. Genus HYPERSTROTIA Hampson, 1910 493. “Hyperstrotia” flavipuncta (Leech, 1889) � genit.: Pl. 76, fig. 6; ♀ genit.: Pl. 228, fig. 3 Ill. Cat. Noctuidae Korea, 1998: 169; No. 437.

NOTE. The generic position of this species is dubious.

Subfamily BAGISARINAE Crumb, 1956 Tribe BAGISARINI Crumb, 1956 Genus SPHRAGIFERA Staudinger, 1892 494. Sphragifera sigillata (Ménétriés, 1859) � genit.: Pl. 77, fig. 1; ♀ genit.: Pl. 228, fig. 5 Ill. Cat. Noctuidae Korea, 1998: 264; No. 703.

495. Sphragifera biplagiata (Walker, 1865)



Genus IMOSCA Sugi, 1984 496. Imosca coreana (Matsumura, 1926) � genit.: Pl. 77, fig. 3; ♀ genit.: Pl. 228, fig. 4 Ill. Cat. Noctuidae Korea, 1998: 251; No. 655.

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Genus BREVIPECTEN Hampson, 1894 497. Brevipecten consanguis Leech, 1900 � genit.: Pl. 77, fig. 4; ♀ genit.: Pl. 230, fig. 7 Ill. Cat. Noctuidae Korea, 1998: 110; No. 268.

Genus XANTHODES Guenée, 1852 498. Xanthodes transversa Guenée, 1852 Ill. Cat. Noctuidae Korea, 1998: 175; No. 451 (Pyralidae, Evergestis extimalis, wrong illustration).

NOTE. The species was reported by Leech (1889a: 518) from Wonsan [Gensan], North Korea. No further documentated records is available since Leech. In the Catalogue of Korean Noctuidae, instead of X. transversa, Evergestis extimalis (Scopoli) was incorrectly illustrated (Pyralidae, Evergestinae). The other species, Xanthodes albago (Fabricius, 1794) (Ill. Cat. Noctuidae Korea, 1998: 174; No. 450) should be excluded from the list of Korean Noctuidae. The Pyralidae species, Evergestis sp., (Evergestinae) was misidentified and incorrectly illustrated, instedad of Xanthodes albago.

Subfamily ACONTIINAE Guenée, 1841 Tribe ACONTIINI Guenée, 1841 Genus ACONTIA Ochsenheimer, 1816 Subgenus ACONTIA Ochsenheimer, 1816 499. Acontia olivacea (Hampson, 1891) � genit.: Pl. 77, fig. 6; ♀ genit.: Pl. 228, fig. 7 Ill. Cat. Noctuidae Korea, 1998: 171; No. 443.

Subgenus EMMELIA Hübner, [1821] 1816 500. Acontia trabealis (Scopoli, 1763) � genit.: Pl. 78, fig. 2; ♀ genit.: Pl. 229, fig. 8 Ill. Cat. Noctuidae Korea, 1998: 172; No. 445.


501. Acontia bicolora Leech, 1889 Ill. Cat. Noctuidae Korea, 1998: 171; No. 444.

Genus AUTOBA Walker, [1863] 1864 502. Autoba tristalis (Leech, 1909) � genit.: Pl. 78, fig. 3; ♀ genit.: Pl. 229, fig. 2 Ill. Cat. Noctuidae Korea, 1998: 153; No. 387.

503. Autoba trilinea (de Joannis, 1909)



NOTE. The male specimen was unavailable for dissection. Tribe AEDIINI Beck, 1960 Genus AEDIA Hübner, [1823] 1816 504. Aedia leucomelas (Linnaeus, 1758) � genit.: Pl. 78, fig. 4; ♀ genit.: Pl. 229, fig. 4 Ill. Cat. Noctuidae Korea, 1998: 94; No. 216.

84

Subfamily PANTHEINAE Smith, 1898 Genus PANTHEA Hübner, [1820] 1816 505. Panthea coenobita (Esper, 1785) � genit.: Pl. 79, fig. 1; ♀ genit.: Pl. 229, fig. 6 Ill. Cat. Noctuidae Korea, 1998: 177; No. 456.

Genus PANTHAUMA Staudinger, 1892 506. Panthauma egregia Staudinger, 1892 � genit.: Pl. 79, fig. 2; ♀ genit.: Pl. 229, fig. 5 Ill. Cat. Noctuidae Korea, 1998: 175; No. 452.

Genus COLOCASIA Ochsenheimer, 1816 507. Colocasia mus (Oberthür), 1884 � genit.: Pl. 79, fig. 3; ♀ genit.: Pl. 229, fig. 7 Ill. Cat. Noctuidae Korea, 1998: 178; No. 459.

Genus TRICHOSEA Grote, 1875 508. Trichosea ludifica (Linnaeus, 1758) � genit.: Pl. 79, fig. 4; ♀ genit.: Pl. 229, fig. 8 Ill. Cat. Noctuidae Korea, 1998: 178; No. 457.

509. Trichosea champa (Moore, 1879)



Genus ANACRONICTA Warren, 1909 510. Anacronicta caliginea (Butler, 1881) � genit.: Pl. 80, fig. 1; ♀ genit.: Pl. 230, fig. 2 Ill. Cat. Noctuidae Korea, 1998: 177; No. 455.

Genus XANTHOMANTIS Warren, 1909 511. Xanthomantis cornelia (Staudinger, 1888) � genit.: Pl. 80, fig. 2; ♀ genit.: Pl. 230, fig. 3 Ill. Cat. Noctuidae Korea, 1998: 176; No. 453.

512. Xanthomantis contaminata (Draudt, 1937)



TRISULOIDES Butler, 1881 (Type-species: Trisuloides sericea Butler, 1881 [India: Darjiling, Assam, Shillong]) 513. Trisuloides rotundipennis Sugi, 1976* � genit.: Pl. 80, fig. 5; ♀ genit.: Pl. 230, fig. 5 Trisuloides rotundipennis Sugi, 1976, Tinea, 10(5): 57, figs 4-6 (TL: Japan, Kyushu, Fukuoka Pref., Yoshii [NIAES, Tsukuba]). Ill. Cat. Noctuidae Korea, 1998 _____. Han, Li and Byun, 2007.

DISTRIBUTION. S Korea, Japan.

Subfamily RAPHIINAE Beck, 1996 Genus RAPHIA Hübner, [1821] 1816 514. Raphia peustera Püngeler, 1906 � genit.: Pl. 80, fig. 4; ♀ genit.: Pl. 230, fig. 6 Ill. Cat. Noctuidae Korea, 1998: 179; No. 460.

85

Subfamily BALSINAE Grote, 1896 Genus BALSA Walker, 1860 (pos. prov.) 515. Balsa leodura (Staudinger, 1887) � genit.: Pl. 80, fig. 6; ♀ genit.: Pl. 249, fig. 8 Ill. Cat. Noctuidae Korea, 1998: 260; No. 690.

Subfamily ACRONICTINAE Heinemann, 1859 Genus CYMATOPHOROPSIS Hampson, 1894 516. Cymatophoropsis trimaculata (Bremer, 1861) � genit.: Pl. 81, fig. 1; ♀ genit.: Pl. 231, fig. 1 Ill. Cat. Noctuidae Korea, 1998: 179; No. 461.

517. Cymatophoropsis unca (Houlbert, 1921)



Genus NACNA Nye, 1975 518. Nacna malachitis (Oberthür, 1881) � genit.: Pl. 81, fig. 3; ♀ genit.: Pl. 231, fig. 3 Ill. Cat. Noctuidae Korea, 1998: 183; No. 470.

Genus BELCIADES Kozh., 1950 519. Belciades niveola (Motschulsky, 1866) � genit.: Pl. 81, fig. 4; ♀ genit.: Pl. 231, fig. 4 Ill. Cat. Noctuidae Korea, 1998: 181; No. 464.

Genus BELCIANA Walker, 1862 520. Belciana staudingeri (Leech, 1900) � genit.: Pl. 81, fig. 5; ♀ genit.: Pl. 231, fig. 5 Ill. Cat. Noctuidae Korea, 1998: 180; No. 463.

Genus SUBLEUCONYCTA Kozh., 1950 521. Subleuconycta palshkovi (Filipjev, 1937) � genit.: Pl. 82, fig. 1; ♀ genit.: Pl. 231, fig. 6 Ill. Cat. Noctuidae Korea, 1998: 195; No. 498.

Genus NARCOTICA Sugi, 1982 522. Narcotica niveosparsa (Matsumura, 1926) � genit.: Pl. 82, fig. 2; ♀ genit.: Pl. 231, fig. 7 Ill. Cat. Noctuidae Korea, 1998: 197; No. 506.

Genus GERBATHODES Warren, 1911 523. Gerbathodes paupera (Staudinger, 1892) � genit.: Pl. 82, fig. 3; ♀ genit.: Pl. 231, fig. 8 Ill. Cat. Noctuidae Korea, 1998: 183; No. 471.

524. Gerbathodes angusta (Butler, 1879) Ill. Cat. Noctuidae Korea, 1998: 184; No. 472.

NOTE. The male specimen was not available for dissection.

86


Genus EUROMOIA Staudinger, 1892 525. Euromoia mixta Staudinger, 1892 � genit.: Pl. 82, fig. 4; ♀ genit.: Pl. 232, fig. 2 Ill. Cat. Noctuidae Korea, 1998: 181; No. 465.

526. Euromoia subpulchra (Alphéraky, 1897)



Genus MOMA Hübner, [1820] 1816 527. Moma alpium (Osbeck, 1778) � genit.: Pl. 83, fig. 1; ♀ genit.: Pl. 232, fig. 4 Ill. Cat. Noctuidae Korea, 1998: 182; No. 467.

528. Moma kolthoffi (Bryk, 1948)



529. Moma tsushimana Sugi, 1982



Genus ACRONICTA Ochsenheimer, 1816 Subgenus ACRONICTA Ochsenheimer, 1816 530. Acronicta vulpina (Grote, 1883) � genit.: Pl. 83, fig. 4; ♀ genit.: Pl. 232, fig. 7 Ill. Cat. Noctuidae Korea, 1998: 185; No. 473.

531. Acronicta major (Bremer, 1861)



Subgenus HYBOMA Hübner, [1829] 1816 532. Acronicta adaucta (Warren, 1909) � genit.: Pl. 84, fig. 1; ♀ genit.: Pl. 233, fig. 1 Ill. Cat. Noctuidae Korea, 1998: 186; No. 475.

533. Acronicta strigosa ([Den. & Schiff.], 1775)



534. Acronicta jozana (Matsumura, 1926)



Subgenus MOLYBDONYCTA Sugi, 1979 535. Acronicta bellula (Alphéraky, 1895) � genit.: Pl. 84, fig. 4; ♀ genit.: Pl. 233, fig. 4 Ill. Cat. Noctuidae Korea, 1998: 187; No. 478.

Subgenus PLATAPLECTA 536. Acronicta pulverosa (Hampson, 1909) � genit.: Pl. 84, fig. 5; ♀ genit.: Pl. 233, fig. 5 Ill. Cat. Noctuidae Korea, 1998: 188; No. 479.

537. Acronicta pruinosa (Guenée, 1852)



87

Subgenus TRIAENA Hübner, 1818 538. Acronicta tridens ([Den. & Schiff.], 1775) � genit.: Pl. 85, fig. 1; ♀ genit.: Pl. 233, fig. 7 Ill. Cat. Noctuidae Korea, 1998: 189; No. 481.

539. Acronicta sugii (Kinoshita, 1990)



540. Acronicta intermedia (Warren, 1909)



541. Acronicta cuspis (Hübner, [1813])



542. Acronicta leucocuspis (Butler, 1878)



Subgenus JOCHEAERA 543. Acronicta alni (Linnaeus, 1767) � genit.: Pl. 86, fig. 1; ♀ genit.: Pl. 234, fig. 4 Ill. Cat. Noctuidae Korea, 1998: 191; No. 486.

Subgenus HYLONYCTA Sugi, 1979 544. Acronicta subornata (Leech, 1889) � genit.: Pl. 86, fig. 2; ♀ genit.: Pl. 234, fig. 5 Ill. Cat. Noctuidae Korea, 1998: 191; No. 487.

545. Acronicta carbonaria (Graeser, 1889 [1890])



546. Acronicta catocaloida (Graeser, 1888 [1889])



547. Acronicta hercules (Felder & Rogenhofer, 1874) � genit.: Pl. 86, fig. 5; ♀ genit.: Pl. 234, fig. 8 Ill. Cat. Noctuidae Korea, 1998: 192; No. 490.

Subgenus VIMINIA Chapman., 1890 548. Acronicta rumicis (Linnaeus) � genit.: Pl. 87, fig. 1; ♀ genit.: Pl. 235, fig. 1 Ill. Cat. Noctuidae Korea, 1998: 192; No. 492.

549. Acronicta lutea (Bremer & Grey, 1852)



550. Acronicta digna (Butler, 1881)



551. Acronicta raphael (Oberthür, 1884) Ill. Cat. Noctuidae Korea, 1998: 193; No. 495.

88


Genus SUBACRONYCTA Kozh., 1950 552. Subacronycta concerpta (Draudt, 1937) � genit.: Pl. 87, fig. 5; ♀ genit.: Pl. 235, fig. 5 Ill. Cat. Noctuidae Korea, 1998: 192; No. 491 (Acronicta (Subacronicta) concrepta, misspelling).

Genus SIMYRA Ochsenheimer, 1816 553. Simyra nervosa ([Den. & Schiff.], 1775) � genit.: Pl. 87, fig. 6; ♀ genit.: Pl. 235, fig. 6 Ill. Cat. Noctuidae Korea, 1998: 194; No. 496 (wrong illustration).

NOTE. The other species, S. albovenosa (Goeze, 1781) was erroneously illustraded in the Catalogue of Korean Noctuidae (Kononenko et al., 1998), instead of S. nervosa. 554. Simyra splendida Staudinger, 1888



Genus CRANIONYCTA de Lattin, 1949 555. Cranionycta jankowskii (Oberthür, 1880) � genit.: Pl. 88, fig. 2; ♀ genit.: Pl. 235, fig. 8 Ill. Cat. Noctuidae Korea, 1998: 197; No. 503.

556. Cranionycta albonigra (Herz, 1904)



557. Cranionycta oda deLattin, 1949



Genus CRANIOPHORA Snell., 1867 558. Craniophora ligustri (Den. & Schiff., 1775) � genit.: Pl. 88, fig. 5; ♀ genit.: Pl. 236, fig. 3 Ill. Cat. Noctuidae Korea, 1998: 195; No. 499.

559. Craniophora praeclara (Graeser, 1890)



560. Craniophora pacifica Filipjev, 1927



561. Craniophora fasciata (Moore, 1887)



Subfamily METOPONIINAE Herrich-Schüffer, [1851] Genus USBECA Püngeler, 1914 562. Usbeca kulmburgi (Rebel, 1918) � genit.: Pl. 89, fig. 4; ♀ genit.: Pl. 236, fig. 7 Ill. Cat. Noctuidae Korea, 1998: 261; No. 692.

89

Subfamily SINOCHARINAE Speidel, Fänger, & Naumann, 1996 Genus SINOCHARIS Püngeler, 1912 563. Sinocharis korbae Püngeler, 1912 � genit.: Pl. 89, fig. 5; ♀ genit.: Pl. 236, fig. 8 Ill. Cat. Noctuidae Korea, 1998: 172; No. 446.

Subfamily AGARISTINAE Herrich-Schüffer, [1858] Genus MIMEUSEMIA Butler, 1875 564. Mimeusemia persimilis Butler, 1875 � genit.: Pl. 90, fig. 1; ♀ genit.: Pl. 237, fig. 1 Ill. Cat. Noctuidae Korea, 1998: 203; No. 524.

Genus SARBANISSA Walker, 1865 565. Sarbanissa subflava (Moore, 1877) � genit.: Pl. 90, fig. 2; ♀ genit.: Pl. 237, fig. 3 Ill. Cat. Noctuidae Korea, 1998: 203; No. 525.

566. Sarbanissa venusta (Leech, [1889])



Genus CHELONOMORPHA Motschulsky, [1861] 1860 567. Chelonomorpha japana Motschulsky, [1861] � genit.: Pl. 90, fig. 4♀ genit.: Pl. 237, fig. 2 Ill. Cat. Noctuidae Korea, 1998: 204; No. 527.

NOTE. The female specimen was not avilable for dissection.

Subfamily CUCULLIINAE Herrich-Schäffer, [1850] Genus CUCULLIA Schrank, 1802 568. Cucullia scopariae Dorfmeister, 1853 � genit.: Pl. 91, fig. 1; ♀ genit.: Pl. 237, fig. 5 Ill. Cat. Noctuidae Korea, 1998: 266; No. 705.

569. Cucullia fuschiana Eversmann, 1842



570. Cucullia fraudatrix Eversmann, 1837



571. Cucullia argentea (Hufnagel, 1766)



572. Cucullia jankowskii Oberthür, 1884



573. Cucullia artemisiae (Hufnagel, 1766) 90



574. Cucullia humilis Boursin, 1941



575. Cucullia maculosa Staudinger, 1888



576. Cucullia hostilis Boursin, 1934*


Cucullia hostilis Boursin, 1934, Rev. franc. Entomol. 1934: 143, Pl. 2: 3 (TL: RFE, Primorye: Sutschansky rudnik [LM, Karlsruhe]). Ill. Cat. Noctuidae Korea, 1998. _____. Han, Sohn and Weon, 2006.

DISTRIBUTION. N Korea, N China, RFE (Primorye). 577. Cucullia perforata Bremer, 1861



578. Cucullia mandschuriae Oberthür, 1884



579. Cucullia fraterna Butler, 1878



580. Cucullia umbratica (Linnaeus, 1758)



581. Cucullia kurilullia Bryk, 1942



582. Cucullia elongata Butler, 1880



Subfamily ONCOCNEMIDINAE Forbes & Franclemont, 1954 Genus CALOPHASIA Stephens, 1829 583. Calophasia lunula (Hufnagel, 1766) � genit.: Pl. 94, fig. 1; ♀ genit.: Pl. 239, fig. 4 Ill. Cat. Noctuidae Korea, 1998: 270; No. 719.

Genus ONCOCNEMIS Lederer, 1853 584. Oncocnemis campicola Lederer, 1853 � genit.: Pl. 94, fig. 2; ♀ genit.: Pl. 239, fig. 5 Ill. Cat. Noctuidae Korea, 1998: 272; No. 722.

Genus CALLIERGES Hübner, [1821] 1816 585. Callierges ramosula (Staudinger, 1888) � genit.: Pl. 94, fig. 3; ♀ genit.: Pl. 239, fig. 6 Ill. Cat. Noctuidae Korea, 1998: 271; No. 721.

91

Genus STILBINA Staudinger, 1892 586. Stilbina koreana Draudt, 1934 � genit.: Pl. 94, fig. 4; ♀ genit.: Pl. 239, fig. 7 Ill. Cat. Noctuidae Korea, 1998: 271; No. 720.

Subfamily AMPHIPYRINAE Guenée, 1837 Genus AMPHIPYRA Ochsenheimer, 1816 587. Amphipyra acheron Draudt, 1950 � genit.: Pl. 94, fig. 5; ♀ genit.: Pl. 240, fig. 1 Ill. Cat. Noctuidae Korea, 1998: 243; No. 629.

588. Amphipyra pyramidea (Linnaeus, 1758)



589. Amphipyra monolitha (Guenée, 1852)



590. Amphipyra schrenckii Ménétriés, 1859, 1884



591. Amphipyra jankowskii Oberthür, 1884



592. Amphipyra tripartita Butler, 1878



593. Amphipyra erebina Butler, 1878



594. Amphipyra livida ([Den. & Schiff.], 1775)



Subfamily PSAPHIDINAE Grote, 1896 Tribe PSAPHIDINI Grote, 1896 Genus BRACHIONYCHA Hübner, [1819] 1816 595. Brachionycha nubeculosa (Esper, 1785) � genit.: Pl. 96, fig. 3; ♀ genit.: Pl. 241, fig. 2 Ill. Cat. Noctuidae Korea, 1998: 273; No. 725.

Genus VALERIA Stephens, 1829 (Type-species: Bombyx oleagina [Denis & Schiffermüller], 1775 [Austria]). 596. Valeria tricristata Draudt, 1934* � genit.: Pl. 96, fig. 4 Valeria tricristata Draudt, 1934, in Seitz; 3, suppl.: 141, Pl. 17l (TL: China, Nanking [MNHU, Berlin]). Ill. Cat. Noctuidae Korea, 1998 _____. Han, Kononenko and Park, 2005.

92

DISTRIBUTION. S Korea, China. NOTE. The female specimen was unavailable for dissection. Genus DASEOCHAETA Warren, 1907 597. Daseochaeta viridis (Leech, 1889)



NOTE. The female specimen was unavailable for dissection. Genus MEGANEPHRIA Hübner, [1821] 1816 Subgenus BELOSTICTA Butler, 1879 598. Meganephria cinerea (Butler, 1881) � genit.: Pl. 97, fig. 1; ♀ genit.: Pl. 241, fig. 3 Ill. Cat. Noctuidae Korea, 1998: 273; No. 726.

599. Meganephria funesta (Leech, 1889)



600. Meganephria extensa (Butler, 1879)



601. Meganephria parki Ronkay & Kononenko, 1998



Subgenus MEGANEPHRIA Hübner, [1821] 1816 602. Meganephria kononenkoi Poole, 1989 � genit.: Pl. 97, fig. 5; ♀ genit.: Pl. 242, fig. 1 Ill. Cat. Noctuidae Korea, 1998: 275; No. 730.

603. Meganephria tancrei (Graeser, [1889] 1888)



Tribe FERALIINI Poole, 1995 Genus FERALIA Grote, 1874 604. Feralia sauberi (Graeser, 1892) � genit.: Pl. 98, fig. 1; ♀ genit.: Pl. 241, fig. 1 Ill. Cat. Noctuidae Korea, 1998: 272; No. 723.

Subfamily HELIOTHINAE Boisduval, 1828 Genus HELIOTHIS Ochsenheimer, 1816 605. Heliothis maritima Graslin, 1855 � genit.: Pl. 98, fig. 2; ♀ genit.: Pl. 242, fig. 3 Ill. Cat. Noctuidae Korea, 1998: 372; No. 953.

606. Heliothis ononis ([Den. & Schiff.], 1775)



93

607. Heliothis cruentata (Moore, 1881)*


Curubasa cruentata Moore, 1881, Proc. Zool. Soc. Lond. 1881: 367 (TL: India, Kaschmir, Noashera, BMNH, London) Curubasa marginata Moore, 1881, Proc. Zool. Soc. Lond. 1881: 367 (TL: India, NW Himalaya [BMNH, London]). Ill. Cat. Noctuidae Korea, 1998 _____.

DISTRIBUTION. S Korea, China, Japan, Tibet, Nepal, N India. NOTE. The species is reported for Korea for the first time. Only a female from Korea is known. It was refered as Masalia cruentana and Heliothis cruentana by Sugi (1982, 2000). The correct spelling of the name is Heliothis cruentata. Material examined. 1 female, Korea, Wando, 8 IX 2002 (Gab-Jae Weon). Genus HELIOCHEILUS Grote, 1865 608. Heliocheilus fervens (Butler, 1881) � genit.: Pl. 98, fig. 4; ♀ genit.: Pl. 242, fig. 5 Ill. Cat. Noctuidae Korea, 1998: 375; No. 956.

Genus PROTOSCHINIA Hardw., 1970 609. Schinia scutosa (Goeze, 1781) � genit.: Pl. 98, fig. 5; ♀ genit.: Pl. 242, fig. 6 Ill. Cat. Noctuidae Korea, 1998: 374; No. 955.

Genus HELICOVERPA Hardw., 1965 610. Helicoverpa armigera (Hübner, [1808]) � genit.: Pl. 98, fig. 6; ♀ genit.: Pl. 243, fig. 1 Ill. Cat. Noctuidae Korea, 1998: 371; No. 951.

611. Helicoverpa assulta (Guenée, 1852)



Genus PYRRHIA Hübner, [1821] 1816 612. Pyrrhia umbra (Hufnagel, 1766) � genit.: Pl. 99, fig. 2; ♀ genit.: Pl. 243, fig. 3 Ill. Cat. Noctuidae Korea, 1998: 375; No. 957.

613. Pyrrhia exprimens (Walker, 1857)



614. Pyrrhia hedemanni (Staudinger, 1892)



615. Pyrrhia bifasciata (Staudinger, 1888)



Genus ADISURA Moore, 1881 616. Adisura atkinsoni Moore, 1881 � genit.: Pl. 99, fig. 6; ♀ genit.: Pl. 243, fig. 7 Ill. Cat. Noctuidae Korea, 1998: 377; No. 961.

94

Subfamily CONDICINAE Poole, 1995 Tribe CONDICINI Poole, 1995 Genus CONDICA Walker, 1856 617. Condica illecta (Walker, 1865) � genit.: Pl. 100, fig. 1; ♀ genit.: Pl. 244, fig. 1 Ill. Cat. Noctuidae Korea, 1998: 257; No. 681.

618. Condica illustrata (Staudinger, 1888)



619. Condica fuliginosa (Leech, 1900) Ahn Kononenko and Park, 1994a: 37, misidentification of Condica cyclicoides. Ill. Cat. Noctuidae Korea, 1998: 258; No. 681.

NOTE. First reported for Korea by Park and Yoo (1986: 41) and later by Ahn et al. (1994a: 37). The last record is a misidentification of Condica cyclicoides. The distribution of the species in Korea is possible, but require further confirmation. The specimens were not available for dissection. 620. Condica cyclicoides (Draudt, 1950)*


Perigea cyclicoides Draudt, 1950, Mitt. Münch. entomol. Ges. 40: 94, pl. 6, fig. 16 (TL: China, Hoeng-shan, West Tienmushan [ZFMK, Bonn]). Ill. Cat. Noctuidae Korea, 1998 _____. See No. 683 (Prospalta cyclica).

DISTRIBUTION. S Korea, China. NOTE. The species was misidentified as Condica fuliginosa by Ahn et al. (1994a) and by Kononenko et al. (1998), as Prospalta cyclica. The present identification is based on examination of the type series of Perigea cyclicoides. First report for Korea. Additional material examined: 1 male, Korea, Gajengri, Chuncheon, KW, 17. VI 2002 (Han Huy Lin). 621. Condica sp.*


Ill. Cat. Noctuidae Korea, 1998 _____.

DISTRIBUTION. S Korea, RFE (Prinorye terr.). NOTE. Probably an undescribed species of the genus Condica. For the description of the species the revision of its allied is necessary. Genus PROSPALTA Walker, [1858] 1857 622. Prospalta cyclica (Hampson, 1908)


Ill. Cat. Noctuidae Korea, 1998: 258; No. 683 (misidentification of Condica cyclicoides).

NOTE. First reported for Korea by Pak (1959: 15). The species was misidentified in the Catalogue of Korean Noctuidae: C. cyclicoides was illustrated instead of P. cyclica. The distribution of the species in Korea is possible, but require a further confirmation. The female specimen was not available for dissection.

95

Genus CHYTONIX Grote, 1874 623. Chytonix albonotata (Staudinger, 1892) � genit.: Pl. 100, fig. 6; ♀ genit.: Pl. 244, fig. 5 Ill. Cat. Noctuidae Korea, 1998: 254; No. 669.

Genus NIPHONYX Sugi, 1982 624. Niphonyx segregata (Butler, 1878) � genit.: Pl. 101, fig. 1; ♀ genit.: Pl. 244, fig. 6 Ill. Cat. Noctuidae Korea, 1998: 254; No. 670.

Genus OLIGONYX Sugi, 1982 625. Oligonyx vulnerata (Butler, 1878) � genit.: Pl. 101, fig. 2; ♀ genit.: Pl. 244, fig. 7 Ill. Cat. Noctuidae Korea, 1998: 255; No. 671.

Genus NIKARA Moore, 1882 626. Nikara castanea Moore, 1881*


Nikara castanea Moore, 1882, in Hewitson and Moore, Descr. new Indian lepid. Insects Colln late Mr W. S. Afkinson: 126, pi. 4, fig. 24 (TL: India, Sikkim, Darjiling, MNHU, Berlin) Ill. Cat. Noctuidae Korea, 1998 _____.

DISTRIBUTION. Korea (first record), China, Vietnam, India, Nepal. NOTE. First report for Korea. Material examined: 1 male, North Korea, Myohyang-san, August 1991, genit. prep. 6886 LR. HNHM. Genus PYRRHIDIVALVA Sugi, 1982 627. Pyrrhidivalva sordida (Butler, 1881) � genit.: Pl. 101, fig. 3; ♀ genit.: Pl. 244, fig. 8 Ill. Cat. Noctuidae Korea, 1998: 255; No. 672.

Genus DYSMILICHIA Speis., 1902 628. Dysmilichia gemella (Leech, 1889) � genit.: Pl. 101, fig. 4; ♀ genit.: Pl. 245, fig. 1 Ill. Cat. Noctuidae Korea, 1998: 257; No. 678.

629. Dysmilichia fukudai Sugi, 1963



Genus EUCARTA Lederer, 1857 630. Eucarta fasciata (Butler, 1878) � genit.: Pl. 101, fig. 6; ♀ genit.: Pl. 245, fig. 3 Ill. Cat. Noctuidae Korea, 1998: 256; No. 673.

631. Eucarta arcta (Lederer, 1853)



632. Eucarta arctides (Staudinger, 1888)



633. Eucarta virgo (Treitschke, 1835) Ill. Cat. Noctuidae Korea, 1998: 256; No. 676.

96


634. Eucarta amethystina (Hübner, [1803])



Genus ACOSMETIA Stephens, 1829 635. Acosmetia chinensis (Wallengren, 1860) � genit.: Pl. 102, fig. 5; ♀ genit.: Pl. 245, fig. 8 Ill. Cat. Noctuidae Korea, 1998: 259; No. 685 (Hajina sinensis).

NOTE: Transferred to the genus Acosmetia by Kononenko (2005). 636. Acosmetia biguttula (Motschulsky, 1866)


Ill. Cat. Noctuidae Korea, 1998: 259; No. 684 (Hajina biguttula).

NOTE. Reported for Korea from Gensan [Wonsan] by Leech (1889a: 491, 1900: 116) and also by Yoon et al. (1982: 30). The distribution of the species in Korea require a further confirmation. Transferred to the genus Acosmetia by Kononenko (2005). Genus IAMBIA Walker, 1863 637. Iambia transversa (Moore, 1882) � genit.: Pl. 103, fig. 1; ♀ genit.: Pl. 246, fig. 1 Ill. Cat. Noctuidae Korea, 1998: 259; No. 686.

638. Iambia japonica Sugi, 1958



Genus PLUSILLA Staudinger, 1892 639. Plusilla rosalia Staudinger, 1892 � genit.: Pl. 103, fig. 3; ♀ genit.: Pl. 246, fig. 3 Ill. Cat. Noctuidae Korea, 1998: 261; No. 691.

Genus PROMETOPUS Guenée, 1852 (pos. prov.) 640. Prometopus flavicollis (Leech, 1889) � genit.: Pl. 103, fig. 5; ♀ genit.: Pl. 246, fig. 5 Ill. Cat. Noctuidae Korea, 1998: 260; No. 688.

Subfamily ERIOPINAE Herrich-Schäffer, [1851] Genus CALLOPISTRIA Hübner, [1821] 1816 641. Callopistria juventina (Stoll, 1782) � genit.: Pl. 103, fig. 6; ♀ genit.: Pl. 246, fig. 6 Ill. Cat. Noctuidae Korea, 1998: 262; No. 693.

642. Callopistria repleta Walker, 1858



643. Callopistria duplicans Walker, [1858] Ill. Cat. Noctuidae Korea, 1998: 262; No. 695.

NOTE. The specimens were not available for dissection. 644. Callopistria aethiops Butler, 1878

� genit.: Pl. 104, fig. 5 97


NOTE. The female specimen was not available for dissection. 645. Callopistria albolineola (Graeser, 1888 [1889])




646. Callopistria japonibia Inoue & Sugi, 1958 Ill. Cat. Noctuidae Korea, 1998: 263; No. 698.

NOTE. The male specimen was not available for dissection. ♀ genit.: Pl. 247, fig. 3

647. Callopistria rivularis Walker, [1858] 1857 Ill. Cat. Noctuidae Korea, 1998: 263; No. 699.

NOTE. The male specimen was not available for dissection. 648. Callopistria placodoides (Guenée, 1852)



649. Callopistria argyrosticta (Butler, 1881)



650. “Callopistria” miracula Herz, 1904



NOTE. The generic position of the species is uncertain. It is not congeneric with Callopistria.

Subfamily BRYOPHILINAE Guenée, 1852 Genus BRYOMOIA Staudinger, 1892 651 . Bryomoia melachlora (Staudinger, 1892) � genit.: Pl. 107, fig. 5; ♀ genit.: Pl. 247, fig. 6 Ill. Cat. Noctuidae Korea, 1998: 197; No. 507.

Genus CRYPHIA Hübner, 1818 652. Cryphia bryophasma (Boursin, 1951) � genit.: Pl. 104, fig. 6; ♀ genit.: Pl. 247, fig. 7 Ill. Cat. Noctuidae Korea, 1998: 198; No. 509.

653. Cryphia mitsuhashi (Marumo, 1917)



654. Cryphia mediofusca (Sugi, 1959)



655. Cryphia griseola (Nagano, 1918) Ill. Cat. Noctuidae Korea, 1998: 199; No. 511.

98


656. Cryphia minutissima (Draudt, 1950)



657. Cryphia sugitanii Boursin, 1961



Genus BRYOLEUCA Hampson, 1908 658. Bryoleuca granitalis (Butler, 1881) � genit.: Pl. 105, fig. 6; ♀ genit.: Pl. 248, fig. 5 Ill. Cat. Noctuidae Korea, 1998: 200; No. 514 (Bryophila granitalis).

659. Bryoleuca albimixta Sugi, 1980


Ill. Cat. Noctuidae Korea, 1998: 200; No. 515 (Bryophila albimixta).

660. Bryoleuca orthogramma (Boursin, 1954)


Ill. Cat. Noctuidae Korea, 1998: 200; No. 516 (Bryophila orthogramma).

Genus STENOLOBA Staudinger, 1892 661. Stenoloba oculata Draudt, 1950 � genit.: Pl. 106, fig. 3; ♀ genit.: Pl. 248, fig. 8 Ill. Cat. Noctuidae Korea, 1998: 201; No. 518.

662. Stenoloba clara (Leech, 1889)



663. Stenoloba assimilis (Warren, 1909)



664. Stenoloba manleyi (Leech, 1889)



665. Stenoloba fontinalis Ronkay & Kononenko, 1998 � genit.: Pl. 106, fig. 1; ♀ genit.: Pl. 249, fig. 4 Ill. Cat. Noctuidae Korea, 1998: 202; No. 522.

666. Stenoloba glauca Kononenko & Ronkay, 2001* � genit.: Pl. 107, fig. 1; ♀ genit.: Pl. 249, fig. 6 Stenoloba glauca Kononenko & Ronkay, 2001, Ins. Koreana, 18(12): 103, figs 11, 29 (TL: N Vietnam [HNHM, Budapest]). ssp. kuytekparki Sohn & Han, 2005, stat. n. Stenoloba kuytekparki Sohn & Han, 2005, Tinea 19(1): 23, figs 1, 2, 4, 6, 7. Ill. Cat. Noctuidae Korea, 1998 _____. Sohn and Han, 2005 (Stenoloba kuytekparki).

DISTRIBUTION. S Korea, N Vietnam. NOTE. The senior author consider the taxon kuytekparki as a subspecies of the Stenoloba glauca Kononenko and Ronkay, 2001. 667. Stenoloba jankowskii (Oberthür, 1884)



99

Genus ATHAUMASTA Hampson, 1906 668. Athaumasta koreana Ronkay & Kononenko, 1998 � genit.: Pl. 107, fig. 3; ♀ genit.: Pl. 249, fig. 7 Ill. Cat. Noctuidae Korea, 1998: 201; No. 517.

Subfamily XYLENINAE Guenée, 1837 Tribe PSEUDEUSTROTIINI Beck, 1996 Genus PSEUDEUSTROTIA Warren, 1913 669. Pseudeustrotia candidula ([Den. & Schiff.], 1775) � genit.: Pl. 107, fig. 6; ♀ genit.: Pl. 250, fig. 1 Ill. Cat. Noctuidae Korea, 1998: 165; No. 424.

Genus ANTERASTRIA Sugi, 1982 670. Anterastria atrata (Butler, 1881) � genit.: Pl. 108, fig1; ♀ genit.: Pl. 250, fig. 2 Ill. Cat. Noctuidae Korea, 1998: 167; No. 434.

Genus PHYLLOPHILA Guenée, 1852 671. Phyllophila obliterata (Rambur, 183) � genit.: Pl. 108, fig. 2; ♀ genit.: Pl. 250, fig. 3 Ill. Cat. Noctuidae Korea, 1998: 170; No. 440.

Tribe PRODENIINI Forbes, 1954 Genus SPODOPTERA Guenée, 1852 672. Spodoptera exigua (Hübner, [1808]) � genit.: Pl. 108, fig. 4; ♀ genit.: Pl. 250, fig. 4 Ill. Cat. Noctuidae Korea, 1998: 233; No. 606.

673. Spodoptera litura (Fabricius, 1775)



674. Spodoptera depravata (Butler, 1879)



675. Spodoptera pecten Guenée, 1852



NOTE. Warren (1914) reported Spodoptera pecten from Wonsan [Gensan]. The species was refered in Korean literature and included in the Check list of insects from Korea (1994: 373) on the basis of the previous records. The distribution of S. pecten in Korea requires a further confirmation. 676. Spodoptera apertura (Walker, 1856)



NOTE. First reported for Korea by Ahn et al. (1994a: 36) on the basis of single male, collected in Gwangneung. Probably the species is tropical migrant in Korea. The female specimen was not available for dissection.

100

677. Spodoptera mauritia (Boisduval, 1833)*


Hadena mauritia Boisduval, 1833, Nouv. Annls. Mus. Paris, 2: 240, Pl. 13: 9 and Fauna ent. Madag., Bourbon & Maurice, (Lép.): 92, Pl. 1: 4 (TL: Mauritius, Réunion). Hadena filium Guenée, 1852, Hist. nat. Insectes (Lépid.) 5: 155, (TL: Indonesia, Java). Hadena nubes Guenée, 1852, Hist. nat. Insectes (Lépid.) 5: 155, (TL: Indonesia, Java; Central India). Prodenia infecta Walker, 1856, List. Spec. lep. Ins. B. M. 9: 196 (HT: N India, Himalaya [BMNH, London]). Prodenia insignata Walker, 1856, List Specimens lepid. Insects Colln. Br. Mus. 9: 197 (HT: N India, Himalaya [BMNH, London]). Prodenia permuda Walker, 1857, List Specimens lepid. Insects Colln. Br. Mus. 10: 327 (HT: Thailand [BMNH, London]). Agrotis transducta Walker, 1857, List Specimens lepid. Insects Colln. Br. Mus. 10: 344 (HT: Sri Lanka, BMNH, London). Agrotis aliena Walker, 1865, List Specimens lepid. Insects Colln. Br. Mus. (HT: Mauritius [BMNH, London]). Spodoptera mauritia acronyctoides Guenée, 1852, Hist. nat. Insectes (Lépid.) 5: 155 (HT: Tahiti). Spodoptera mauritia acronyctiformis Guenée, 1852, Hist. nat. Insectes (Lépid.) 7: 399 (HT: Tahiti). Euxoa ogasawarensis Matsumura, 1926, Insect. matsum. 1: 6, Pl. 1: 19 (TL: Japan, Bonin I., Ogasawara I. [EIHU, Sapporo]). Ill. Cat. Noctuidae Korea, 1998 _____. Sohn and Ronkay, 2001.

DISTRIBUTION. Korea, Japan, China, Taiwan, India, SE Asia, Australia, Africa, Madagascar, Saudi Arabia, Yemen. NOTE. The species is probably tropical migrant in Korea. Tribe ELAPHRIINAE Beck, 1996 Genus ELAPHRIA Hübner, 1818 678. Elaphria venustula (Hübner, 1790) � genit.: Pl. 117, fig. 6; ♀ genit.: Pl. 251, fig. 2 Ill. Cat. Noctuidae Korea, 1998: 168; No. 435.

Tribe CARADRININI Boisduval, 1840 Subtribe CARADRININA Boisduval, 1840 Genus CARADRINA Ochsenheimer, 1816 Subgenus PARADRINA Boursin, 1937 679. Caradrina chinensis Leech, 1900 � genit.: Pl. 109, fig. 3; ♀ genit.: Pl. 251, fig. 3 Caradrina chinensis Leech, 1900. Trans. Zool. Soc. London: 117. [TL: Western China Pu-tsu-fon, Ta-chien-lu, Ni-ton] [BMNH, London]. Caradrina fusca Leech, 1900: Trans. Zool. Soc. London, 1900: 118. (TL: Western China, Ta-chien-lu, Ni-ton] [BMNH]). Athetis heptarchia Boursin, 1936 Bull. Soc. entomol. France, 1936: 93 (HT: China, Yunnan, Tali, [MNHN, Paris]). Ill. Cat. Noctuidae Korea, 1998: 242; No. 627 (Caradrina sp.).

DISTRIBUTION. Korea, West and Central China. NOTE. A male specimen of this species was reported from North Korea, Shuotsu and illustrated by Bryk (1948) as a paratype of Caradrina morosa permorosa Bryk, 1948. We dissected, examined and identified it hereby as Caradrina chinensis Leech, 1900. Subgenus EREMODRINA Boursin, 1937 680. Caradrina morosa (Lederer, 1853) � genit.: Pl. 109, fig. 5; ♀ genit.: Pl. 251, fig. 5 Ill. Cat. Noctuidae Korea, 1998: 242; No. 628.

101

Subgenus PLATYPERIGEA Smith, 1894 681. Caradrina montana (Bremer, 1861) � genit.: Pl. 109, fig. 4; ♀ genit.: Pl. 251, fig. 4 Ill. Cat. Noctuidae Korea, 1998: 240; No. 626.

Genus HOPLODRINA Boursin, 1937 682. Hoplodrina octogenaria (Goeze, 1781) � genit.: Pl. 109, fig. 6; ♀ genit.: Pl. 251, fig. 6 Ill. Cat. Noctuidae Korea, 1998: 239; No. 624.

683. Hoplodrina euryptera Boursin, 1937



Genus STYGIODRINA Boursin, 1937 684. Stygiodrina maurella (Staudinger, 1888) � genit.: Pl. 110, fig. 2; ♀ genit.: Pl. 251, fig. 8 Ill. Cat. Noctuidae Korea, 1998: 235; No. 611.

Subtribe ATHETINA Fibiger & Lafontaine, 2005 Genus ATHETIS Hübner, [1821] 1816 685. Athetis cinerascens (Motschulsky, [1861] 1860) � genit.: Pl. 110, fig. 3; ♀ genit.: Pl. 252, fig. 1 Ill. Cat. Noctuidae Korea, 1998: 236; No. 612.

686. Athetis furvula (Hübner, [1808])



687. Athetis gluteosa (Treitschke, 1835)



688. Athetis funesta (Staudinger, 1888)



689. Athetis lapidea Wileman, 1911



690. Athetis lepigone (Möschler, 1860)



691. Athetis correpta (Püngeler, 1907)



692. Athetis dissimilis (Hampson, 1909)



693. Athetis albisignata (Oberthür, 1879)



694. Athetis pallidipennis Sugi, 1982

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695. Athetis stellata (Moore, 1882)



696. Athetis lineosa (Moore, 1881)



Tribe DYPTERYGIINI Forbes, 1954 Genus DIPTERIGIA Stephens, 1829 697. Dipterigia caliginosa (Walker, 1858) � genit.: Pl. 118, fig. 2; ♀ genit.: Pl. 257, fig. 8 Ill. Cat. Noctuidae Korea, 1998: 229; No. 596.

Genus DIPTERYGINA Sugi, 1954 698. Dipterygina cupreotincta Sugi, 1954 � genit.: Pl. 120, fig. 1; ♀ genit.: Pl. 258, fig. 1 Ill. Cat. Noctuidae Korea, 1998: 232; No. 603.

Genus TRACHEA Ochsnheimer, 1816 (HERAEMA Staudinger, 1892, syn. n.) NOTE. The examination of the external morphology and male and female genitalia of Heraema mandschurica, Trachea atriplicis and the other representatives of the genus Trachea revealed their congenerity. Hereby the generic name Heraema (syn. n.) is synonimised with Trachea. 699. Trachea atriplicis (Linnaeus, 1758)



700. Trachea punkikonis Matsumura, 1927



701. Trachea melanospila Kollar, [1844]



702. Trachea tokiensis (Butler, 1884)



703. Trachea mandschurica (Bremer, 1861), comb. n.


Ill. Cat. Noctuidae Korea, 1998: 228; No. 593 (Heraema mandschurica).

Genus OLIVENEBULA Kishida & Yoshimoto, 1977 (CHLOROTHALPA Beck, 1996) 704. Olivenebula oberthueri (Staudinger, 1892) � genit.: Pl. 120, fig. 3; ♀ genit.: Pl. 259, fig. 1 Ill. Cat. Noctuidae Korea, 1998: 224; No. 582.

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Genus POLYPHAENIS Boisduval, 1840 705. Polyphaenis subviridis (Butler, 1878) � genit.: Pl. 120, fig. 4; ♀ genit.: Pl. 259, fig. 2 Ill. Cat. Noctuidae Korea, 1998: 225; No. 583.

Genus XENOTRACHEA Sugi, 1958 706. Xenotrachea niphonica Kish. & Yoshim., 1979 � genit.: Pl. 121, fig. 4; ♀ genit.: Pl. 259, fig. 7 Ill. Cat. Noctuidae Korea, 1998: 227; No. 590.

Genus KARANA Moore, 1882 707. Karana laetevirens (Oberthür, 1884) � genit.: Pl. 121, fig. 5; ♀ genit.: Pl. 259, fig. 8 Ill. Cat. Noctuidae Korea, 1998: 231; No. 602.

Genus TRIPHAENOPSIS Butler, 1878 708. Triphaenopsis lucilla Butler, 1878 � genit.: Pl. 121, fig. 6; ♀ genit.: Pl. 260, fig. 2 Ill. Cat. Noctuidae Korea, 1998: 223; No. 578.

709. Triphaenopsis jezoensis Sugi, 1962



710. Triphaenopsis cinerescens Butler, 1885



711. Triphaenopsis postflava (Leech, 1900)



Genus MORMO Ochsenheimer, 1816 (pos. prov.) 712. Mormo muscivirens Butler, 1878 � genit.: Pl. 118, fig. 5; ♀ genit.: Pl. 258, fig. 4 Ill. Cat. Noctuidae Korea, 1998: 245; No. 637.

Genus ORTHOGONIA Felder & Felder, 1862 713. Orthogonia sera Felder & Felder, 1862 � genit.: Pl. 119, fig. 1; ♀ genit.: Pl. 258, fig. 5 Ill. Cat. Noctuidae Korea, 1998: 246; No. 638.

714. Orthogonia tapaishana (Draudt, 1939)



NOTE. The distribution of this rare speices in Korea is reconfirmed. Material examined: 1male, 1female, “North Korea”, Kumgan-san, 16-19. VII 1985, leg. E. Palik (ZFMK, Bonn). 715. Orthogonia basimacula (Draudt, 1939)*


Orthogonica basimacula Draudt, 1939, Ent. Rdsch. 56(15): 146, Pl. 1: 3 (TL: China, westl. Tien-mu-shan [ZFMK, Bonn]). Ill. Cat. Noctuidae Korea, 1998 _____. Sohn, Ronkay and Kononenko, 2004.

DISTRIBUTION. Korea, China.

104

Tribe ACTINOTIINI Beck, 1996 Genus HYPPA Duponchel, 1844 716. Hyppa rectilinea (Esper, 1788) � genit.: Pl. 118, fig. 4; ♀ genit.: Pl. 258, fig. 3 Ill. Cat. Noctuidae Korea, 1998: 228; No. 591.

Genus ACTINOTIA Hübner, [1823] 1816 717. Actinotia intermediata (Bremer, 1861) � genit.: Pl. 120, fig. 1; ♀ genit.: Pl. 258, fig. 7 Ill. Cat. Noctuidae Korea, 1998: 229; No. 594.

718. Actinotia polyodon (Clerck, 1759)



Tribe PHLOGOPHORINI Hampson, 1918 Genus PHLOGOPHORA Treitschke, 1825 719. Phlogophora beatrix Butler, 1878 � genit.: Pl. 121, fig. 2; ♀ genit.: Pl. 259, fig. 5 Ill. Cat. Noctuidae Korea, 1998: 227; No. 587.

720. Phlogophora illustrata (Graeser, 1888 [1889])



Genus EUPLEXIA Stephens, 1829 721. Euplexia lucipara (Linnaeus, 1758) � genit.: Pl. 120, fig. 5; ♀ genit.: Pl. 259, fig. 3 Ill. Cat. Noctuidae Korea, 1998: 225; No. 584.

722. Euplexia koreaeplexia Bryk, 1948



Genus EUPLEXIDIA Hampson, 1896 723. Euplexidia angusta Yoshimoto, 1987 Ill. Cat. Noctuidae Korea, 1998: 227; No. 589.

NOTE. The specimen was not available for dissection. Genus SASUNAGA Moore, 1881 724. Sasunaga longiplaga Warren, 1912 � genit.: Pl. 118, fig. 6; ♀ genit.: Pl. 260, fig. 1 Ill. Cat. Noctuidae Korea, 1998: 232; No. 604.

Genus AUCHMIS Hübner, [1821] 1816 725. Auchmis saga (Butler, 1878) � genit.: Pl. 118, fig. 3; ♀ genit.: Pl. 258, fig. 2 Ill. Cat. Noctuidae Korea, 1998: 227; No. 592.

105

Tribe APAMEINI Guenée, 1841 (NONAGRIINI Guenée, 1837, suppressed senior syn.) Subtribe OXYTRIPINA Goznamy, 1970 Genus OXYTRIPIA Staudinger, 1871 726. Oxytripia orbiculosa (Esper, 1799) � genit.: Pl. 122, fig. 4; ♀ genit.: Pl. 260, fig. 6 Ill. Cat. Noctuidae Korea, 1998: 230; No. 597.

Subtribe APAMEINA Guenée, 1841 Genus SIDEMIA Staudinger, 1892 727. Sidemia bremeri (Erschoff, 1870) � genit.: Pl. 122, fig. 5; ♀ genit.: Pl. 260, fig. 7 Ill. Cat. Noctuidae Korea, 1998: 204; No. 528.

728. Sidemia spilogramma (Rambur, 1871)



Genus BAMBUSIPHILA Sugi, 1958 729. Bambusiphila vulgaris (Butler, 1886)



NOTE. First reported for Korea by Pak (1959: 11), his record was repeated in the Korean literature (Nam, 1982: 362, 750; Check List of Insects from Korea, 1994: 371). The male specimen was not available for examination and dissection. Genus STAUROPHORA R. L., 1817 730. Staurophora celsia (Linnaeus, 1758) � genit.: Pl. 123, fig. 2; ♀ genit.: Pl. 261, fig. 1 Ill. Cat. Noctuidae Korea, 1998: 215; No. 555.

Genus HELOTROPHA Lederer, 1857 731. Helotropha leucostigma (Hübner, [1808]) � genit.: Pl. 123, fig. 3; ♀ genit.: Pl. 261, fig. 2 Ill. Cat. Noctuidae Korea, 1998: 218; No. 563.

Genus GORTYNA Ochsenheimer, 1816 732. Gortyna fortis (Butler, 1878) � genit.: Pl. 123, fig. 4; ♀ genit.: Pl. 261, fig. 3 Ill. Cat. Noctuidae Korea, 1998: 219; No. 564.

733. Gortyna basalipunctata Graeser, 1888 [1889]



Genus HYDRAECIA Guenée, 1841 734. Hydraecia petasitis Doubleday, 1847 � genit.: Pl. 124, fig. 1; ♀ genit.: Pl. 261, fig. 5 Ill. Cat. Noctuidae Korea, 1998: 216; No. 557.

Genus AMPHIPOEA Billberg, 1820 735. Amphipoea fucosa (Freyer, 1830) � genit.: Pl. 124, fig. 2; ♀ genit.: Pl. 261, fig. 6 106


736. Amphipoea lucens (Freyer, 1845)



737. Amphipoea burrowsi (Chapman, 1912)



738. Amphipoea ussuriensis (Petersen, 1914)



739. Amphipoea asiatica (Burrows, 1912)



Genus RHIZEDRA Warren, 1911 740. Rhizedra lutosa (Hübner, [1803]) � genit.: Pl. 125, fig. 3; ♀ genit.: Pl. 262, fig. 6 Ill. Cat. Noctuidae Korea, 1998: 220; No. 570.

Genus ARCHANARA Walker, 1866 741. Archanara neurica (Hübner, [1808]) � genit.: Pl. 125, fig. 2; ♀ genit.: Pl. 262, fig. 3 Ill. Cat. Noctuidae Korea, 1998: 219; No. 565.

Genus CTENOSTOLA Sugi, 1982 742. Ctenostola sparganoides (Bang-Haas, 1927) � genit.: Pl. 125, fig. 4; ♀ genit.: Pl. 262, fig. 5 Ill. Cat. Noctuidae Korea, 1998: 221; No. 571.

Genus COENOBIA Stephens, 1850 743. Coenobia orientalis Sugi, 1982



Genus CAPSULA Fibiger, Zilli, Ronkay & Golstein, 2005 (Type species: Phalaena Noctua algae Esper, 1789 [Europe]). 744. Capsula aerata (Butler, 1878) � genit.: Pl. 125, fig. 5; ♀ genit.: Pl. 262, fig. 7 Ill. Cat. Noctuidae Korea, 1998: 219; No. 567.

745. Capsula sparganii (Esper, 1790)



Genus PABULATRIX Sugi, 1982 746. Pabulatrix pabulatricula (Brahm, 1791) � genit.: Pl. 126, fig. 1; ♀ genit.: Pl. 263, fig. 1 Ill. Cat. Noctuidae Korea, 1998: 210; No. 542.

Genus ASIDEMIA Sugi, 1982 (Type species: Apamea inexpecta Sugi, 1963 [Japan]). 747. Asidemia inexpecta (Sugi, 1963) � genit.: Pl. 126, fig. 2; ♀ genit.: Pl. 265, fig. 4 Apamea inexpecta Sugi, 1963, Tinea 6: 12, pl. 4: 9 (TL: Japan, Niigata Pref., Mt. Yahiko, coll. S. Sugi, Tokyo). Asidemia

107

inexpecta ssp. inexpecta (Sugi, 1963). - Japan (Honshu), Korea. Apamea inexpecta ssp. insulicola (Sugi, 1963) Tinea, 6: 12, pl. 4: 10, 9 (TL: Japan, Mikura I., coll. S. Sugi, Tokyo). Sidemia inexpecta ssp. insulicola (Sugi, 1963). - Japan: Izu I., Miyaka I., Mikura I. Ill. Cat. Noctuidae Korea, 1998: 223; No. 576 (Kumasia kumaso, misidentification).

DISTRIBUTION. Korea (Central, Cheju I.), Japan (Honshu, Miyaka I., Mikura I.). NOTE. The species was incorrectly reported for Korea (Ahn et al., 1994: 35, Kononenko et al., 1998: Nr. 576, fig. 576) as Kumasia kumaso (Sugi, 1963). Genus FELINIOPSIS Roepke, 1938 (Type-species: Feliniopsis incerta Roepke, 1938 [Indonesia, Sulawesi]) 748. Feliniopsis indistans (Guenée, 1852)* � genit.: Pl. 126, fig. 3 Hadena indistans Guenée, 1852, in Boisduval and Guenée, Hist. nat. Insectes (Lépid.) 6: 87 (TL: Central India [BMNH, London]). Hadena subdistans Guenée, 1852, in Boisduval and Guenée, Hist. nat. Insectes (Lépid.) 7: 399. Toxocampa alikangiae Strand, 1920, Arch. Naturgesh. 84A(12): 136 (TL: Taiwan [EI, Eberswalde]). Eutamsia asahinai Sugi, 1982 Moths of Japan, 1: 762, 2: 369, Pl. 187: 21, 22 (TL: Japan [NIAES, Tsukuba]). Ill. Cat. Noctuidae Korea, 1998 _____. Sohn, Ronkay, Kim and Cho, 2005.

DISTRIBUTION. Korea, Japan, China, Taiwan, Vietnam, Nepal, India, Sri Lanka. Genus ATRACHEA Warren, 1911 749. Atrachea nitens (Butler, 1878) � genit.: Pl. 126, fig. 4; ♀ genit.: Pl. 263, fig. 2 Ill. Cat. Noctuidae Korea, 1998: 209; No. 539.

750. Atrachea jankowskii (Oberthür, 1879)



751. Atrachea alpherakyi Kononenko, 1986



Genus APAMEA Ochsenheimer, 1816 752. Apamea aquila Donzel, 1837 � genit.: Pl. 127, fig. 2; ♀ genit.: Pl. 263, fig. 5 Ill. Cat. Noctuidae Korea, 1998: 205; No. 531.

753. Apamea striata Haruta & Sugi, 1958



754. Apamea sodalis (Butler, 1878)*


Xylophasia sodalis Butler, 1878, Ann. Mag. nat. Hist. (5)1: 83 (TL: Japan, Yokohama [BMNH, London]). Hadena obliquiorbis Moore, 1882, in Hewitson and Moore, Descr. new Indian lepid. Insects Colln late Mr W. S. Afkinson: 109, (TL: India, Sikkim, Darjiling [BMNH, London]). China, Japan, India, Assam, Himalaya, Sikkim, Nepal, Taiwan. Ill. Cat. Noctuidae Korea, 1998 _____. Kim and Choi, 2000.

DISTRIBUTION. Korea, China, Taiwan, Japan, Philippines, Indonesia, India, Nepal. NOTE. Reported as a new for Korea by Kim and Choi (2000). The male specimen was not

108

available for dissection. Additiona material examined: 1 male, 1 female, Korea, Prov. Gangwon, Chuncheon, Gangwon Natl. Univ. 8. IX 2000 (J. C. Sohn). 755. Apamea crenata (Hufnagel, 1766)*


Phalaena crenata Hufnagel, 1766, Berl. Mag. 3: 402 (TL: Germany, Berlin). - Apamea crenata ssp. crenata (Hufnagel). Eurasia. Noctua rurea (Fabricius, 1775) Syst. ent.: 618 (TL: England). Phalaena, Noctua lutulenta Esper, 1788, Die Schmett. n. Abb. Nat. 4(1): pl. 133: 3, 4 (TL: Germany, Leipzig, [ZSM, München]). Phalaena, Noctua alopecurus Esper, 1788, Die Schmett. n. Abb. Nat. 4(2): pl. 472: 3 (TL.: Austria, Tirol [ZSM, München]). Noctua combusta Hübner, 1803-1808, Samml. Eur. Schmett. Noct. 2: pl. 79: 366 (TL: Europe). Noctua hirticornus Haworth, 1812, Trans. ent. Soc. London 1: 336 (TL: Great Britain). Hadena crenata ssp. corsa Schawerda, 1928, Z Öst. ent. Ver. 13: 43 (TL: Korsika, Mt. Incudine [LM, Karlsruhe]). Apamea crenata ssp. corsa (Schawerda). - Korsika. Ill. Cat. Noctuidae Korea, 1998 _____.

DISTRIBUTION. Korea (North, first record), Japan, China (to Tibet), RFE (Primorye terr., Khabarovsk terr., Amur reg., Sakhalin, Kuriles), S Siberia (Transbaikalia, Baikal, Tuva, Altai), W Siberia, Ural, European part of Russia, E and W Europe (from Fennoscandia to Spain and Italy), Caucasus, Turkey. NOTE. The speices is reported here as a new for Korea. Material examined:1 male [Korea] Choangy [ZFMK, Bonn]. 756. Apamea sordens (Hufnagel, 1766)



757. Apamea brunnescens Kononenko, 1985



758. Apamea hampsoni Sugi, 1963

� genit.: Pl. 128, fig. 2 ♀ genit.: Pl. 264, fig. 2


759. Apamea veterina (Lederer, 1853)

� genit.: Pl. 128 fig. 4; ♀ genit.: Pl. 264, fig. 4


760. Apamea rubrirena (Treitschke, 1825)



761. Apamea scolopacina (Esper, 1788)



Genus LATEROLIGIA Zilli, Ronkay & Fibiger, 2005 (Type species: Noctua ophiogramma Esper, 1794 [Europe]). 762. Lateroligia ophiogramma (Esper, 1794) � genit.: Pl. 129, fig. 1; ♀ genit.: Pl. 264, fig. 7 Ill. Cat. Noctuidae Korea, 1998: 208; No. 538 (Apamea ophiogramma).

109

Genus LEUCAPAMEA Sugi, 1982 763. Leucapamea kawadai (Sugi, 1955) � genit.: Pl. 129, fig. 2; ♀ genit.: Pl. 264, fig. 8 Ill. Cat. Noctuidae Korea, 1998: 210; No. 543.

764. Leucapamea askoldis (Oberthür, 1880)



Genus ANAPAMEA Sugi, 1982 765. Anapamea cuneatoides Poole, 1989



766. Anapamea apameoides (Draudt, 1950)



767. Anapamea incerta (Staudinger, 1892)



Genus XENAPAMEA Sugi, 1970 768. Xenapamea pacifica (Sugi, 1970)



Genus CHANDATA Moore, 1882 769. Chandata bella (Butler, 1881) � genit.: Pl. 130, fig. 1; ♀ genit.: Pl. 265, fig. 5 Ill. Cat. Noctuidae Korea, 1998: 226; No. 586.

Genus MESAPAMEA Heineman, 1959 770. Mesapamea concinnata Heinicke, 1959 � genit.: Pl. 130, fig. 2; ♀ genit.: Pl. 265, fig. 6 Ill. Cat. Noctuidae Korea, 1998: 213; No. 548.

Genus XYLOMOIA Staudinger, 1892 771. Xylomoia graminea (Graeser, 1888 [1889]) � genit.: Pl. 130, fig. 3; ♀ genit.: Pl. 265, fig. 7 Ill. Cat. Noctuidae Korea, 1998: 215; No. 556.

Genus LITOLIGIA Beck, 1999 (Type species: Noctua literosa Haworth, 1809 [England]). 772. Litoligia fodinae (Oberthür, 1880), comb. n. � genit.: Pl. 130, fig. 4; ♀ genit.: Pl. 265, fig. 8 Ill. Cat. Noctuidae Korea, 1998: 211; No. 545.

NOTE. The species is congeneric with Litholigia literosa (Haworth, 1809), which is the type species of the genus Litoligia. Genus OLIGIA Hübner, [1821] 1816 773. “Oligia” leuconephra Hampson, 1908 � genit.: Pl. 130, fig. 5; ♀ genit.: Pl. 266, fig. 3 Ill. Cat. Noctuidae Korea, 1998: 211; No. 546.

NOTE. The species is not congeneric with Oligia strigilis (Linnaeus, 1758), which is the type

110

species of the genus Oligia. Its systematic position requires a further revision. Genus SAPPORIA Sugi, 1982 774. Sapporia repetita (Butler, 1885) � genit.: Pl. 130, fig. 6; ♀ genit.: Pl. 266, fig. 1 Ill. Cat. Noctuidae Korea, 1998: 213; No. 549.

Genus XANTHOGRAPTA Hampson, 1910 775. “Xanthograpta” basinigra Sugi, 1982 � genit.: Pl. 131, fig. 1; ♀ genit.: Pl. 266, fig. 2 Ill. Cat. Noctuidae Korea, 1998: 169; No. 439.

NOTE. Sugi (1982) described basinigra in the genus Xanthograpta in Acontiinae (sensu auctorum). However, the species seems to be not congeneric with Xanthograpta trilatalis Walker, [1866] 1865 [India]. Its systematic position requires a further revision. Subtribe SESAMIINA Fibiger & Lafontaine, 2005 Genus VIRGO Staudinger, 1892 (pos. prov.) 776. Virgo datanidia (Butler, 1885) � genit.: Pl. 131, fig. 2; ♀ genit.: Pl. 266, fig. 4 Ill. Cat. Noctuidae Korea, 1998: 260; No. 689.

Genus SESAMIA Guenée, 1852 777. Sesamia turpis (Butler, 1879) � genit.: Pl. 131, fig. 3; ♀ genit.: Pl. 266, fig. 5 Ill. Cat. Noctuidae Korea, 1998: 221; No. 573.

778. Sesamia pseudoturpis Kononenko & Ahn, 1998



779. Sesamia confusa (Sugi, 1982)*


Epipsammia confusa Sugi, 1982, Moths of Japan, 1: 752, Pl. 185: 16-18 (TL: Japan, Tsushima I. Mitsushima-cho, Gaya [NIAES, Tsukuba]). Ill. Cat. Noctuidae Korea, 1998 _____.

DISTRIBUTION. Korea, Japan, South of RFE (Primorye terr.). NOTE. First record in Korea. Material examined: 1 male: Korea, Prov. Kunggi, DMZ. Daseongdong, 15-16. V. 1997 (H. S. Kim); 1 female, Korea, Beakryung Is., Junghwa-dong GG, 23. VIII 1995 (M. S. Go and G. G. Lee). 780. Sesamia inferens (Walker, 1856)



Genus ACRAPEX Hampson, 1894 781. Acrapex azumai Sugi, 1970



NOTE. The female specimen was not available for dissection. 782. Acrapex klapperichi Kononenko, 1999

� genit.: Pl. 132, fig. 2 111

Acrapex klapperichi Kononenko, 1999, Japan Heterecerist’s J. 206: 112, figs 1-2 (TL: China, Provinces Jangsu, Fujian [ZFMK, Bonn]). Ill. Cat. Noctuidae Korea, 1998 _____.

DISTRIBUTION. Korea, SE China (Prov. Jangsu, Fujian). NOTE. The species has been described from South East China. Ths is first record after its description. First report for Korea. Material examined: 1 male, S Korea, Is. Yeongjiong-do (Yongyu), 22. VI 1993 (E. J. Hong) � genit.: Pl. 132, fig. 3

783. Acrapex owadai Kononenko, 1999

Acrapex owadai Kononenko, 1999, Japan Heterecerist’s J. 207: 112, figs 5-6 (TL: N Thailand [NSM, Tokyo]). Ill. Cat. Noctuidae Korea, 1998 _____.

DISTRIBUTION. Korea, North Thailand. NOTE. First report for Korea. Material examined: 1 male, S Korea, Is. Yeongjiong-do (Yongyu), Incheon, 22. VI 1993 (S. H. Yoo). Tribe XYLENINI Guenée, 1837 Subtribe XYLENINA Guenée, 1837 Genus PARASTICHTIS Hübner, [1821] 1816 784. Parastichtis suspecta (Hübner, [1817]) � genit.: Pl. 133, fig. 6; ♀ genit.: Pl. 267, fig. 1 Ill. Cat. Noctuidae Korea, 1998: 212; No. 547.

Genus TILIACEA Tutt, 1896 785. Tiliacea japonago (Wileman & West, 1929) � genit.: Pl. 132, fig. 4; ♀ genit.: Pl. 267, fig. 2 Ill. Cat. Noctuidae Korea, 1998: 294; No. 785 [Xanthia (Tiliacea) japonago].

786. Tiliacea auragides (Draudt, 1950)


Cosmia auragides Draudt, 1950, Mitt. Münch, ent. Ges. 40: 82, Taf. 6: 2 (TL: China, West Tien-mu-shan [ZFMK, Bonn]). Cirrhia tigrina Kononenko, 1978, Ent. Obozr. 57(4): 898, figs 12, 13 (TL: RFE: Primorye terr., Tigrovoi [ZI, St. Petersburg]). Ill. Cat. Noctuidae Korea, 1998: 294; No. 786 [Xanthia (Tiliacea) tigrina].

NOTE. For the synonymy cited see Kononenko (2005). Genus XANTHIA Ochsenheimer, 1816 787. Xanthia togata (Esper, 1788) � genit.: Pl. 132, fig. 6; ♀ genit.: Pl. 267, fig. 4 Ill. Cat. Noctuidae Korea, 1998: 293; No. 780.

Genus CIRRHIA Hübner, [1821] 1816 788. Cirrhia fasciata (Kononenko, 1978) � genit.: Pl. 133, fig. 1; ♀ genit.: Pl. 267, fig. 5 Ill. Cat. Noctuidae Korea, 1998: 293; No. 781 [Xanthia (Cirrhia) fasciata].

789. Cirrhia icteritia (Hufnagel, 1766)


Ill. Cat. Noctuidae Korea, 1998: 293; No. 782 [Xanthia (Cirrhia) icteritia].

112

790. Cirrhia tunicata Graeser, 1889 [1890]


Ill. Cat. Noctuidae Korea, 1998: 294; No. 783 [Xanthia (Cirrhia) tunicata].

791. Cirrhia ocellaris (Borkhausen, 1792)


Ill. Cat. Noctuidae Korea, 1998: 294; No. 784 [Xanthia (Cirrhia) ocellaris].

Genus AGROCHOLA Hübner [1821] 1816 792. Agrochola evelina (Butler, 1879) � genit.: Pl. 133, fig. 5; ♀ genit.: Pl. 268, fig. 2 Ill. Cat. Noctuidae Korea, 1998: 289; No. 771.

NOTE. The species was misplaced to the genus Himalistra Hacker, 1992 by Kononenko (2005). 793. Agrochola vulpecula (Lederer, 1853)



Genus HYALOBOLE Warren, 1911 794. Hyalobole albimacula (Kononenko, 1978) � genit.: Pl. 134, fig. 1; ♀ genit.: Pl. 268, fig. 3 Ill. Cat. Noctuidae Korea, 1998: 290; No. 772.

Genus TELORTA Warren, 1910 795. Telorta edentata (Leech, 1889) � genit.: Pl. 134, fig. 2; ♀ genit.: Pl. 268, fig. 4 Ill. Cat. Noctuidae Korea, 1998: 295; No. 787.

796. Telorta divergens (Butler, 1879)



797. Telorta acuminata (Butler, 1878)



Genus CONISTRA Hübner, [1821] 1816 Subgenus CONISTRA Hübner, [1821] 1816 798. Conistra ardescens (Butler, 1879) � genit.: Pl. 134, fig. 5; ♀ genit.: Pl. 268, fig. 7 Ill. Cat. Noctuidae Korea, 1998: 290; No. 773.

799. Conistra fletcheri Sugi, 1958



800. Conistra grisescens Draudt, 1950



Subgenus DASYCAMPA Guenée, 1837 801. Conistra albipuncta (Leech, 1889) � genit.: Pl. 135, fig. 3; ♀ genit.: Pl. 269, fig. 1 Ill. Cat. Noctuidae Korea, 1998: 291; No. 776.

802. Conistra filipjevi Kononenko, 1978



113

803. Conistra castaneofasciata (Motsch., [1861] 1860)



804. Jodia sericea (Butler, 1878)

Genus JODIA Hübner, 1818 � genit.: Pl. 136, fig. 1; ♀ genit.: Pl. 269, fig. 4


Genus TERATOGLAEA Sugi, 1958 805. Teratoglaea pacifica Sugi, 1958 � genit.: Pl. 136, fig. 2; ♀ genit.: Pl. 269, fig. 5 Ill. Cat. Noctuidae Korea, 1998: 287; No. 768.

Genus LITHOPHANE Hübner, [1821] 1816 806. Lithophane ustulata (Butler, 1878) � genit.: Pl. 136, fig. 3; ♀ genit.: Pl. 269, fig. 6 Ill. Cat. Noctuidae Korea, 1998: 279; No. 744.

807. Lithophane pruinosa (Butler, 1878)



808. Lithophane lamda (Fabricius, 1787)



809. Lithophane consocia (Borkhausen, 1792)


Ill. Cat. Noctuidae Korea, 1998: 280; No. 747 (Lithophane hepatica).

810. Lithophane plumbealis (Matsumura, 1926)



811. Lithophane venusta (Leech, 1889)



812. Lithophane rosinae (Püngeler, 1906)



813. Lithophane socia


Ill. Cat. Noctuidae Korea, 1998: 281; No. 751 (Lithophane hepatica).

NOTE. According to Mikkola and Honey (1985) and Mikkola (1993), Phalaena hepatica Cl., 1759 is a senior synonym of Lithophane socia Hfn., 1766 (a lectotype of the former has been designated), but Fibiger and Hacker (1991), Nowacky and Fibiger (1998) and Fibiger (pers. comm.) consider the lectotype designation by Mikkola and Honey (1985) is doubtful, and regarded hepatica as a senior synonym of Polia tincta (Brahm, 1790). In such case of different opinions between specialists, the ICZN should be asked for a decision to establish stability. M. Fibiger (pers. comm.) informed me that he applied to International Comission on Zoological Nomencalature for a solution of this problem. Pending the decision of ICZN and following to Ronkay et al., 2001 and Hacker and Fibiger, 2005, we use the name socia for this taxon, instead of hepatica taking in account the conservation of the name which has long been used for this taxon. 114

814. Lithophane remota Hreblay & Ronkay, 1998



815. Lithophane nagaii Sugi, 1958



816. Lithophane pacifica Kononenko, 1978



Genus LITHOMOIA Hübner, [1821] 1816 817. Lithomoia solidaginis (Hübner [1803]) � genit.: Pl. 138, fig. 4; ♀ genit.: Pl. 271, fig. 1 Ill. Cat. Noctuidae Korea, 1998: 279; No. 743.

Genus XYLENA Ochsenheimer, 1816 818. Xylena exsoleta (Butler, 1878) � genit.: Pl. 138, fig. 5; ♀ genit.: Pl. 271, fig. 2 Ill. Cat. Noctuidae Korea, 1998: 275; No. 732.

819. Xylena formosa (Butler, 1878)



820. Xylena confusa Kononenko & Ronkay, 1998




821. Xylena nihonica Höne, 1917*

Xylena japonica Höne, 1917, Ent. Mag. Kyoto, 3: 48, Pl. 1: 3a, 3b (TL: Japan, Kobe [ZFMK, Bonn]), nec Neuberger, 1903. Xylena nihonica Höne, 1917, Ent. Mag. Kyoto, 3: 50, Pl. 1: 4a, 4b, repl. name. Ill. Cat. Noctuidae Korea, 1998 _____. Sohn, Ronkay and Choi, 2005.

DISTRIBUTION. Korea, Japan, Taiwan. NOTE. The female specimen was not available for dissection. Genus ORBONA Hübner, [1821] 1816 822. Orbona fragariae (Vieweg, 1790) � genit.: Pl. 139, fig. 4; ♀ genit.: Pl. 271, fig. 5 Ill. Cat. Noctuidae Korea, 1998: 276; No. 735.

Genus EUPSILIA Hübner, [1821] 1816 823. Eupsilia transversa (Hufnagel, 1766) � genit.: Pl. 139, fig. 5; ♀ genit.: Pl. 271, fig. 6 Ill. Cat. Noctuidae Korea, 1998: 277; No. 736.

824. Eupsilia quadrilinea (Leech, 1889)



825. Eupsilia strigifera Butler, 1879



115

826. Eupsilia silla Kononenko & Ahn, 1998



827. Eupsilia tripunctata Butler, 1878



828. Eupsilia boursini Sugi, 1958*


Eupsilia boursini Sugi, 1958, Tinea, 4: 205, Pl. 31: 17 (HTm: Japan, Nagano Pref., Shiga Hights [NIAES, Tsukuba]). Ill. Cat. Noctuidae Korea, 1998 _____. Han, Kononenko and Park, 2005.

DISTRIBUTION. Korea, Japan, RFE (Primorye terr.). 829. Eupsilia hidakaensis Sugi, 1987



830. Eupsilia contracta (Butler, 1878)



Genus PYGOPTERYX Staudinger, 1887 831. Pygopteryx suava Staudinger, 1887 � genit.: Pl. 141, fig. 3; ♀ genit.: Pl. 272, fig. 6 Ill. Cat. Noctuidae Korea, 1998: 232; No. 605.

Genus ANTIVALERIA Sugi, 1958 832. Antivaleria viridimacula (Graeser, 1888 [1889]) � genit.: Pl. 141, fig. 4; ♀ genit.: Pl. 272, fig. 7 Ill. Cat. Noctuidae Korea, 1998: 286; No. 764.

Genus NYCTYCIA Hampson, 1906 833. Nyctycia sugii Ronkay, 1990 � genit.: Pl. 142, fig. 3; ♀ genit.: Pl. 273, fig. 4 Ill. Cat. Noctuidae Korea, 1998: 287; No. 765.

Genus RHYNCHAGLAEA Hampson, 1906 834. Rhynchaglaea fuscipennis Sugi, 1958 � genit.: Pl. 142, fig. 4; ♀ genit.: Pl. 273, fig. 5 Ill. Cat. Noctuidae Korea, 1998: 287; No. 766.

835. Rhynchaglaea scitula (Butler, 1879)*


Xylophasia scitula Butler, 1879, Ann. Mag. nat. Hist. (5)4: 359 (TL: Japan, Yokohama, [BMNH, London]). Xylophasia kumamotonis Matsumura, 1926, Ins. matsum. 1: 54, Pl. 1: 30 (TL: Japan, Kyushu, Kumamoto [EIHU, Sapporo]). Xylophasia scitula f. albibasis Draudt, 1934, in Seitz; 3, suppl. 148, Pl. 18e, form. Ill. Cat. Noctuidae Korea, 1998 _____. Sohn, Ronkay and Choi, 2005.

DISTRIBUTION. Korea, Japan, China, Taiwan. Genus HEMIGLAEA Sugi, 1980 836. Hemiglaea costalis (Butler, 1789) Ill. Cat. Noctuidae Korea, 1998: 287; No. 767.

116


NOTE. The male specimen was not available for dissection. Genus SUGITANIA Matsumura, 1926 837. Sugitania clara Sugi, 1990 � genit.: Pl. 143, fig. 1; ♀ genit.: Pl. 273, fig. 7 Ill. Cat. Noctuidae Korea, 1998: 288; No. 769.

Subtribe COSMIINA Guenée, 1852 (IPIMORPHINI Beck, 1989) Genus ENARGIA Hübner, [1821] 1816 838. Enargia paleacea (Esper, 1788) � genit.: Pl. 112, fig. 3; ♀ genit.: Pl. 253, fig. 6 Ill. Cat. Noctuidae Korea, 1998: 246; No. 641.

Genus IPIMORPHA Hübner, [1821] 1816 839. Ipimorpha retusa (Linnaeus, 1761) � genit.: Pl. 112, fig. 4; ♀ genit.: Pl. 253, fig. 7 Ill. Cat. Noctuidae Korea, 1998: 251; No. 656.

840. Ipimorpha subtusa ([Den. & Schiff.], 1775)



841. Ipimorpha contusa (Freyer, 1849)



Genus COSMIA Ochsenheimer, 1816 Subgenus CALYMNIA Hübner, [1823] 1816 842. Cosmia affinis (Linnaeus, 1767) � genit.: Pl. 113, fig. 1; ♀ genit.: Pl. 254, fig. 2 Ill. Cat. Noctuidae Korea, 1998: 247; No. 642.

843. Cosmia unicolor (Staudinger, 1892)



844. Cosmia cara (Butler, 1881)



845. Cosmia restituta Walker, 1857



846. Cosmia pyralina ([Den. & Schiff.], 1775)



847. Cosmia inconspicua (Draudt, 1950)



848. Cosmia achatina Butler, 1879



117

Subgenus COSMIA Ochsenheimer, 1816 849. Cosmia camptostigma (Ménétriés, 1859) � genit.: Pl. 114, fig. 1; ♀ genit.: Pl. 255, fig. 1 Ill. Cat. Noctuidae Korea, 1998: 248; No. 648.

850. Cosmia trapezina (Linnaeus, 1758)



851. Cosmia moderata (Staudinger, 1888)



852. Cosmia sanguinea Sugi, 1955



Genus DIMORPHICOSMIA Sugi, 1982 853. Dimorphicosmia variegata (Oberthür, 1879) � genit.: Pl. 114, fig. 6; ♀ genit.: Pl. 255, fig. 5 Ill. Cat. Noctuidae Korea, 1998: 250; No. 654.

Genus XANTHOCOSMIA Sugi, 1982 854. Xanthocosmia jankowskii (Oberthür, 1884) � genit.: Pl. 115, fig. 1; ♀ genit.: Pl. 255, fig. 6 Ill. Cat. Noctuidae Korea, 1998: 250; No. 653.

Genus CHASMINODES Hampson, 1908 855. Chasminodes albonitens (Bremer, 1861) � genit.: Pl. 115, fig. 2; ♀ genit.: Pl. 255, fig. 7 Ill. Cat. Noctuidae Korea, 1998: 252; No. 659.

856. Chasminodes bremeri Sugi & Kononenko, 1981



857. Chasminodes sugii Kononenko, 1981



858. Chasminodes aino Sugi, 1956



859. Chasminodes pseudalbonitens Sugi, 1955



860. Chasminodes ussurica Kononenko, 1982



861. Chasminodes cilia (Staudinger, 1888)



862. Chasminodes nigrilinea (Leech, 1889) Ill. Cat. Noctuidae Korea, 1998: 253; No. 666.

118


863. Chasminodes atrata (Butler, 1884)



Genus PSEUDOCOSMIA Kon., 1985 864. Pseudocosmia maculata Kononenko, 1985



NOTE. The female specimen was not available for dissection.

865. Antha grata (Butler, 1881)

Genus ANTHA Staudinger, 1892 � genit.: Pl. 116, fig. 6; ♀ genit.: Pl. 257, fig. 1


Genus GYROSPILARA Kon., 1989 866. Gyrospilara formosa (Graeser, 1888 [1889]) � genit.: Pl. 117, fig. 1; ♀ genit.: Pl. 257, fig. 2 Ill. Cat. Noctuidae Korea, 1998: 223; No. 577.

Subtribe ANTITYPINA Forbes & Franclemont, 1954 Genus DRYOBOTODES Warren, 1910 867. Dryobotodes pryeri (Leech, 1900) � genit.: Pl. 141, fig. 5; ♀ genit.: Pl. 273, fig. 1 Ill. Cat. Noctuidae Korea, 1998: 283; No. 755.

868. Dryobotodes intermissa (Butler, 1886)



869. Dryobotodes angusta Sugi, 1980



Genus DASYPOLIA Guenée, 1852 870. Dasypolia fani Staudinger, 1892 � genit.: Pl. 143, fig. 2; ♀ genit.: Pl. 273, fig. 8 Ill. Cat. Noctuidae Korea, 1998: 284; No. 758.

Genus POLYMIXIS Hübner, [1820] 1816 871. Polymixis mandshurica Boursin, 1970 � genit.: Pl. 143, fig. 3; ♀ genit.: Pl. 274, fig. 1 Ill. Cat. Noctuidae Korea, 1998: 286; No. 763.

Genus BLEPHARITA Hampson, 1907 872. Blepharita amica (Treitschke, 1825) � genit.: Pl. 143, fig. 4; ♀ genit.: Pl. 274, fig. 2 Ill. Cat. Noctuidae Korea, 1998: 284; No. 759.

Genus MNIOTYPE Franclemont, 1941 873. Mniotype bathensis (Lutzau, 1900) � genit.: Pl. 144, fig. 1; ♀ genit.: Pl. 274, fig. 3 Ill. Cat. Noctuidae Korea, 1998: 285; No. 760.

874. Mniotype satura ([Den. & Schiff.], 1775)

� genit.: Pl. 144, fig. 2; ♀ genit.: Pl. 274, fig. 4 119


875. Mniotype melanodonta (Hampson, 1906)



Subfamily HADENINAE Guenée, 1837 Tribe ORTHOSIINI Guenée, 1837 Genus PANOLIS Hübner, 1816 876. Panolis japonica Draudt, 1935 � genit.: Pl. 144, fig. 4; ♀ genit.: Pl. 274, fig. 7 Ill. Cat. Noctuidae Korea, 1998: 333; No. 878.

Genus DIOSZEGHYANA Hreblay, 1993 877. Dioszeghyana mirabilis (Sugi, 1955) � genit.: Pl. 145, fig. 1; ♀ genit.: Pl. 274, fig. 8 Ill. Cat. Noctuidae Korea, 1998: 333; No. 877.

Genus CLAVIPALPULA Staudinger, 1892 878. Clavipalpula aurariae (Oberthür, 1880) � genit.: Pl. 145, fig. 2; ♀ genit.: Pl. 275, fig. 1 Ill. Cat. Noctuidae Korea, 1998: 325; No. 855.

Genus XYLOPOLIA Sugi, 1982 879. Xylopolia bellula Kononenko & Ronkay, 1995 � genit.: Pl. 145, fig. 3; ♀ genit.: Pl. 275, fig. 2 Ill. Cat. Noctuidae Korea, 1998: 325; No. 856.

880. Xylopolia bella (Butler, 1881)



Genus ORTHOSIA Ochsenheimer, 1816 Subgenus ORTHOSIA Ochsenheimer, 1816 881. Orthosia incerta (Hufnagel, 1766) � genit.: Pl. 145, fig. 5; ♀ genit.: Pl. 275, fig. 4 Ill. Cat. Noctuidae Korea, 1998: 329; No. 864.

882. Orthosia evanida (Butler, 1879)



Subgenus MICRORTHOSIA 883. Orthosia paromoea (Hampson, 1905) � genit.: Pl. 146, fig. 3; ♀ genit.: Pl. 275, fig. 7 Ill. Cat. Noctuidae Korea, 1998: 320; No. 867.

Subgenus CORORTHOSIA Berio, 1980 884. Orthosia ella (Butler, 1878) � genit.: Pl. 146, fig. 4; ♀ genit.: Pl. 275, fig. 8 Ill. Cat. Noctuidae Korea, 1998: 320; No. 869.

885. Orthosia lizetta (Butler, 1878) 120



Subgenus EUCHORISTEA Warren, 1910 886. Orthosia limbata (Butler, 1879) � genit.: Pl. 146, fig. 5; ♀ genit.: Pl. 276, fig. 1 Ill. Cat. Noctuidae Korea, 1998: 331; No. 870.


887. Orthosia fausta Leech, 1889*

Orthosia fausta Leech, 1889, Proc. zool. Soc. London 1889: 513, Pl. 50: 2 (TL: Japan, Yokohama, BMNH, London). Ill. Cat. Noctuidae Korea, 1998 _____. Sohn and Kim, 2003.

DISTRIBUTION. Korea, Japan. 888. Orthosia nigromaculata (Höne, 1917)*


Monima nigromaculata Höne, 1917, Ent. Mag. Kyoto, 3: 47, Pl. 1: 2 (TL: Japan, Kobe [ZFMK, Bonn]). Monima pfennigschmidti Püngeler, 1925, Iris, 39: 232 (TL: Japan, Yokohama [MNHU, Berlin]). Ill. Cat. Noctuidae Korea, 1998 _____. Sohn and Ronkay, 2001.

DISTRIBUTION. Korea, Japan, China, Taiwan. Subgenus ERYTHROTIS Bryk, 1948871. 889. Orthosia cedermarki (Bryk, 1948) � genit.: Pl. 147, fig. 3; ♀ genit.: Pl. 276, fig. 3 Ill. Cat. Noctuidae Korea, 1998: 331; No. 871.

890. Orthosia carnipennis (Butler, 1878)




891. Orthosia satoi Sugi, 1960*

Orthosia satoi Sugi, 1960, Tinea, 5: 328, figs 1, 2 (HT: Japan, Tokyo, [NIAES, Tsukuba]). Ill. Cat. Noctuidae Korea, 1998 ?. Han, Kononenko and Park, 2005.

DISTRIBUTION. Korea, Japan, RFE (Primorye terr.). Subgenus SEMIOPHORA Stephens, 1829 892. Orthosia askoldensis (Staudinger, 1892) � genit.: Pl. 148, fig. 1; ♀ genit.: Pl. 276, fig. 6 Ill. Cat. Noctuidae Korea, 1998: 331; No. 873.

893. Orthosia odiosa (Butler, 1878)

Subgenus (undescribed) � genit.: Pl. 148, fig. 2; ♀ genit.: Pl. 276, fig. 7


Subgenus (undescribed) 894. Orthosia coniortota (Filipjev, 1927) � genit.: Pl. 148, fig. 3; ♀ genit.: Pl. 276, fig. 8 Ill. Cat. Noctuidae Korea, 1998: 332; No. 875.

Genus ANORTHOA Berio, 1980 895. Anorthoa munda ([Den. & Schiff.], 1775) � genit.: Pl. 148, fig. 4; ♀ genit.: Pl. 277, fig. 1 Ill. Cat. Noctuidae Korea, 1998: 327; No. 860 [Perigrapha (Anorthoa) munda].

121

896. Anorthoa angustipennis (Matsumura, 1926)


Ill. Cat. Noctuidae Korea, 1998: 330; No. 868 [Orthosia (Microrthosia) angustipennis].

Genus HARUTAEOGRAPHA Yoshimoto, 1993 897. Harutaeographa stenoptera (Staudinger, 1892) � genit.: Pl. 149, fig. 1; ♀ genit.: Pl. 277, fig. 3 Ill. Cat. Noctuidae Korea, 1998: 332; No. 876 (Orthosia stenoptera).

Genus PERIGRAPHA Lederer, 1857 898. Perigrapha hoenei Püngeler, 1914 � genit.: Pl. 149, fig. 2; ♀ genit.: Pl. 277, fig. 4 Ill. Cat. Noctuidae Korea, 1998: 327; No. 861.

899. Perigrapha extincta Kononenko, 1989



900. Perigrapha circumducta (Lederer, 1855)



901. Perigrapha uniformis Draudt, 1950



Genus PSEUDOPANOLIS Inaba, 1927 902. Pseudopanolis heterogyna (O. Bang-Haas, 1927) � genit.: Pl. 150, fig. 1; ♀ genit.: Pl. 277, fig. 8 Ill. Cat. Noctuidae Korea, 1998: 328; No. 863.

Tribe HADENINI Guenée, 1852 Genus PROTOMISELIA Sugi, 1958 903. Protomiselia bilinea (Hampson, 1905) � genit.: Pl. 150, fig. 2; ♀ genit.: Pl. 278, fig. 1 Ill. Cat. Noctuidae Korea, 1998: 325; No. 854.

NOTE. The female specimen was unavailable for dissection. Genus ANARTA Ochsenheimer, 1816 (TRICHOCLEA Grote, 1883, subg., HADULA Staudinger, 1889, DISCESTRA Hampson, 1905) Subgenus TRICHOCLEA Grote, 1883 (DISCESTRA Hampson, 1905) NOTE. The generic name Anarta (Hadula auct.) and generic synonymy are treated according to revision and subsequent corrections by Hacker (1998b), Hacker et al. (2001) and Fibiger and Hacker (2005). 904. Anarta trifolii (Hufnagel, 1766)



905. Anarta stigmosa (Christoph, 1889)*


Mamestra stigmosa Christoph, 1889, in Rom; Mém. Lép. 3: 70, Pl. 3: 12 (TL: S Russia, ZI, St. Petersbourg)

122

Ill. Cat. Noctuidae Korea, 1998 _____. Sohn, Ronkay, Kim and Cho, 2005.

DISTRIBUTION. Korea, China, Mongolia, Central Asia, Kazakstan, South Siberia, South of European part of Russia, South Europe, Turkey. Genus POLIA Ochsenheimer, 1816 906. Polia bombycina (Hufnagel, 1766) � genit.: Pl. 150, fig. 5; ♀ genit.: Pl. 278, fig. 4 Ill. Cat. Noctuidae Korea, 1998: 308; No. 814.

907. Polia hepatica (Clerck, 1759)


Ill. Cat. Noctuidae Korea, 1998: 309; No. 815 (Polia trimaculosa).

NOTE. According to Mikkola and Honey (1985) and Mikkola (1993), the name trimaculosa is an ojective replacement name for Polia hepatica sensu auctorum, while the name hepatica Cl., 1759 is applied to Lithophane hepatica (with socia Hfn., 1766 in synonymy) (see Note for L. socia). However some other lepidopterists (Fibiger and Hacker, 1990) disagree to this treatment of the names hepatica and trimaculosa by Mikkola. The name hepatica has been applied to Polia hepatica and socia to Lithophane socia in the Lists of European Noctuidae (Hacker and Fibiger, 1990, Nowacki and Fibiger, 1997, Hacker and Fibiger, 2005). M. Fibiger (pers. comm.) informed us that he applied to ICZN for a solution of this nomenclature problem. Pending the final decision of the Comission we use the name Polia hepatica for this taxon. 908. Polia nebulosa (Hufnagel, 1766)



909. Polia goliath (Oberthür, 1880)



910. Polia mortua (Staudinger, 1888)



911. Polia malchani (Draudt, 1934)



912. Polia vespertilio (Draudt, 1934)



Genus LACANOBIA Billberg, 1820 Subgenus LACANOBIA Billberg, 1820 913. Lacanobia dentata (Kononenko, 1981) � genit.: Pl. 152, fig. 2; ♀ genit.: Pl. 279, fig. 4 Ill. Cat. Noctuidae Korea, 1998: 296; No. 296.

Subgenus DIANOBIA Behounek, 1993 914. Lacanobia contrastata (Bryk, 1942) � genit.: Pl. 152, fig. 3; ♀ genit.: Pl. 279, fig. 5 Ill. Cat. Noctuidae Korea, 1998: 297; No. 794.

123

915. Lacanobia contigua ([Den. & Schiff.], 1775)



Subgenus DIATARAXIA Hübner, [1821] 1816 916. Lacanobia oleracea (Linnaeus, 1758) � genit.: Pl. 152, fig. 4; ♀ genit.: Pl. 280, fig. 1 Ill. Cat. Noctuidae Korea, 1998: 297; No. 793.

917. Lacanobia splendens (Hübner, [1808])



Genus MELANCHRA Hübner, [1820] 1816 918. Melanchra persicariae (Linnaeus, 1761) � genit.: Pl. 153, fig. 2; ♀ genit.: Pl. 280, fig. 3 Ill. Cat. Noctuidae Korea, 1998: 305; No. 809.

919. Melanchra postalba Sugi, 1982



Genus HYPORBARATHRA Hampson, 1905 920. Hyporbarathra icterias (Eversmann, 1843) � genit.: Pl. 153, fig. 4; ♀ genit.: Pl. 280, fig. 5 Ill. Cat. Noctuidae Korea, 1998: 298; No. 796.

Genus PAPESTRA Sukhareva, 1973 921. Papestra biren (Goeze, 1781) � genit.: Pl. 153, fig. 5; ♀ genit.: Pl. 280, fig. 6 Ill. Cat. Noctuidae Korea, 1998: 307; No. 813.

Genus HYSSIA Guenée, 1852 922. Hyssia cavernosa (Eversmann, 1842) � genit.: Pl. 154, fig. 1; ♀ genit.: Pl. 280, fig. 7 Ill. Cat. Noctuidae Korea, 1998: 302; No. 801.

Genus MAMESTRA Ochsenheimer, 1816 923. Mamestra brassicae (Linnaeus, 1758) � genit.: Pl. 154, fig. 2; ♀ genit.: Pl. 280, fig. 8 Ill. Cat. Noctuidae Korea, 1998: 306; No. 812.

Genus SIDERIDIS Hübner, [1821] 1816 Subgenus COLONSIDERIDIS Beck, [1992] 1991 924. Sideridis turbida (Esper, [1790]) � genit.: Pl. 154, fig. 3; ♀ genit.: Pl. 281, fig. 4 Ill. Cat. Noctuidae Korea, 1998: 303; No. 802 (Sideridis albicolon).

NOTE. Hacker (1998a) stated the synonymy and priority the name turbida Esper, 1790 ower albicolon Hübner, [1813]. Subgenus SIDERIDIS Hübner, [1821] 1816 925. Sideridis incommoda (Staudinger, 1888) � genit.: Pl. 154, fig. 5; ♀ genit.: Pl. 281, fig. 2 Ill. Cat. Noctuidae Korea, 1998: 303; No. 803.

926. Sideridis unica (Leech, 1889) 124



Subgenus ANEDA Sukhareva, 1973 927. Sideridis honeyi Yoshimoto, 1989 � genit.: Pl. 154, fig. 4; ♀ genit.: Pl. 281, fig. 6 Ill. Cat. Noctuidae Korea, 1998: 303; No. 805.

Genus DICTYESTRA Sugi, 1982 928. Dictyestra dissecta (Walker, 1865) � genit.: Pl. 155, fig. 1; ♀ genit.: Pl. 281, fig. 1 Ill. Cat. Noctuidae Korea, 1998: 304; No. 806.

Genus CONISANIA Hampson, 1905 929. Conisania suavis (Staudinger, 1892) � genit.: Pl. 155, fig. 2; ♀ genit.: Pl. 281, fig. 5 Ill. Cat. Noctuidae Korea, 1998: 304; No. 807.

Genus NIABOMA Nye, 1975 930. Niaboma xena (Staudinger, 1896) Ill. Cat. Noctuidae Korea, 1998: 305; No. 808.

NOTE. The distribution of the species in Korea requires a further confirmation. Genus HADENA Schrank, 1802 Subgenus HADENA Schrank, 1802 931. Hadena compta ([Den. & Schiff.], 1775) � genit.: Pl. 155, fig. 3; ♀ genit.: Pl. 281, fig. 7 Ill. Cat. Noctuidae Korea, 1998: 299; No. 797.

932. Hadena variolata (Smith, 1888)



Subgenus ANEPIA Hampson, 1918 933. Hadena aberrans (Eversmann, 1856) � genit.: Pl. 155, fig. 5; ♀ genit.: Pl. 281, fig. 9 Ill. Cat. Noctuidae Korea, 1998: 301; No. 799.

934. Hadena corrupta (Herz, 1898)



Genus TIRACOLA Moore, 1881 935. Tiracola aureata Holloway, 1989 � genit.: Pl. 156, fig. 1; ♀ genit.: Pl. 282, fig. 2 Ill. Cat. Noctuidae Korea, 1998: 310; No. 310.

Tribe LEUCANINI Guenée, 1837 (MYTHIMNINI Rungs, 1956) Genus SARCOPOLIA Sugi, 1982 936. Sarcopolia illoba (Butler, 1878) � genit.: Pl. 156, fig. 2; ♀ genit.: Pl. 282, fig. 3 Ill. Cat. Noctuidae Korea, 1998: 306; No. 811.

125

Genus MYTHIMNA Ochsenheimer, 1816 Subgenus MYTHIMNA Ochsenheimer, 1816 937. Mythimna turca (Linnaeus, 1761) � genit.: Pl. 156, fig. 3; ♀ genit.: Pl. 282, fig. 4 Ill. Cat. Noctuidae Korea, 1998: 312; No. 822.

938. Mythimna monticola Sugi, 1958



939. Mythimna grandis Butler, 1878



940. Mythimna divergens Butler, 1878



941. Mythimna curvata Leech, 1900,



942. Mythimna placida (Butler, 1878)



943. Mythimna bani (Sugi, 1977)



944. Mythimna velutina (Eversmann, 1846)



945. Mythimna pudorina ([Den. & Schiff.], 1775)



946. Mythimna rufipennis Butler, 1878



947. Mythimna conigera ([Den. & Schiff.], 1775)



948. Mythimna pallens (Linnaeus, 1758)



949. Mythimna impura (Hübner, [1808])



950. Mythimna lineatipes Moore, 1881*



Subgenus PSEUDALETIA Franclemont, 1951 951. Mythimna separata (Walker, 1865) � genit.: Pl. 159, fig. 1; ♀ genit.: Pl. 284, fig. 2 126


Subgenus SABLIA Sukhareva, 1973 952. Mythimna nigrilinea (Leech, 1899) � genit.: Pl. 159, fig. 2; ♀ genit.: Pl. 284, fig. 3 Ill. Cat. Noctuidae Korea, 1998: 322; No. 848.

953. Mythimna albiradiosa (Eersmann, 1852)*


Leucania albiradiosa Eersmann, 1852; Bull. Soc. imp. Nat. Mosc. 25(1): 158 (TL: Russia, Volga region [ZI, St. Petersbourg]). Leucania seifersi RANGNOW, 1930, Dt. ent. Z. 1930: 73 (TL: Russia, SUral, Guberla). Ill. Cat. Noctuidae Korea, 1998 _____.

DISTRIBUTION. Korea, Russia: RFE, S Siberia, Altai, Ural, Wolga region, Mongolia, Kazakstan, Kyrgizia. NOTE. First record for Korea. Subgenus HYPHILARE Hübner, [1821] 1816 954. Mythimna flavostigma (Bremer, 1861) � genit.: Pl. 159, fig. 3; ♀ genit.: Pl. 284, fig. 4 Ill. Cat. Noctuidae Korea, 1998: 318; No. 840.

955. Mythimna radiata (Bremer, 1861)



956. Mythimna iodochra (Sugi, 1982)


Ill. Cat. Noctuidae Korea, 1998: 318; No. 841 (numbers 841 and 847 interchanged on plate).

NOTE. In the Catalogue of Korean Noctuidae (Kononenko et al., 1998), the numbers 841 and 847 on the plate (p. 453) are interchanged. The number 841c and 847a denote Mythimna iodochra; the numbers 841a and 841b denote Mythimna stolida. � genit.: Pl. 160, fig. 1

957. Mythimna salebrosa (Butler, 1878) Ill. Cat. Noctuidae Korea, 1998: 317; No. 837.

958. Mythimna chosenicola (Bryk, 1948)



959. Mythimna simplex (Leech, 1889)



960. Mythimna proxima (Leech, 1900)



Subgenus MORPHOPOLIANA Hreblay & Legrran, 1996 961. Mythimna stolida (Leech, 1889) � genit.: Pl. 160, fig. 5; ♀ genit.: Pl. 285, fig. 3 Ill. Cat. Noctuidae Korea, 1998: 321; No. 847 (number 847 interchanged with 841 on plate).

NOTE. The number 841 and 847 are interchanged on the plate (p. 453) in the Catalogue Korean

127

Noctuidae (Kononenko et al., 1998). The number 841a and 841b denote M. stolida, the number 847a denote M. iodochra. Subgenus ANAPOMA Berio, 1980 962. Mythimna postica (Hampson, 1905) � genit.: Pl. 161, fig. 1; ♀ genit.: Pl. 285, fig. 4 Ill. Cat. Noctuidae Korea, 1998: 323; No. 851.

Subgenus DYSALETIA Sugi, 1982 963. Mythimna inanis (Oberthür, 1880) � genit.: Pl. 161, fig. 2; ♀ genit.: Pl. 285, fig. 5 Ill. Cat. Noctuidae Korea, 1998: 324; No. 852.

Genus LEUCANIA Ochsenheimer, 1816 Subgenus ACANTHOLEUCANIA Rungs, 1953 964. Mythimna loreyi (Duponchel, 1827) � genit.: Pl. 161, fig. 3; ♀ genit.: Pl. 285, fig. 6 Ill. Cat. Noctuidae Korea, 1998: 323; No. 850.

Subgenus LEUCANIA Ochsenheimer, 1816 965. Mythimna compta (Moore, 1881) Ill. Cat. Noctuidae Korea, 1998: 319; No. 843.

NOTE. Reported as a new for Korea by Kononenko et al. (1998). The specimens were unavailable for dissection. 966. Mythimna striata Leech, 1900



967. Mythimna obsoleta (Hübner, [1803])



Genus SENTA Stephens, 1834 968. Senta flammea (Curtis, 1828) � genit.: Pl. 162, fig. 2; ♀ genit.: Pl. 286, fig. 1 Ill. Cat. Noctuidae Korea, 1998: 321; No. 846.

Tribe GLOTTULINI Guenée, 1852 Genus BRITHYS Hübner, 1816 969. Brithys crini (Fabricius, 1775) � genit.: Pl. 162, fig. 3; ♀ genit.: Pl. 286, fig. 2 Ill. Cat. Noctuidae Korea, 1998: 324; No. 853.

Subfamily NOCTUINAE Tribe AGROTINI Rambur, 1848 Subtribe AUSTRANDESIINA Angulo and Olivares, 1990 Genus PERIDROMA Hübner, [1821] 1816 970. Peridroma saucia (Hübner, [1808]) � genit.: Pl. 162, fig. 4; ♀ genit.: Pl. 286, fig. 3 128


Subtribe AGROTINA Rambur, 1848 Genus ACTEBIA Stephens, 1829 971. Actebia praecox (Linnaeus, 1758) � genit.: Pl. 162, fig. 5; ♀ genit.: Pl. 286, fig. 4 Ill. Cat. Noctuidae Korea, 1998: 359; No. 934.

972. Actebia praecurrens (Staudinger, 1888)



973. Actebia fennica (Tauscher, 1806)



Genus DICHAGYRIS Lederer, 1857 NOTE. In treatment of Albocosta as a subgenus of the genus Dichagyris we follow to Lafontaine and Fibiger (2004). Subgenus ALBOCOSTA Fibiger & Lafontaine, 1997 974. Dichagyris stentzi (Lederer, 1853) � genit.: Pl. 163, fig. 3; ♀ genit.: Pl. 287, fig. 1 Ill. Cat. Noctuidae Korea, 1998: 334; No. 880 (Albocosta stentzi).

975. Dichagyris triangularis (Moore, 1867)


Ill. Cat. Noctuidae Korea, 1998: 334; No. 881 (Albocosta triangularis).

Genus EUXOA Hübner, [1821] 1816 Subgenus CHORIZAGROTIS Smith, 1890 976. Euxoa adumbrata (Eversmann, 1842) � genit.: Pl. 163, fig. 5; ♀ genit.: Pl. 287, fig. 3 (Synonyms: inexpectata Alph., 1897; arenacea Kozh., 1923; friedeli Pinker, 1974; lidia auct.). Ill. Cat. Noctuidae Korea, 1998: 361; No. 937 (Euxoa lidia).

NOTE. Lafontaine (1987a) treated the taxon adumbrata as an eastern Palaearctic subspecies of the Holarctic species E. lidia. Fibiger (1997) raised the status of E. adumbrata to full species and separated E. adumbrata and E. lidia. According to Fibiger, E. lidia occurs only in north-eastern Europe. Lafontaine (1998) confirmed that E. lidia and E. adumbrata could be two distinct species. In the persent Checklist, we follow the latest treatment of this species proposed by Fibiger (1996) and accepted in the Checklist of the Noctuidae of Europe (Fibiger, Hacker, 2005). Subgenus EUXOA Hübner, [1821] 1816 977. Euxoa sibirica (Boisduval, 1834) � genit.: Pl. 164, fig. 1; ♀ genit.: Pl. 287, fig. 4 Ill. Cat. Noctuidae Korea, 1998: 362; No. 938.

978. Euxoa ochrogaster (Guenée, 1953)



129

979. Euxoa karschi (Graeser, 1889 [1890])



980. Euxoa novoobscurior Bryk, 1948



Genus AGROTIS Ochsenheimer, 1816 981. Agrotis ipsilon (Hufnagel, 1766) � genit.: Pl. 164, fig. 5; ♀ genit.: Pl. 287, fig. 8 Ill. Cat. Noctuidae Korea, 1998: 366; No. 942.

982. Agrotis exclamationis (Linnaeus, 1758)



983. Agrotis scotacra (Filipjev, 1927)



984. Agrotis tokionis Butler, 1881



985. Agrotis segetum ([Den. & Schiff.], 1775)



986. Agrotis clavis (Hufnagel, 1766)



987. Agrotis ruta (Eversmann, 1851)



988. Agrotis characteristica Alpheraky, 1892



989. Agrotis trifurca Eversmann, 1837



Tribe NOCTUINI Latreille, 1809 Subtribe AXYLIINA Fibiger & Lafontaine, 2005 Genus AXYLIA Hübner, [1821] 1816 990. Axylia putris (Linnaeus, 1761) � genit.: Pl. 166, fig. 4; ♀ genit.: Pl. 289, fig. 2 Ill. Cat. Noctuidae Korea, 1998: 333; No. 879.

Genus OCHROPLEURA Hübner, [1821] 1816 991. Ochropleura plecta (Linnaeus, 1761) � genit.: Pl. 166, fig. 5; ♀ genit.: Pl. 289, fig. 3 Ill. Cat. Noctuidae Korea, 1998: 335; No. 882.

130

Subtribe NOCTUINA Latreille, 1809 Genus DIARSIA Hübner, [1821] 1816 992. Diarsia deparca (Butler, 1879) � genit.: Pl. 167, fig. 1; ♀ genit.: Pl. 289, fig. 4 Ill. Cat. Noctuidae Korea, 1998: 336; No. 883.

993. Diarsia dahlii (Hübner, [1813])



994. Diarsia canescens (Butler, 1878)



995. Diarsia brunnea ([Den. & Schiff.], 1775)



996. Diarsia dewitzi (Graeser, 1888 [1889])



997. Diarsia pacifica Boursin, 1943



998. Diarsia ruficauda (Warren, 1909)



999. Diarsia albipennis (Butler, 1889)



Genus CERASTIS Ochsenheimer, 1816 1000. Cerastis pallescens (Butler, 1878) � genit.: Pl. 168, fig. 4; ♀ genit.: Pl. 290, fig. 4 Ill. Cat. Noctuidae Korea, 1998: 354; No. 924.

1001. Cerastis violetta Boursin, 1955



1002. Cerastis leucographa ([Den. & Schiff.], 1775)



1003. Cerastis rubricosa ([Den. & Schiff.], 1775)*


Noctua rubricosa Denis & Schiffermüller, 1775, Ankündung syst. Werkes Schmett. Wienergegend: 7 (Noctua) (TL: Austria, Wien). Phalaena Noctua mucida Esper, [1790], Die Schmett. n. Abb. Nat. 4(2/1): 482, Pl. 184, Noct. 69, fig. 4 (TL: Austria, Tirol; vicinity of Wien [LT: Nr. 875, ZSM, München]). Phalaena, Noctua pilicornis Brahm, 1791, Ins. Kal. f. Samml. Oek. 2: 193 (TL: not stated). Noctua mista (Hübner, 1827) Samml. Eur. Schmett. Noct. 4: Pl. 109: 509 (TL: Europe). Ill. Cat. Noctuidae Korea, 1998 _____.

DISTRIBUTION. Korea (first record), Japan, RFE (Primorye), S Siberia, Central Asia, European part of Russia, Europe. 131

NOTE. The species is reported for Korea for the first time. Material examined: 1 male, North Korea, Genitalia Preparate 8942 (L. Ronkay), HNHM, Budapest. Genus PARADIARSIA McDunnough, [1829] 1004. Paradiarsia punicea (Hübner, 1803) � genit.: Pl. 169, fig. 2; ♀ genit.: Pl. 290, fig. 7 Ill. Cat. Noctuidae Korea, 1998: 342; No. 899.

Genus LYCOPHOTIA Hübner, [1821] 1816 1005. Lycophotia cissigma (Ménétriés, 1859) � genit.: Pl. 169, fig. 3; ♀ genit.: Pl. 290, fig. 8 Ill. Cat. Noctuidae Korea, 1998: 340; No. 895.

Genus CHERSOTIS Boisduval, 1840 1006. Chersotis deplanata (Eversmann, 1843) � genit.: Pl. 169, fig. 4; ♀ genit.: Pl. 295, fig. 6 Ill. Cat. Noctuidae Korea, 1998: 341; No. 896.

Genus HERMONASSA Walker, 1865 1007. Hermonassa cecilia Butler, 1878 � genit.: Pl. 170, fig. 1; ♀ genit.: Pl. 291, fig. 1 Ill. Cat. Noctuidae Korea, 1998: 341; No. 897.

1008. Hermonassa arenosa (Butler, 1881)



Genus CRYPTOCALA Benjamin, 1921 1009. Cryptocala chardinyi (Boisduval, 1829) � genit.: Pl. 169, fig. 5; ♀ genit.: Pl. 291, fig. 3 Ill. Cat. Noctuidae Korea, 1998: 357; No. 931.

Genus SPAELOTIS Boisduval, 1840 1010. Spaelotis ravida ([Den. & Schiff.], 1775) � genit.: Pl. 170, fig. 3; ♀ genit.: Pl. 291, fig. 4 Ill. Cat. Noctuidae Korea, 1998: 342; No. 901.

1011. Spaelotis nipona (Felder & Rogenhofer, 1874)


Agrotis nipona Felder & Rogenhofer, 1874, Reise öst. Freg. Novara, (Zool.) 2 (Abt. 2): 110, fig. 20 (TL: Japan [BMNH, London]). Graphiphora caliginea Butler, 1878, Ann. Mag. nat. Hist. (5)1: 165 (TL: Japan, Hakodate [BMNH, London]). Spaelotis valida auct., misidentification. Reference: Honey, 1996, Sugi, 2000. Ill. Cat. Noctuidae Korea, 1998: 343; No. 902 (Spaelotis valida Walker, 1865).

DISTRIBUTION. Korea, Japan (Hokkaido, Honshu), China. NOTE. Sugi (2000), based on presentation of M. Honey (1996), stated that Spaelotis niphona is not a synonym of S. ravida (according to Poole, 1989), but a valid taxon, distinct from S. valida. The name caliginea is not a synonym of S. valida, but a synonym of S. niphona. 1012. Spaelotis lucens Butler, 1881 Ill. Cat. Noctuidae Korea, 1998: 343; No. 903.

132


1013. Spaelotis suecia (Aurivillius, 1890)


Ill. Cat. Noctuidae Korea, 1998: 343; No. 904 (Spaelotis clandestina).

NOTE. We accept here the opinion of Fibiger (1997) who treated S. suecia as a Palaearctic species, disinct from the Nearctic S. clandestina. This treatment is also accepted in recent Cheklist of the Noctuidea of Europe (Fibiger, Hacker, 2005). 1014. Spaelotis deplorata (Staudinger, 1896)



Genus EUROIS Hübner, [1821] 1816 1015. Eurois occulta (Linnaeus, 1758) � genit.: Pl. 171, fig. 2; ♀ genit.: Pl. 292, fig. 1 Ill. Cat. Noctuidae Korea, 1998: 342; No. 900.

Genus ANAPLECTOIDES McDunnough, [1929] 1928 1016. Anaplectoides prasina ([Den. & Schiff.], 1775) � genit.: Pl. 171, fig. 3; ♀ genit.: Pl. 292, fig. 2 Ill. Cat. Noctuidae Korea, 1998: 356; No. 929.

1017. Anaplectoides virens (Butler, 1878)



Genus XESTIA Hübner, 1818 Subgenus MEGASEMA Hübner, [1821] 1816 1018. Xestia c-nigrum (Linnaeus, 1758) � genit.: Pl. 172, fig. 1; ♀ genit.: Pl. 292, fig. 4 Ill. Cat. Noctuidae Korea, 1998: 346; No. 906.

1019. Xestia ditrapezium ([Den. & Schiff.], 1775)



1020. Xestia kollari (Lederer, 1853)



1021. Xestia stupenda (Butler, 1878)



1022. Xestia baja ([Den. & Schiff.], 1775)


Noctua baja [Denis & Schiffermüller], 1775, Ank. Syst. Werk. Schmett. Wien, 77 (TL: Austria, Wien). Noctua baja (Fabricius, 1787) Mant. Ins. 2: 175. Phalaena Noctua tricomma (ESPER, [1791]) Schmett. Abb. n. Nat., 4(2/1): 603, Pl. 167, Noct. 88, fig. 6 (TL: Austria, Wien, Nr.861 [ZSM, München, LT designated by Hacker, 1998]). Agrotis punctata (Aurivillius, 1888) Nord. Fjär. 115 (TL: Lappland [RM, Stockholm]). Agrotis baja bajula Staudinger, 1881, Stettin. ent. Ztg. 1881: 411 (TL: “Lepsa” [Kyrgizia] [MNHU, Berlin]). Taeniocampa tabida Butler, 1878, Ann. Mag. nat. Hist. (5)1: 166 (TL: Japan: Yokohama [BMNH, London]). Agrotis baja nisseni Rotschild, 1912, Novit. Zool. 19: 125 (TL: Tunesia, Ain Draham [BMNH, London]). Rhyacia durandi Lucas, 1932, Bull. Soc. ent. Fr. 1932: 185 (TL: Algeria, [MNHN, Paris]). Hypoxestia ohtanensis Matsumura, 1925, J. Coll. Agric. Hokkaido Imp. Univ. 15(3): 144, pl. 8: 6 (TL: Sakhalin, Ichinosawa, Otani, Nyiwo, Rykowskoie [EIHU, Sapporo]).

133

Hypoxestia sachalinensis Matsumura, 1925, J. Coll. Agric. Hokkaido Imp. Univ. 15(3): 144, pl. 9: 1 (TL: RFE: S Sakhalin: Solovjevka [Ichinosawa]). Hypoxestia sachalinensis rikovskensis Matsumura, 1925, J. Coll. Agric. Hokkaido Imp. Univ. 15(3): 145, pl. 2: 17 (TL: RFE: N Sakhalin: Rykovskoe [Tymovskoe] [EIHU, Sapporo]). Agrotis baja cinigera Filipjev, 1927, Ann. Mus. zool. Acad. Sci. USSR, 28: 237 (TL: SE Siberia, Primorye terr., Bezverkhovo [Sidemi] [ZI, St. Petersburg]). Diarsia baja chosenbaja Bryk, 1948, Ark. Zool. 41(A) 1: 68 (TL: South Korea: Chidisan [Jiri Mt.] [NRM, Stockholm]). Ill. Cat. Noctuidae Korea, 1998: 347; No. 910 (Xestia tabida).

DISTRIBUTION. Korea (North, Central, South), Japan (Hokkaido, Honshu), China, Mongolia, Kazakstan, RFE (Primorye terr., S Khabarovsk reg., S Amur reg., Sakhalin, Kuriles, Kamchatka), Siberia, Ural, European part of Russia, Europe, Turkey, N Africa (Algeria, Morokko, Tunesia). NOTE. Transpalaearctic polymorphic species with numerous described local forms. The examination of the material from different part of its distribution revealed conspecifity of the taxa described as bajula, tabida, ohtanensis, sachalinensis, sachalinensis rikovskensis, baja cinigera and baja chosenbaja with Xestia baja (Kononenko, 2005 and present publication). The subspecific splitting of the species is not clarified yet. Reported in the Catalogue of Korean Noctuidae as Xestia tabida. � genit.: Pl. 172, fig. 6; ♀ genit.: Pl. 293, fig. 1

1023. Xestia dilatata (Butler, 1879) Ill. Cat. Noctuidae Korea, 1998: 348; No. 911.

1024. Xestia fuscostigma (Bremer, 1861)



1025. Xestia descripta (Bremer, 1861)



1026. Xestia vidua (Staudinger, 1892)



Subgenus (undescribed) 1027. Xestia semiherbida (Walker, 1857) � genit.: Pl. 173, fig. 4; ♀ genit.: Pl. 293, fig. 5 Ill. Cat. Noctuidae Korea, 1998: 352; No. 919.

1028. Xestia efflorescens (Butler, 1879)



1029. Xestia undosa (Leech, 1889)

Subgenus (undescribed) �� genit.: Pl. 174, fig. 1; ♀ genit.: Pl. 293, fig. 7


Subgenus PACHNOBIA Guenée, 1852 1030. Xestia speciosa (Hübner, [1813]) � genit.: Pl. 174, fig. 2; ♀ genit.: Pl. 293, fig. 8 Ill. Cat. Noctuidae Korea, 1998: 349; No. 349.

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1031. Xestia albonigra (Kononenko, 1981)



1032. Xestia sincera (Herrich-Schäffer, 1851)



1033. Xestia albuncula (Eversmann, 1851)



Genus EUGRAPHE Hübner, [1821] 1816 1034. Eugraphe sigma ([Den. & Schiff.], 1775) � genit.: Pl. 175, fig. 1; ♀ genit.: Pl. 294, fig. 4 Ill. Cat. Noctuidae Korea, 1998: 353; No. 922.

1035. Eugraphe versuta (Püngeler, 1908)



Genus SINEUGRAPHE Boursin, 1954 1036. Sineugraphe exusta (Butler, 1878) � genit.: Pl. 175, fig. 3; ♀ genit.: Pl. 294, fig. 6 Ill. Cat. Noctuidae Korea, 1998: 338; No. 891.

1037. Sineugraphe bipartita (Graeser, 1888 [1889])



1038. Sineugraphe oceanica (Kardakoff, 1928) � genit.: Pl. 176, fig. 1; ♀ genit.: Pl. 295, fig. 1 Ill. Cat. Noctuidae Korea, 1998: 338; No. 893.

1039. Sineugraphe stolidoprocta Boursin, 1954 � genit.: Pl. 176, fig. 2; ♀ genit.: Pl. 295, fig. 2 Ill. Cat. Noctuidae Korea, 1998: 340; No. 894.

Genus PROTOLAMPRA McDunnough, 1928 1040. Protolampra sobrina (Duponchel, 1843) � genit.: Pl. 176, fig. 3; ♀ genit.: Pl. 295, fig. 3 Ill. Cat. Noctuidae Korea, 1998: 357; No. 932.

NOTE. The record of this species in Korea require a further reconfirmation. The Eurasiatic boreal species Protolampra sobrina is distributed in Eastern Asia in Siberia and Northern part of Russian Far East. It has never been found in Japan or in the South of Russian Far East. However, as the taxon P. sobrina shuotsuensis (Bryk, 1948) was described from North Korea, it is included in the present Catalogue. The status of this taxon is yet uncertain. The holotype of Cerastis sobrina shuotsuensis was not found in the collection of NHRM, Stockholm (Kononenko, 1996). Genus NAENIA Stephens, 1827 1041. Naenia contaminata (Walker, 1865) � genit.: Pl. 176, fig. 4; ♀ genit.: Pl. 295, fig. 4 Ill. Cat. Noctuidae Korea, 1998: 355; No. 927.

Genus NYSSOCNEMIS Lederer, 1857 1042. Nyssocnemis eversmanni (Lederer, 1853) � genit.: Pl. 176, fig. 5; ♀ genit.: Pl. 295, fig. 5 Ill. Cat. Noctuidae Korea, 1998: 3550; No. 928.

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REFERENCES Beck, H. 1996. Systematische Liste der Noctuidae Europas (Lepidoptera, Noctuidae). Neue Entomologische Nachrichten, 36: 1-122. Beck, H. 1999-2000. Die Larven der Europäischen Noctuidae. Vol. 1-4: Text (1); Illustrations (2-4). Herbipolitana. Buchreihe zur Lepidopterologie. Band 5/1. Check List of Insects from Korea. 1994. Ent. Soc. Korea and Korean Soc. Appl. Ent. Kon-Kuk Univ. Press. Seoul. 1-744 (in Korean). Choi, S.-W. 2006. Two new records of Nolidae (Lepidoptera) from Korea. Korean J. Syst. Zool. 22: 185-188. Edwards, E.D. 1996. Noctuidae. In: E.S. Nielsen, E.D. Edwards and T.V. Runsi [Eds], Check list of the Lepidoptera of Australia. Monogr. Aust. Lepid. 4: 291-333. Fibiger M. 1990. Noctuinae 1. Noctuidae Europaeae. Vol. 1. Entomological Press, Soro, 1990: 1208. Fibiger M. 1993. Noctuinae 2. Noctuidae Europaeae. Vol. 2. Entomological Press, Soro, 1993: 1230. Fibiger, M. 1997. Noctuinae 3. Noctuidae Europaeae. Vol. 3. Entomological Press, Sorø: 1-418. Fibiger, M. and H. Hacker. 1990. Systematic list of the Noctuidae of Europe. Esperiana 2: 1-111. Fibiger, M. and H. Hacker. 1992. Systematic list of the Noctuidae o f Europe. Corrigenda et addenda 1. Esperiana 3: 507-511. Fibiger, M. and H. Hacker. 1998. Systematic List of the Noctuidae of Europe. Corrigenda et Addenda II. Esperiana 6: 9-40. Fibiger, M. and H. Hacker. 2005. Systematic List of the Noctuoidea of Europe (Notodontidae, Nolidae, Arctiidae, Lymantriidae, Erebidae, Micronoctuidae, and Noctuidae). Esperiana 11: 93205. Fibiger, M. and J.D. Lafontaine. 2005. A review of the higher classification of the Noctuoidea (Lepidoptera) with special reference to the Holarctic fauna. Esperiana, 11: 7-92. Franclemont, J.G. and E.L. Todd. 1983. Noctuidae. pp. 120-159. In Hodges, R. W. et al. Check List of the Lepidoptera of America North of Mexico. xxiv+284 pp. London: E. W. Classey Ltd. and the Wedge Entomological Research Foundation. Goater, B., L. Ronkay and M. Fibiger. 2003. Catocalinae, Plusiinae. Noctuidae Europaeae. Vol. 8 Sorø: Entomological Press. 452 pp. Hacker, H. 1990. Revision der Gattung Hadena Schrank, 1802 (Lepidoptera) Theil I. Esperiana 3: 243-361. Hacker, H. 1996. Revision der Gattung Hadena Schrank, 1802 (Lepidoptera: Noctuidae). Esperiana 5: 7-687. Hacker, H. 1998a. Die typen der von E.J. Ch. Esper (1742-1810) in seinem “Die Schmetterlinge in Abbildungen der Natur” beschhreiebenen Noctuoidea (Lepidoptera). Esperiana 6: 433-468. Hacker, H. 1998b. Revision der Gattung Hadula Staudinger, 1889 (= Discestra Hampson, 1905; Aglossestra, Hampson, 1905; = Cardiestra Boursin, 1963), Anartomorpha Alpheraky, 1892, Trichanarta Hampson, 1895, Anarta Ochsenheimer, 1816 und Cardepia Hampson, 1905 mit Beschreibung einer neuen Gattung Hadumorpha gen. n. (Lepidoptera, Noctuidae). Esperiana 6: 577-843. 136

Hacker, H. 2004. Revision of the genus Caradrina Ochsenheimer, 1816, with notes on other genera of the tribus Caradrini (Lepidoptera, Noctuidae). Esperiana, 10: 7-690. Hacker, H., L. Ronkay and M. Hreblay. 2002. Hadeninae I. Noctuidae Europaeae. Vol. 5. Entomological Press, Sorø: 1-419. Hampson, G.F. 1898-1913. Catalogue of the Lepidoptera Phalaenae in the British Museum, 1-13. Han, H.L., B.K. Byun. 2007. Hepatica nakatatanii (Lepidoptera, Noctuidae) new for China. Japan Heterocerists’ J. 242: 307-308. Han, H.L., K.T. Park and L.S. Lu. 2005. Zanclognatha species in Mt. Changbai, with description of a new species and two unknown species from China (Lepidoptera, Noctuidae). Korean J. Syst. Zool. 21(1): 1-10. Han, H.L., V.S. Kononenko and K.T. Park. 2005. Three unknown species of Noctuidae from Korea (Lepidoptera). Korean J. Appl. Entomol. 44(3): 165-168. Han, H.L., J.C. Sohn and G.J. Weon. 2006. Four species of the family Noctuidae (Lepidoptera) new to Korea. Korean J. Appl. Entomol., 45(2): 101-105. Han, H.L., C.D. Li and B.K. Byun. 2007. New record of Trisuloides rotundipennis Sugi (Lepidoptera: Noctuidae: Pantheinae) from Korea. Entomol. Res. 37: 36-38. Hardwick D.F. 1950. Preparation of slide mounts of lepidopterous genitalia. Can. Ent. 82: 231-235. Holloway, J.D. 1998. The classification of the Sarrothripinae, Chloephorinae, Camptolominae and Nolinae as Nolidae (Lepidoptera: Noctuoidea). Quadrifina. 1: 247-276. Holloway, J.D. 2005. The moths of Borneo: family Noctuidae, subfamiliy Catocalinae. Malay. Nat. J. 58(1-4), 1-529 pp. Holloway, J.D. and S.E. Miller. 2003. The composition, generic placement and host-plant relationship of the joviana-group in the Parallelia generic complex (Lepidoptera: Noctuidae, Catocalinae). Invertebrate Systematics, 17: 111-128. Honey, M.R. 1996. Curation of the national collection of Lepidoptera. Presented to poster session at 10th European Congress of Lepidopterology, Miraflores. Unpublished. Inoue, H. 1979b. A new species of the Noctuidae (Lepidoptera) from Japan and Korea. Tinea. 10 (28): 286-289. Ju, J.S. 2005. Additional records on the fauna of lepidopterous insects from Is. Jinbo, Jellanam-do, Korea (II). Lucanus 5: 13-15. Kim, S.S. 1999. Moths collected from Is. Ullung-do, Korea. J. Lepid. Soc. Korea. 11: 19-28. Kim, S.S. and Y.G. Choi. 2000. Some moths taken at caves, South Korea. J. Lepid. Soc. Korea. 13: 49-51. Kishida Y. and H. Yoshimoto. 1988. On Toxocampa lilacina, a senior synonym of Lygephila ogatai. Japan Heterocerist’s J. 150: 395-396. Kitching, I.J. 1984. An historical review of the higher classification of the Noctuidae (Lepidoptera). Bulletin of the British Museum (Natural History), 49: 153-234. Kitching, I.J. and Rawlins, J.E. 1999 [1998]. The Noctuoidea. pp. 355-401. In Kristensen, N. P. (Ed.). Lepidoptera: Moths and Butterflies. Volume 1: Evolution, systematics and biogeography. Handbook of Zoology/Handbuch der Zoologie. 491 pp. Berlin/New York: Walter de Gruyter. Kljuchko, Z.F. 2003. Noctuidae. Subfamily Plusiinae. In: P.A. Lehr (ed). Key to insects of Russian Far East. Vol. V. Trichoptera and Lepidoptera. Pt. 4. Vladivostok. Dal’nauka. P. 187-214. Koçak, A.Ö. 1980. On the nomenclature of some genus- and species-group names of Lepidoptera. Nota Lepid. 2(4): 139-146. Kononenko, V.S. 1990. Synonimic check list of the Noctuidae (Lepidoptera) of the Primorye 137

Territory, the Far East of U. S. S. R. Tinea 13, Suppl. 1: 1-40. Kononenko, V.S. 1996. A revised catalogue of types of the Noctuidae (Lepidoptera) described by F. Bryk (1948) from the Korean Peninsula. Ins. Koreana 13: 1-26. Kononenko, V.S. 1998. A revision of the collection of Noctuidae (Lepidoptera) reported by O. Herz from Korea in 1904. Ins. Koreana. 15: 77-93. Kononenko, V.S. 2003. Noctuidae. Introduction, Subfamilies Euteliinae, Acontiinae, Pantheinae, Acronictinae, Bryophilinae, Agaristinae, Amphypirinae, Cuculliinae, Hadeninae, Noctuinae, Heliothinae. In: P. A. Lehr. (ed). Key to insects of Russian Far East. Vol. V. Trichoptera and Lepidoptera. Pt. 4. Vladivostok. Dal’nauka. p. 11-33, 215-217, 237-602. Kononenko, V.S. 2005. Noctuidae Sibiricae Vol. 1. An annotated check list of the Noctuidae (s. l. ) (Lepidoptera, Noctuoidae: Nolidae, Erebidae, Micronoctuidae, Noctuidae) of the Asian part of Russia and the Ural region. Entomological Press. pp. 1-243. Kononenko, V.S. and A. Prinratana. 2005. Moths of Thailand. Vol. 3. Noctuidae: An illustrated catalogue of the Noctuidae (Insecta, Lepidoptera) in Thailand, Pt. 1: Subfamilies Herminiinae, Rivulinae, Hypeninae, Catocalinae, Aganainae, Euteliinae, Stictopterinae, Plusiinae, Pantheinae, Acronictinae and Agaristinae. Brothers of Saint Gabriel in Thailand. Bangkok. pp. 261. Kononenko, V.S., S.-B. Ahn and L. Ronkay. 1998. Illustrated catalog of Noctuidae in Korea (Lepidoptera). In: K.T. Park (ed. ): Insects of Korea [3]: 1-509. Lödl, M. 1994. Remarks on the classification of the genera Hypena, Dichromia and Harita. Nota Lepid. 16(3/4): 241-250. Lödl, M. 1998. First approach on the identity and taxonomic position of Bleptinodes perumbrosa (Hampson, 1898) (Lepidoptera, Noctuidae: Herminiinae). Quadrifina 1: 245-246. Lödl, M. and B. Mayerl. 1996. Rhinchodontodes Warren, 1913-jüngeres subjectives Synonym von Zekelita Walker, 1863 (Lepidoptera: Noctuidae, Hypeninae). Ent. Z., Essen 107: 236-254. Lödl, M. and B. Mayerl. 1998. Revision der Gattung Zekelita Walker, 1863 (= Rhynchodontodes Warren, 1913) (Lepidoptera: Noctuidae, Hypeninae). Quadrifina 1: 115-230. Lödl, M., S. and Gaal. 1998. Revision of the genus Anoratha (Lepidoptera: Noctuidae: Hypeninae). Quadrifina 1: -5-24. Lafontaine, J.D. 1987. Noctuoidea: Noctuidae (part). In: Dominick, R.B., et al., The Moths of America North of Mexico, fasc. 27. 2: 1-237. Washington: The Wedge Entomological Research Foundation. Lafontaine, J.D. 1998. Noctuoidea Noctuidae (part) Noctuinae (part-Noctuini). In: Dominick, R.B., et al., The Moths of America North of Mexico, fasc. 27. 3: 1-348. Washington: The Wedge Entomological Research Foundation. Lafontaine, J.D. 2004. Noctuoidea: Noctuidae (part), Agrotinae. In: Hodges, R. W. (Ed. ), The Moths of North America, fasc. 27. 1: 1-394. Washington: The Wedge Entomological Research Foundation. Lafontaine J.D. and K. Mikkola 1987. Lock-and-key system in the inner genitalia of Noctuidae (Lepidoptera) as taxonomic character. Ent. Meddr. 55: 161-167. Lafontaine, J.D. and M. Fibiger. 2004. Classification and distribution of the Agrotini. pp. 17-24. In: Lafontaine, J.D. Noctuoidea: Noctuidae (part), Agrotinae. In: Hodges, R.W. (Ed. ), The Moths of North America, fasc. 27. 1: 1-394. Washington: The Wedge Entomological Research Foundation. Lafontaine, J.D. and M. Fibiger. 2006. Revised higher classification of the Noctuoidea. Can. Entomol. 138: 610-635. Leech, J.H. 1889a. On the Lepidoptera of Japan and Korea. Part III. Heterocera, sect. II. Proc. zool. 138

Soc. Lond. 1889: 474-571. Leech, J.H. 1889b. New species of Deltoids and Pyrales from Korea, North China and Japan. Entomologist. 22: 62-71, pls. 2-4. Leech, J.H. 1900. Lepidoptera Heterocera from northern China, Japan and Corea. Part. 3. Trans. ent. Soc. Lond. 1900: 9-161; Part 4. Ibidem. 1900: 511-663. Matthews, M. 1991. Classification of the Heliothinae. Bull. nat. Resour. Inst. 44: i-v, 1-195. Mayerl B. and M. Lödl. 1999. Revision der Gattung Rhynchina Guenée, 1854) (Lepidoptera: Noctuidae, Hypeninae). Quadrifina 2: 1-124. McCabe T.L. 1980. A reclassification of the Polia complex for North America (Lepidoptera: Noctuidae) New York State Mus. Bull. 42: 1-141 p. Mikkola, K. 1993. Lithophane hepatica Clerck, 1759 - a valid combination (Lepidoptera, Noctuidae). Nota lepid. 16(2): 139-144. Mitchell, A., C. Mitter and J.C. Regier. 2000. More taxa or more characters revisited: combining data from nuclear protein-encoding genes for phylogenetic analyses of Noctuoidea (Insecta: Lepidoptera). Systematic Biology, 49: 202-224. Mitchell, A., S. Cho, J.C. Regier, C. Mitter, R.W. Poole, and M. Matthews. 1997. Phylogenetic utility of elongation factor-1” in Noctuoidea (Insecta: Lepidoptera): the limits of synonymous substitution. Molecular Biology and Evolution, 14: 381-390. Mitchell, A., C. Mitter, and J.C. Regier. 2000. More taxa or more characters revisited: combining data from nuclear protein-encoding genes for phylogenetic analyses of Noctuoidea (Insecta: Lepidoptera). Systematic Biology, 49: 202-224. Mitchell, A., C. Mitter, and J.C. Regier. 2006. Systematics and evolution of the cutworm moths (Lepidoptera: Noctuidae): evidence from two protein-coding nuclear genes. Systematic Entomology, 31: 21-46, -. doi: 10. 1111/j. 1365-3113. 2005. 00306. x. Nam, S.H. 1982. Family Noctuidae Stephens, 1829, pp. 282-534, 713-813, pls. 30-71. In: Kim, C. W.S., H. Nam and S.M. Lee. Illustrated Flora and Fauna of Korea. 26. Insecta (8): 1-919, pls. 192 [in Korean with synonimic list in English]. Nowacki, J. and M. Fibiger. 1997. Noctuidae. A distributional Checklist. In: J. Razovsky and M. Fibiger [Eds]. The Lepidoptera of Europe: 250-294, 332-339. Nye, I.W.B. 1975. The generic names of mothds of the world, Volume 1 Noctuoidea (part): Noctuidae, Agaristidae, and Nolidae. 1-568. London: British Museum (Natural History). Oh, S.-H. 1991. Systematics of the family Nolidae (Lepidoptera) in Korea. (I). Thesis for the degree of Doctor of Philosophy. Dept. Agrobiol. Grad. School Kangweon Natnl. Univ. 139 p. (manuscript). Oh, S.-H. 2001. A review of the subfamily Nolinae (Lepidoptera, Noctuidae) in Korea. (I): genus Nola Leech. Ins. Koreana, 18(2): 123-127. Owada, M. 1982. Noctuidae (Herminiinae). In: Inoue H., S. Sugi, H. Kuroko, S. Moriuti, and A. Kawabe. Moths of Japan, 1: 913-935; 2: 405-408, pls. 224-226, 356, 381-392. Kodansha, Tokyo. Owada, M. 1987. A taxonomic study of the subfamily Herminiinae of Japan (Lepidoptera, Noctuidaea). Natn. Sci. Mus. Tokyo: ii+1-208. Owada, M. 1988. On the Lygephila-recta-complex in the collection of Kyushu-University. Japan Heterocerist’s J. 148: 364-365. Owada, M. 1992. Synonymic notes on the herminiine moths (Noctuidae) of Japan, with description of three new species. Tinea. 13(18): 183-203. Owada, M. 2006. Notes on some type specimens of herminiine moths (Lepidoptera, Noctuidae) 139

described by Felix Bryk. Jpn. J. syst. Entom. 121-126. Owada, M., H. Kobayashi, W. Min., C.-M. Fu, H.-R. Tzuoo, C.-S. Lin, M.-L. Chan., T. H Thinh, C. Liu-sheng and H. Yoshimoto. 2006 [2007]. A revision of winter cuculliine moths of the genus Rhynchaglaea (Lepidoptera, Noctuidae) in East and Southeast Asia. Mem. Natn. Sci. Mus., Tokyo. 44: 39-57. Pak, S.W. 1959. A hand list of Korean moths. Forest. Exp. Sta. Korea 1-37. (in Korean). Park, K.T., L. Ronkay, L. Przybylowicz, A. Kun and L. Peregovits. 2001. Moths of North Korea. In: Park, K.T. (eds): Insects of Korea [Series 7] 1-443 pp. Park, K.T., H.L. Han and L. Ronkay. 2006. Catocala of Korea (Lepidoptera, Noctuidae). In Park, K. T. (ed.) Insects of Korea [Series 10], pp. 111. Pogue, M.G. 2002. A world revision of the genus Spodoptera Guenée (Lepidoptera: Noctuidae). Memoirs of the American Entomological Society, 43: 1-202. Poole, R.W. 1989. Noctuidae. Lepidopterorum Catalogues (New Series). Fascicle 118, pt 1: v-xii + 1-500; pt. 2: 501-1013; pt. 3: 1014-1314. E. J. Brill, Leiden. Poole, R.W. 1995. Noctuoidea: Noctuidae (part), Cuculliinae, Stiriinae, Psaphidinae (part). In: Dominick, R. B. et al., The Moths of America North of Mexico, fasc. 26. 1: 1-249. Ronkay, G. and L. Ronkay. 1994. Cuculliinae I. Noctuidae Europaeae. Vol. 6. Entomological Press, Sorø, 1-282. +10 pls. Ronkay, G. and L. Ronkay. 1995. Cuculliinae II. Noctuidae Europaeae. Vol. 7. Entomological Press, Sorø, 1-224 pp. +4 pls. Ronkay, L., J.L. Yela and M. Hreblay. 2001. Hadeninae II. Noctuidae Europaeae. Vol. 4. Entomological Press. Sorø. 452 pp. Sohn, J.C. and L. Ronkay. 2001. New records of Korean Noctuidae (Lepidoptera) with description of a new species. Ins. Koreana. 18(3): 219-227. Sohn, J.C. and S.S. Kim. 2003. Three species of Noctuidae (Lepidoptera) new to Korea. Ins. Koreana. 20(3, 4): 295-300. Sohn, J.C., L. Ronkay and V.S. Kononenko. 2004. Three newly recorded species and four little known species of Korean Noctuidae (Lepidoptera). Entomol. Res. 34(2): 65-72. Sohn, J.C. and H.L. Han. 2005. A new species of Stenoloba Staudinger (Noctuidae, Bryophilinae) from Korea. Tinea 19(1): 22-27. Sohn, J.C., L. Ronkay, S.S. Kim and S.W. Cho. 2005. A taxonomic report of six Noctuidae species (Lepidoptera) new to Korea. Entomol. Res. 35(4): 219-226. Sohn, J.C., L. Ronkay and S.W. Choi. 2005. First report of five Noctuid species (Lepidoptera: Noctuidae) from Korea. J. Asia-Pacific Entomol. 8(2): 147-152. Sohn, J.C., S. and S.W. Cho. 2005. A new record of Ischyja manlia (Cramer) (Lepidoptera: Noctuidae: Catocalinae) from Korea. Entomol. Res., 35(2): 107-110. Speidel, W. and C.M. Naumann. 2005. A survey of family-group names in noctuoid moths (Insecta: Lepidoptera). Systematics and Biodiversity, 2: 191-221. Speidel, W., H. Fänger and C.M. Naumann. 1996. The phylogeny of the Noctuidae (Lepidoptera). Systematic Entomology, 21: 219-251. Sugi, S. and U. Jinbo. 2004. ‘Post-MJ’. Additions of species and changes in names of Japanese moths. Supplement 1. Edn. 2. Japan Heterocerist’s Soc. Tokyo: i-ii +1-60 (in Japanese). Sugi, S. Noctuidae. 2000. ‘Post-MJ’. Additions of species and changes in names of Japanese moths. Edn. 2. Japan Heterocerist’s Soc. Tokyo: I-VII+1-171 (in Japanese). Sugi, S. 1982. Noctuidae (except Herminiinae). In: Inoue H., S. Sugi, H. Kuroko, S. Moriuti, and A. 140

Kawabe. Moths of Japan 1: 669-913, 2: 334-405, pls. 37, 164-223, 229, 278, 355-280. Kodansha, Tokyo (in Japanese with English synopsis). Sugi, S. 1989. Noctuidae. In: Hirashima (ed). The Check List of the Japanese Insects 2: 1042-1088 (in Japanese). Sugi, S. 1992a. Noctuidae (except Aganainae, Herminiinae and Nolinae). Checklist of the Lepidoptera of Taiwan. In: Heppner, J.B. and H. Inoue. [Eds], Lepid. of Taiwan. 1. (2): 171-202. Sugi, S. 1994. ‘Post-MJ’. Additions of species and changes in names of Japanese moths. Edn 1. Japan Heterocerist’s Soc. Tokyo. 1-97 (in Japanese). Sugi, S. 1996. ‘Post-MJ’. Additions of species and changes in names of Japanese moths. Edn. 1 (Suppl. 2). Japan Heterocerist’s Soc. 1-16 (in Japanese). Sviridov, A.V. 1990. On the type material of noctuid species described by F. Bryk from Korea. Novosti faunistiki i sistematiki. Kiev: 97-100 (in Russian). Sviridov, A.V. 1991. To the knowledge on Noctuidae (Lepidoptera) of the Far East from the genera Zanclognatha and Herminia. Zool. zhurn. 70: 55-65 (in Russian with English summary). Sviridov, A.V. 2003. Noctuidae. Subfamilies Herminiinae, Rivulinae, Hypeninae, Catocalinae, Sarrothripinae, Chloephorinae. In: P.A. Lehr (ed). Key to insects of Russian Far East. Vol. V. Trichoptera and Lepidoptera. Pt. 4. Vladivostok. Dal’nauka. P. 35-187, 217-236. Tikhomirov, A.M. 1979. Phylogenetic links in the family Noctuidae (Lepidoptera) on the base of the functional morphology of copulatuve apparatus of males. Proc. Zool. Inst. 89: 75-99. Tikhomirov, A.M. 1989. The system and pylogeny of Palaearctic Noctuidae (Lepidoptera) on the base of the functional morphology of copulatuve apparatus of males. Entomol. obozrenie. 58: 373-386. Tuxen, S.L. 1970. Taxonomist’s glossary of genitalia in insects. (2nd edition 1970). Munksgaard, Copenhagen. 359 p. Ueda, K. 1984. A revision of the genus Deltote R. L. and its allied genera from Japan and Taiwan (Lepidoptera: Noctuidae, Acontiinae). Pt 1. A generic classification of the genus Deltote R. L. and its allied genera. Bull. Kitakyushu nat. Hist. Mus. 5: 91-133. Ueda, K. 1987. A revision of the genus Deltote R. L. and its allied genera from Japan and Taiwan (Lepidoptera: Noctuidae, Acontiinae). Pt 2. Systematics of the genus Deltote R. L. and its allied genera. Bull. Kitakyushu nat. Hist. Mus. 6: 1-117. Weller, S.J., D.P. Pashlev Y, J.A. Martin, and J.L. Constable. 1994. Phylogeny of noctuoid moths and the utility of combining independent nuclear and mitochondrial genes. Systematic Biology, 43: 194-211. Yoshimatsu, S. 1994. A Revision of the Genus Mythimna (Lepidoptera: Noctuidae) from Japan and Taiwan. Bull. Nat. Inst. Agro-Environ. Sci. 11: 81-323. Zilli, A. and W. Hogenes. 2002. An annotated list of the fruit-percing moth genus Eudocima Billberg, 1820 (sensu Poole, 1989) with description of four new species (Lepidoptera: Noctuidae, Catocalinae). Quadrifina 5: 153-207. Zilli, A., L. Ronkay, and M. Fibiger. 2005. Noctuidae Europaeae. Vol. 8. Apameini. 1-323. Sorø: Entomological Press. Zolotarenko, G.S. and V.V. Dubatolov. 1996. A new species og the genus Maliattha Walker, 1863 (Lepidoptera, Noctuidae) from South Primorye (Russian Far East). Atalanta 26(1/2): 299-301.

141

INDEX

A abamita Bremer & Grey 58 abducalis Walker 47 aberrans Eversmann 125 ABROSTOLA Ochsenheimer 75 abrostolina Butler 75 ABROSTOLINI Eichlin & Cunningham 27, 75 absentimacula Guenée 56 ACANTHOLEUCANIA Rungs 128 ACANTHOPLUSIA Duf. 75 accipiter Felder & Rogenhofer 54 ACHAEA Hübner 57 achatina Butler 117 acheron Draudt 92 ACONTIA Ochsenheimer 84 ACONTIINAE 22, 24, 28, 84 ACONTIINI Guenée 28, 84 ACOSMETIA Stephens 25, 97 ACRAPEX Hampson 111 ACRONICTA Ochsenheimer 87 ACRONICTINAE 22, 25, 28, 86 acronyctiformis Guenée 101 acronyctoides Guenée 101 actaea Felder & Rogenhofer 59 ACTEBIA Stephens 26, 129 ACTINOTIA Hübner 105 ACTINOTIINI Beck 28, 105 acuminata Butler 113 adaucta Warren 87 ADISURA Moore 94 adrasta Felder & Rogenhofer 76 adulatricoides Mell 65 adulatrix Hübner 65 adultera Ménétriés 57 adumbrata Eversmann 129 Aedia Hübner 84 AEDIINI Beck 28, 84 AEDINAE 22 aegrota Butler 44 aenescens Moore 79 aerata Butler 107 aerugula Hübner 8, 66 aethiops Butler 97

affinis Linnaeus 117 AFRIDINAE 22 AGANAIDAE 22 AGANAINAE 22 AGARISTINAE 22, 25, 28, 90 agitatrix Graeser 59 agnata Staudinger 75 AGROCHOLA 113 AGROTINA Rambur 28, 129 AGROTINI Rambur 28, 128 AGROTIS Ochsenheimer 130 ain Hochenwarth 79 aino Sugi 118 albicincta Hampson 8, 82 albicolon Hübner 124 albimacula Kononenko 113 albimixta Sugi 99 albipennis Butler 131 albipuncta Leech 113 albipuncta Wileman 65 albiradiosa Eersmann 127 albisignata Oberthür 102 ALBOCOSTA Fibiger & Lafontaine 129 albolineola Graeser 98 albolunata Moore 49 albomaculalis Bremer 48 albonigra Herz 89 albonigra Kononenko 135 albonitens Bremer 118 albonotata Staudinger 96 albopunctalis Leech 46 albosignata Staudinger 32 albostriata Bremer & Grey 75 albovenosa Goeze 89 albula Den. & Schiff 70 albuncula Eversmann 135 aliena Walker 62, 101 alikangiae Strand 108 alni Linnaeus 88 alopecurus Esper 109 alpherakyi Kononenko 108 alpium Osbeck 87 amasina Eversmann 32

amata Bremer & Grey 53 amata Butler 48 amboinensis Felder 63 amethystina Hübner 97 amica Butler 48 amica Treitschke 119 ammonia Cramer 57 amoena Inoue 31 amphidecta Butler 37 AMPHIPOEA Billberg 106 AMPHIPYRA Ochsenheimer 92 AMPHIPYRINAE 22, 28, 92 AMPHITROGIA Hampson 23, 37 amplifascia Warren 60 amurica Staudinger 79 AMYNA Guenée 24, 83 ANACHROSTIS Hampson 23, 31 ANACRONICTA Warren 85 ANADEVIDIA Kostrowicky 76 ANAPAMEA Sugi 110 ANAPLECTOIDES McDunnough 133 ANAPOMA Berio 128 ANARTA Ochsenheimer 26, 122 ANATATHA Hampson 23, 35 anceps Staudinger 38 ancilla Warren 55 ANEDA Sukhareva 125 ANEPIA Hampson 125 angulata Sugi 72 angulata Wileman 49 angulina Guenée 36 angusta Butler 86 angusta Sugi 119 angusta Yoshimoto 105 angustipennis Matsumura 122 annetta Butler 55 ANOCALA Scott 61 ANOMINI Grote 27, 51 ANOMIS Hübner 51 ANORATHA Moore 23, 45 ANORTHOA Berio 121 anpingicola Strand 66 ANTERASTRIA Sugi 24, 25, 100

449

ANTHA Staudinger 119 ANTIVALERIA Sugi 116 ANTOCULEORA Ichinose 77 ANUGA Guenée 65 APAMEA Ochsenheimer 108 APAMEINA 28 APAMEINI Guenée 28, 106 apameoides Draudt 110 aperta Walker 70 apertura Walker 100 apicalis Guenée 63 apicalis Moore 83 APOPESTES Hübner 54 aquila Donzel 108 ARAEOPTERON Hampson 31 ARAEOPTERONINAE 22, 27 ARAEOPTERONINI 31 ARCHANARA Walker 107 arcta Lederer 96 ARCTE Kollar 51 ARCTEINI Berio 27, 51 arctides Staudinger 96 ARCTIIDAE 22 ARCTIINAE 22, 24, 27, 74 arctotaenia Guenée 56 arcuata Moore 56 arcuinna Hübner 31 ardescens Butler 113 arefacta Butler 81 arenacea Kozh 129 arenosa Butler 44, 132 argentalis Moore 68 argentata Butler 73, 81 argentea Butler 73 argentea Hufnagel 90 argillacea Butler 32 ARGYROGRAMMATINI Eichlin & Cunningham 27, 75 argyrosticta Butler 98 ARIOLICA Walker 73 ARIOLICINAE 27 armigera Hübner 94 artemisiae Hufnagel 90 ARTENA Fabricius 55 ARYTRURA John 54 ARYTRURINI Fibiger & Lafontaine 27, 54 asahinai Sugi 108 asiatica Burrows 107 asiatica Krulikowski 72

450

ASIDEMIA Sugi 107 askoldensis Staudinger 121 askoldis Oberthür 110 assimilis Warren 99 assulta Guenée 94 astrigera Butler 60 ASYMBATA Gerstaecker 61 ATACIRA Swinhoe 65 ATHAUMASTA Hampson 100 ATHETINA 28 ATHETIS Hübner 102 atkinsoni Moore 94 atomosa Bremer 67 ATRACHEA Warren 108 atrata Butler 100, 119 atricolor Ahn 80 atriplicis Linnaeus 103 ATUNTSEA Berio 37 Atuntsea Berio 7 Atuntsea kogii 37 AUCHMIS Hübner 105 auragides Draudt 112 aurariae Oberthür 120 aureata Holloway 125 AUSTRANDESIINA 28 AUTOBA Walker 84 AUTOGRAPHA Hübner 78 AUTOPHILA Hübner 54 AUTOPLUSIINA 27 AVENTIINAE 22, 23, 27, 50 AVENTIOLA Staudinger 34 axis Guenée 83 AXYLIA Hübner 130 AXYLIINA 28 azumai Sugi 111 B BAGISARINAE 22, 24, 28, 83 BAGISARINI Crumb 83 baja bajula Staudinger 133 baja chosenbaja Bryk 134 baja cinigera Filipjev 134 baja Den. & Schiff. 133 baja nisseni Rotschild 133 BALSA Walker 86 BALSINAE 22, 25, 28, 86 BAMBUSIPHILA 94 banghaasi sumi Inoue 68 banghaasi West 8, 68 bani Sugi 126

bankiana Fabricius 80 basalipunctata Graeser 106 basalis Moore 46 basimacula Draudt 104 basinigra Kononenko & Ahn 35 basinigra Sugi 111 BASTILLA Swinhoe 24, 56 bathensis Lutzau 119 beatalis Felder & Rogenhofer 47 beatrix Butler 105 BELCIADES Kozh 86 BELCIANA Walker 25, 86 belinda Butler 46 bella Butler 59, 110, 120 bella Staudinger 81 bellula Alphéraky 87 bellula Kononenko & Ronkay 120 BELOSTICTA Butler 93 BERTULA Walker 41 bicolora Leech 84 bicoloralis Graeser 46 bieti Oberthür 77 bifasciata Staudinger 94 bifidalis Leech 8, 38 biguttula Motschulsky 25, 97 bilinea Hampson 122 bimaculata Staudinger 32 bipartita Graeser 135 biplagiata Walker 83 biren Goeze 124 bisacutum Hampson 45 bistriarts Hübner 56 bistriga Möschler 66 bistrigata Staudinger 41 BLASTICORHINUS Butler 63 BLENINA Walker 74 BLENINAE 22 BLENINI 27, 74 BLEPHARITA Hampson 119 blolneata Wileman & South 40 boarmoides Guenée 62 bokhaica Kononenko 60 BOLETOBIINAE 22 bolinoides Guenée 64 bombycina Hufnagel 123 BOMOLOCHA Hübner 46 boursini Sugi 8, 116 BRACHIONYCHA Hübner 92 brassicae Linnaeus 124 bremeri Erschoff 106

bremeri Sugi & Kononenko 118 BREVIPECTEN Hampson 24, 84 BRITHYS Hübner 128 brunnea Den. & Schiff. 131 brunnea Leech 33 brunnescens Kononenko 109 BRYOLEUCA Hampson 25, 99 BRYOMOIA Staudinger 25, 98 bryophasma Boursin 98 bryophilalis Staudinger 69 BRYOPHILINA Staudinger 24, 82 BRYOPHILINAE 22, 25, 28, 98 buraetica Staudinger 79 burrowsi Chapman 107 butleri Leech 37 C caeneusalis Walker 42 calamina Butler 46 CALESIA Guenée 61 calida Rebel 32 calida Staudinger 32 caliginea Butler 85, 132 caliginosa Walker 103 CALLIERGES Hübner 91 CALLISTEGE Hübner 55 CALLOPISTRIA Hübner 97 CALOPHASIA Stephens 91 CALPINAE 22, 27, 51 CALPINI Boisduval 27, 51 CALYMNIA Hübner 117 CALYPTRA Ochsenheimer 51 campana Guenée 55 campicola Lederer 91 CAMPTOLOMA Felder 73 CAMPTOLOMINA 27 CAMPTOLOMINAE 22 camptostigma Ménétriés 118 candida Butler 66 candidalis Staudinger 67 candidula Den. & Schiff. 100 canescens Butler 131 CAPSULA Fibiger, Zilli 107 cara Butler 117 Caradrina 25 CARADRINA Ochsenheimer 101 CARADRININA 28 CARADRININI Boisduval 28, 101 carbonaria Graeser 88 carnipennis Butler 121

casta Butler 52 castanea Moore 7, 8, 96 castaneofasciata Motsch 114 CATOCALA Schrank 57 CATOCALINAE 22, 24, 27, 53 CATOCALINI Boisduval 27, 57 catocaloida Graeser 88 cavernosa Eversmann 124 cecilia Butler 132 cedermarki Bryk 121 CELAMA Walker 66 celsia Linnaeus 106 centonalis Hübner 67 CERASTIS Ochsenheimer 131 cervina Moore 49 chalcogramma Bryk 81 champa Moore 85 CHANDATA Moore 25, 110 changyangis Mayerl & Lödl 45 CHARACOMA Walker 72 characteristica Alpheraky 130 chardinyi Boisduval 132 CHASMINODES Hampson 118 cheiranthi Tauscher 78 CHELONOMORPHA Motschulsky 90 CHERSOTIS Boisduval 132 CHIBIDOKUGA Matsumura 46 chinensis Leech 101 chinensis Wallengren 97 CHLAMIFERA Hübner 66 CHLOEPHORINA 27 CHLOEPHORINAE 22 CHLOEPHORINI Staiton 27, 71 CHLOROTHALPA Beck 103 CHORIZAGROTIS Smith 129 chosenicola Bryk 127 chrysitis Linnaeus 77 CHRYSODEIXIS Hübner 76 chryson Esper 77 CHRYSORITHRUM Butler 53 CHYTONIX Grote 25, 96 CIASA Walker 65 CIDARIPLURA Butler 39 cilia Staudinger 118 cineracea Butler 64 cinerascens Motschulsky 102 cinerea Butler 38, 93 cinerescens Butler 104 circumducta Lederer 122

CIRRHIA Hübner 112 cissigma Ménétriés 132 clara Leech 99 clara Sugi 117 claripennis Butler 48 CLAVIPALPULA Staudinger 120 clavis Hufnagel 130 CLETHROPHORA Hampson 72 c-nigrum Linnaeus 133 cochylioides Guenée 8, 32 COCYTIINAE 22 coelonota Kollar 52 COENOBIA Stephens 107 coenobita Esper 85 coerula Guenée 51 COLLOMELINAE 22 Colobochyla 23 COLOBOCHYLA Hübner 49 COLOCASIA Ochsenheimer 85 COLONSIDERIDIS Beck 124 combusta Hübner 109 commixtalis Zeller 47 commoda Butler 51 comosa Guenée 61 compta Den. & Schiff. 125 compta Moore 128 concerpta Draudt 89 concinnata Heinicke 110 Condica sp. 7, 95 CONDICA Walker 95 CONDICINAE 22, 25, 28, 95 CONDICINI Poole 95 confusa Kononenko & Ronkay 7, 115 confusa Stephens 76 confusa Sugi 111 confusa Wileman 82 confusalis fumosensis Danie 8, l 67 confusalis Herrich-Schäffer 67 confuscata Rebel 32 conigera Den. & Schiff. 126 coniortota Filipjev 121 CONISANIA Hampson 125 CONISTRA Hübner 113 consanguis Leech 84 consocia Borkhausen 114 conspersalis Staudinger 48 conspicua Dyar 68 contaminata Draudt 85 contaminata Walker 135 contigua Den. & Schiff. 124

451

contigua Leech 40 contigua Wileman 8, 61, 62 contracta Butler 116 contrastata Bryk 123 contusa Freyer 117 coreae Inoue & Sugi 77 corealis Leech 34 coreana Leech 56, 72 coreana Matsumura 83 CORGATHA Walker 23, 32 cornelia Staudinger 85 CORORTHOSIA Berio 120 correpta Püngeler 102 corrupta Herz 125 CORSA Walker 23, 34 CORTYTA Walker 60 COSMIA Ochsenheimer 117, 118 COSMIINA 28 costaemacula Staudinger 36 costaestrigalis Stephens 30 costalis Butler 8, 116 costalis Moore 45 costalis Staudinger 69 costalis Sugi 72 costata Moore 65 costimacula Oberth 82 costimacula Staudinger 32 costinotata Butler 36 craccae Den. & Schiff. 53 cramboides Butler 45 CRANIONYCTA de Lattin 89 CRANIOPHORA Snell 89 crassisigna Warren 77 cremata Butler 35 crenata corsa Schawerda 109 crenata Hufnagel 7, 8, 109 crini Fabricius 128 cristatula Hübner 8, 66 cristulalis Staudinger 44 cruentata Moore 7, 8, 94 CRYPHIA Hübner 25, 98 CRYPTOCALA Benjamin 132 CTENOPLUSIA Duf 75 CTENOSTOLA Sugi 107 cucullatella Linnaeus 66 CUCULLIA Schrank 25, 90 CUCULLIINAE 22, 25, 28, 90 culta Butler 81 cuneatoides Poole 110 cuprea Esp 78

452

cupreotincta Sugi 103 cupreoviridis Walker 74 curtalis Walker 36 curvata Leech 126 curvifera Walker 29 curvilinea Han & Park 42 curvilinea Sugi 30 curvilinea Wileman 82 curvipalpis Butler 40 cuspis Hübner 88 cyclica Draudt 95 cyclicoides Draudt 7, 8, 95 CYMATOPHOROPSIS Hampson D DADDALA Walker 24, 60 dahlii Hübner 131 danilovi Bang-Haas 59 DASEOCHAETA Warren 93 DASYCAMPA Guenée 113 DASYPOLIA Guenée 119 dasypterus Kollar 8, 61 datanidia Butler 111 decipiens Butler 71 deflorata Fabricius 61 degenerana Hübner 72 DELTOTE R.L. 24, 80 dentata Kononenko 123 dentata Staudinger 54 deparca Butler 131 deplanata Eversmann 132 deplorata Staudinger 133 deponens Walker 63 depravata Butler 100 derogata Walker 32 descripta Bremer 134 deuteronympha Staudinger 58 dewitzi Graeser 131 DIACHRYSIA Hübner 77 DIANOBIA Behounek 123 DIARSIA Hübner 131 DIATARAXIA Hübner 124 DICHAGYRIS Lederer 26, 129 dictaria Walker 32 DICTYESTRA Sugi 125 diffusa Sugi 38 digna Butler 88 dilatata Butler 134 DILOBINAE 22 DIMONA Walker 66

86

DIMORPHICOSMIA Sugi 118 DINUMMA Walker 63 DIOMEA Walker 23, 35 DIOSZEGHYANA Hreblay 120 DIPHTHERINAE 22 DIPTERIGIA Stephens 103 DIPTERYGINA Sugi 103 DISCESTRA Hampson 122 discisigna Sugi 35 disjungens Walker 55 disruptalis Walker 36 dissecta Walker 125 dissimilis Bremer 59 dissimilis Hampson 102 distincta Leech 72 distinguenda Staudinger 79 ditrapezium Den. & Schiff. 133 divergens Butler 113,126 divisa Walker 33 doerriesi Staudinger 58 dolosa Butler 44 dotata Fabricius 55 DRYOBOTODES Warren 119 dubia Butler 34, 63 dula Bremer 58 dulcis Butler 56 duplicans Walker 97 duplicata Butler 59 durandi Lucas 133 DYPTERYGIINI Forbes 28, 103 DYSALETIA Sugi 128 DYSEDIA Rogenhofer 64 Dysgonia 24 DYSGONIA Hübner 56 DYSMILICHIA Speis 25, 96 E EARIADINAE 22 EARIADINI Hampson 27, 73 EARIAS Hübner 73 ebatoi Inoue 8, 67 ECTOGONIA Hampson 23, 37 edentata Leech 113 EDESSENA Walker 39 efflorescens Butler 134 egregia Staudinger 85 ELAPHRIA Hübner 101 ELAPHRIINAE Beck 101 ELAPHRIINI 28 electa Vieveg 58

electaria Bremer 55 ELIGMA Hübner 74 ELIGMINAE 22 ELIGMINI 27, 74 ella Butler 59, 120 elongata Butler 91 emarginata Fabricius 52 eminens Staudinger 59 eminentissima Bryk 71 EMMELIA Hübner 84 emortualis Den. & Schiff. 34 ENARGIA Hübner 117 ENISPA Walker 23, 32 ephesperis Hübner 50 EPIZEUCTIS Meyrick 66 erastroides Draudt 80 ERASTROIDES Hampson 82 ERCHEIA Walker 62 ERCHEINI Berio 27, 62 EREBIDAE 22 Erebidae 23 erebina Butler 49, 92 EREBINAE Leech 22, 27, 50 EREBINI Leech 27, 50 EREBUS Latreille 50 EREMODRINA Boursin 101 ERICEIA Walker 62 ERIOPINAE 22, 28, 97 eriosoma Doubleday 76 ERYGIA Guenée 24, 63 ERYTHROPLUSIA Ichinose 76 ERYTHROTIS Bryk 121 esbiahni Sohn & Ronkay 8, 34 esmeralda Oberthür 78 EUBLEMMA Hübner 23, 32 EUBLEMMINAE 22, 23, 27, 31 EUBLEMMINI Forbes 27, 31 EUCARTA Lederer 96 EUCHACIINA 28 EUCHALCIA Hübner 77 EUCHORISTEA Warren 121 EUCLIDIA Ochsenheimer 54 EUCLIDINI Guenée 27, 54 EUCOCYTIINAE 22 EUDOCIMA Billberg 52 EUGRAPHE Hübner 135 EULOCASTRA Butler 24, 82 EUPLEXIA Stephens 105 EUPLEXIDIA Hampson 105 EUPSILIA Hübner 115

EUROIS Hübner 133 EUROMOIA Staudinger 87 euryptera Boursin 102 eurytaenia Prout 62 EUSTROTIINAE 22, 24, 28, 79 EUTELIA Hübner 65 EUTELIINAE 22, 27, 65 EUWILEMANIA 23, 49 EUXOA Hübner 129 evanida Butler 120 evelina Butler 113 eversmanni Lederer 135 Evia ferrinalis Walker 65 EVIA Walker 65 EVONIMA Walker 70 excavata Butler 52 excelsa Kretschmar 79 excisa Swinhoe 82 exclamationis Linnaeus 130 exigua Hübner 100 exprimens Walker 94 exsoleta Butler 115 extensa Butler 93 extincta Kononenko 122 extrema Walker 73 exusta Butler 71, 135 F fagana Fabricius 72 fallax Butler 30 falonia Linnaeus 52 falsa Butler 80 fani Staudinger 119 fascialis Leech 40 fasciata Butler 96 fasciata Kononenko 112 fasciata Moore 89 fasciata Sugi 29 fasciata Walker 66 fausta Leech 8, 121 FELINIOPSIS Roepke 108 fennica Tauscher 129 fentoni Butler 40, 82 FERALIA Grote 93 FERALIINI Poole 28, 93 ferreilutea Hampson 72 fervens Butler 71, 94 festucae Linnaeus 79 figurata Moore 49 filipjevi Kononenko 113

filium Guenée 101 flaccida Inoue 7, 31 flammea Curtis 128 flava Fabricius 51 flavicollis Leech 97 flaviplaga Warren 37 flavipuncta Leech 83 flavomacula Oberthür 38 flavomacula Staudinger 36 flavomaculata Bremer 53 flavostigma Bremer 127 fletcheri Berio 52 fletcheri Sugi 113 flexula Den. & Schiff. 50 FODINA Guenée 61 fodinae Oberthür 7, 110 fontinalis Ronkay & Kononenko formosa Butler 115 formosa Graeser 119 formosenis Strand 62 fortis Butler 106 fractalis Guenée 44 fragariae Vieweg 115 fragilis Swinhoe 66 fragmenta Inoue 7, 31 fraterna Butler 91 fraudatrix Eversmann 90 fraxini Linnaeus 57 friedeli Pinker 129 fucosa Freyer 106 fukudai Sugi 96 fuliginosa Leech 95 fulminea Scopoli 58 fumidalis Zeller 47 fumosa Butler 8, 42, 60, 69 funesta Leech 93 funesta Staudinger 102 furva Vileman 47 furvula Hübner 102 fusca Leech 101 fusca Wileman 62 fuschiana Eversmann 90 fuscipennis Sugi 116 fuscofasciata Inoue 69 fuscostigma Bremer 134 G GABALA Walker 73 gamma Linnaeus 78 gastropachoides Guenée

99

61

453

GELASTOCERA Butler 71 gemella Leech 96 gensanalis Leech 34 gentiusalis Walker 39 geometrica Fabricius 8, 57 GERBATHODES Warren 86 geyeri Felder & Rogenhofer 65 gigantoides Inoue 8, 69 gigas Butler 8, 70 GIRPA Walker 62 gladiata Butler 39 glauca Kononenko & Ronkay 8, 99 glauca kuytekparki 7 glaucopera Hampson 83 glaucotorna Hampson 33 GLOTTULINAE 22 GLOTTULINI Guenée 28, 128 gluteosa Treitschke 102 goliath Oberthür 123 GONEPATICA Sugi 23, 38 gonioptila Prout 62 GORTYNA Ochsenheimer 106 grabczewskii Püngeler 65 GRAMMODES Guenée 57 grandis Butler 126 granitalis Butler 99 grata Butler 119 grisea Leech 60 grisealis Den. & Schiff. 44 griselda Butler 42 griseola Nagano 98 griseola Staudinger 36 griseolineata Warren 51 grisescens Draudt 113 gruesa Draudt 52 GYNAEPHILA Staudinger 23, 40 GYROSPILARA Kon 119 H HADENA Schrank 26, 125 HADENINAE Guenée 22, 25, 28, 120 HADENINI Guenée 28, 122 HADENNIA Moore 39 HADUALA Staudinger 122 hallasana Ronkay 71 hamada Felder & Rogenhofer 39 hampsoni Sugi 109 hampsoni Wileman 71 hamulatrix Draudt 65

454

haplochrosta Hulstaert 47 HARITA Moore 46 HARUTAEOGRAPHA Yoshimoto 122 hebetata Butler 76 hedemanni Staudinger 94 helena Eversmann 58 helicina Hübner 50 HELICOVERPA Hardw 94 HELIOCHEILUS Grote 94 HELIOTHINAE 22, 25, 28, 93 HELIOTHIS Ochsenheimer 93 HELOTROPHA Lederer 106 helva Butler 42 HEMIGLAEA Sugi 116 HEMIPSECTRA Hampson 30 hepatica Clerck 23, 114, 123 HEPATICA Staudinger 38 heptarchia Boursin 101 HERAEMA Staudinger 7, 25, 103 HERCHUNDA Swinhoe 64 hercules Butler 60 hercules Felder & Rogenhofer 88 HERMINIA Latreille 44 HERMINIINAE 22, 23, 27, 39 HERMONASSA Walker 132 heterogyna O. Bang-Haas 122 hidakaensis Sugi 116 Himalistra Hacker 113 HIPOEPA Walker 44 hirticornus Haworth 109 hirtissima Walker 64 hoenei Berio 63 hoenei Püngeler 122 hokkaida Wileman 51 HOLOCRYPTIS Lucas 23, 39 honeyi Yoshimoto 125 HOPLODRINA Boursin 102 hostilis Boursin 8, 91 HULODINI Guenée 27, 62 humidalis Doubleday 30 humilis Boursin 91 HYALOBOLE Warren 113 HYBOMA Hübner 87 hybrida Ronkay 76 HYDRAECIA Guenée 106 HYDRILLODES Guenée 40 HYLONYCTA Sugi 88 HYPENA Schrank 46 Hypena sp. 1 48

HYPENINAE 22, 23, 27, 45 HYPENODES Doubleday 23, 30 hypenodes Inoue 46 HYPENODINAE 22, 23, 27, 30 HYPENOMORPHA Sugi 46 HYPERSTROTIA Hampson 24, 83 HYPERSYPNOIDES Berio 24, 60 HYPHILARE Hübner 127 HYPOCALA Guenée 61 HYPOCALINI Guenée 27, 61 HYPOPYRA Guenée 50 HYPOPYRINI Guenée 27, 50 HYPORBARATHRA Hampson 124 HYPOSADA Hampson 23, 33 HYPOSEMANSIS Hampson 24, 65 HYPOSPILA Guenée 24, 64 HYPOSTROTIA Hampson 23, 38 HYPPA Duponchel 105 HYSSIA Guenée 124 I IAMBIA Walker 25, 97 ichinosei Dufay 76 icterias Eversmann 124 icteritia Hufnagel 112 IDIA Hübner 40 idiostygia Sugi 80 illecta Walker 95 illoba Butler 125 illustrata Graeser 105 illustrata Staudinger 95 IMOSCA Sugi 24, 83 imparata Walker 49 impura Hübner 126 INABAIA Sugi 81 inamoena Hübner 31 inanis Oberthür 128 incerta Hufnagel 120 incerta Roepke 108 incerta Staudinger 110 inchoata Wileman 35 incommoda Staudinger 124 incongruens Butler 39 inconspicua Butler 29, 54 inconspicua Draudt 117 indica Moore 54 indicatalis Walker 48 indistans Guenée 8, 108 inexpecta Sugi 107 inexpectata Alph 129

inexpectata Ueda 79 infecta Walker 101 inferens Walker 111 innocens Butler 44 innocua Butler 8, 66, 68 innocua japonibia Strand 67 insignata Walker 101 insulicola Sugi 108 interiorata Walker 73 intermedia Warren 88 intermediata Bremer 105 intermissa Butler 119 intermixta Warren 75 interrogationis Linnaeus 79 intrahens Walker 8, 64 invaria Walker 61 involuta Walker 51 iodochra Sugi 127,128 iopolia Fletcher 62 IPIMORPHA Hübner 117 IPIMORPHINI Beck 117 ipsilon Hufnagel 130 IRAGAODES Matsumura 71 ISCADIA Hampson 74 ISCHYIA Agassiz 63 ISCHYJA Hübner 63 ITMAHARELA Nye 23, 46 J janata Linnaeus 57 jankowskii Oberthür 35, 89, 90, 92, 99, 108, 118 japana Motschulsky 90 japonago Wileman & West 112 japonibia Inoue & Sugi 8, 98 japonibia Strand 67 japonica Butler 44 japonica Draudt 120 japonica Höne 115 japonica Sugi 97 japonica Warren 82 jessica Butler 77 jezoensis Sugi 104 JOCHEAERA 88 JODIA Hübner 114 jonasii Butler 58 jozana Matsumura 87 juno Dalman 55 jussalis Walker 47 juvenilis Bremer 7, 54

LEIOSTOLA Hampson 23, 38 lentalis Guenée 8, 41 K leodura Staudinger 86 leonina Oberthür 77 KARANA Moore 25, 104 lepigone Möschler 102 karschi Graeser 130 LEUCANIA Ochsenheimer 128 karsholti Fibiger 29 LEUCANINI Guenée 28, 125 kawadai Sugi 110 leucanoides Walker 29 kengkalis Bremer 47 LEUCAPAMEA Sugi 110 KERALA Moore 71 khasanica Zolotarenko & Dubatolov leucocuspis Butler 88 leucographa Den. & Schiff. 131 8, 81 leucolopha Prout 51 klapperichi Kononenko 7, 111 LEUCOMELAS Hampson 7, 54 kogii Inoue 7, 30 leucomelas Linnaeus 84 kogii Sugi 37 leuconephra Hampson 110 kollari Lederer 133 leucostigma Hübner 106 kolthoffi Bryk 87 leucostigma Kollar 61 kononenkoi Poole 93 libatrix Linnaeus 52 korbae Püngeler 90 lidia auct 129 koreaeplexia Bryk 105 lignea Butler 35 koreana Draudt 92 ligustri Den. & Schiff. 89 koreana Ronkay & Kononenko 100 lilacina Butler 7, 42, 54, 121 koreana Staudinger 59 lineatipes Moore 126 koreognatha Bryk 42, 43 lineosa Moore 103 kosemponis Strand 62 liparisalis Walker 66 KOYAGA Ueda 80 LIRA Billberg 66 kulmburgi Rebel 89 literosa Haworth 110 kumamotonis Matsumura 116 LITHOMOIA Hübner 115 kumaso Sugi 108 LITHOPHANE Hübner 114 kunsuki Ahn 33 LITOLIGIA Beck 110 kurilullia Bryk 91 litura Fabricius 101 L livida Den. & Schiff. 92 lizetta Butler 120 LACANOBIA Billberg 123 locuples Oberthür 77 LACERA Guenée 57 longicosta Oh 8, 67 lactaria Graeser 71 longipennis Sugi 51 laetevirens Oberthür 104 longiplaga Warren 105 lagusalis Walker 38 longtpennis Moore 65 lamda Fabricius 114 Lophomilia 23 LAMPROTES R.L. 78 Lophomilia sp. 7, 37 lapidea Wileman 102 LOPHOMILIA Warren 37 lara Bremer 57 LOPHONYCTINAE 22 LASPEYRIA Tutt 23, 50 LOPHOPTERA Guenée 65 lata Butler 51 LOPHORUZA Hampson 23, 33 LATEROLIGIA Zilli 109 loreyi Duponchel 128 latifascialis Walker 66 lucens Butler 132 LATIROSTRUM Hampson 45 lucens Freyer 107 lbibasis Draudt 116 lucilla Butler 60, 104 LEBENA Walker 66 lucipara Linnaeus 105 LECASIA Fawcett 61 ludicra Hübner 53 leichardtii Koch 62 juventina Stoll

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ludifica Linnaeus 85 lui Han & Park 8, 42 lunalis Scopoli 42 lunula Hufnagel 91 lunulata Sterz 36 lutea Bremer & Grey 88 lutefascialis Leech 32 lutosa Hübner 107 lutulenta Esper 109 LYCOPHOTIA Hübner 132 LYGEPHILA Billberg 53 LYMANTRIIDAE 22 LYMANTRIINAE 22, 24, 27, 75 M MACDUNNOUGHIA Kostrowicky 76 MACROCHTHONIA Butler 71 macrops Linnaeus 50 maculana Ahn 80 maculata Kononenko 119 maculata Staudinger 70 maculifera Staudinger 40 maculosa Staudinger 91 magnumisma Ahn 80 MAGUDA Walker 23, 35 major Bremer 87 major Dufay 75 major Hampson 69 malachitis Oberthür 86 malchani Draudt 123 malgassica Heyden 49 MALIATTHA Walker 81 MAMESTRA Ochsenheimer 124 mandarina Freyer 79 mandschuriae Oberthür 91 mandschuriana Oberthür 8, 70, 71 mandschuriana Staudinger 56 mandschurica Bremer 7, 103 mandshurica Boursin 119 manleyi Leech 99 manlia Cramer 8, 63 marginata Moore 94 marginata Walker 61 maritima Graslin 93 marmorata Staudinger 36 marumoi Sugi 33 MATAEOMERA Butler 23, 34 maturata Walker 56 maurella Staudinger 102

456

mauritia Boisduval 101 maxima Berio 51 maxima Bremer 53 MECODINA Guenée 24, 64 MECYRA Walker 61 mediofascia Inoue 7, 8, 69 mediofusca Sugi 98 MEGANEPHRIA Hübner 93 MEGANOLA Dyar 66, 68 MEGASEMA Hübner 133 melachlora Staudinger 98 MELANCHRA Hübner 124 melanica Sugi 46 melanocosta Inoue 69 melanodonta Hampson 120 melanospila Kollar 103 MELAPIA Sugi 55 MESAPAMEA Heineman 110 mesogona Walker 51 mesozona Lucas 66 METOPONIINAE 22, 28, 89 METOPTA Swinhoe 50 mi Clerck 55 MICARDIA Oberthür MICREREMITES Warren 23, 30 MICRONOCTUA Fibiger 23, 29 Micronoctua sp. 7 MICRONOCTUIDAE 22, 23, 27, 29 microphasma Butler 8, 68 MICRORTHOSIA 120 Microxyla 24 MICROXYLA Sugi 82 mikadina Butler 78 MIMACHROSTIA Sugi 23, 29 MIMERASTRIA Butler 70 MIMEUSEMIA Butler 90 minima Ahn 80 minna Butler 47 MINNAGARA Walker 66 minuali Leech 66 minutissima Draudt 99 minutissima Sugi 31 mira Butler 33 mirabilis Bryk 53 mirabilis Sugi 120 miracula Herz 98 misae Sugi 35 mista Hübner 131 mitsuhashi Marumo 98

mixta Staudinger 87 MNIOTYPE Franclemont 119 MOCIS Hübner 55 moderata Staudinger 118 modesta Hübner 78 modestoides Poole 78 moellendorfi Herz 53 moestalis Walker 49 mollicula Graeser 82 mollis Butler 38 MOLYBDONYCTA Sugi 87 MOMA Hübner 87 monolitha Guenée 92 montana Bremer 102 monticola Sugi 126 MORMO Ochsenheimer 25, 104 morosa Butler 40 morosa Lederer 101 morosa permorosa Bryk 101 MORPHOPOLIANA Hreblay & Legrran 127 mortua Staudinger 123 MOSOPIA Walker 39 mucida Esper 131 multiplicans Walker 65 multisignata Christoph 34 munda Den. & Schiff. 121 mus Oberthür 85 muscivirens Butler 104 musculus Ménétriés 54 musmi Hampson 58 MYTHIMNA Ochsenheimer 126 MYTHIMNINI Rungs 125 N NACNA Nye 86 nadeja Oberthür 77 NAENIA Stephens 135 nagaii Sugi 115 NAGANOELLA Sugi 23, 37 nagioides Wileman 59 nakatanii Sugi 8, 38 nami Inoue 8, 67 NARANGA Moore 23, 24, 79 narcissus Cramer 74 NARCOTICA Sugi 86 nebulosa Hufnagel 123 nebulosa Inoue 31 NEGRITOTHRIPA Inoue 71 nemorum Oberthür 81

nervosa Den. & Schiff. 89 neurica Hübner 107 NEUSTROTIA Sugi 24, 82 ni Hübner 75 NIABOMA Nye 125 nigricostata Graeser 54 nigrilinea Leech 118, 127 nigripuncta Wileman 44 nigripunctalis Wileman 31 nigrisigna Leech 44 nigrisigna Walker 78 nigrobasalis Herz 46 nigromaculata Höne 8, 121 nigromaculata Nagano 70 nihonica Höne 8, 115 NIKARA Moore 25, 96 niphona Butler 41 niphonica Kish. & Yoshim 104 NIPHONYX Sugi 25, 96 niphostena Hampson 66 nipona Felder & Rogenhofer 132 nitens Butler 32, 108 nivea Butler 57 niveola Motschulsky 86 niveosparsa Matsumura 86 niveostrigata Warren 63 nobilis Staudinger 71 Noctua algae Esper 107 Noctua tricomma ESPER 133 NOCTUIDAE 22, 27, 29 NOCTUINA 28 NOCTUINAE 22, 26, 28, 128 NOCTUINI Latreille 28, 130 NOCTUOIDEA 29 NODARIA Guenée 39, 41 NOLA Leach 66 NOLATHRIPA Inoue 71 NOLIDAE 22, 24 NOLINAE 22, 27, 66 NOLINI Bruand 27, 66 noloides Butler 82 NONAGRIINI Guenée 106 NOTODONTIDAE 27 novoobscurior Bryk 130 nubeculosa Esper 92 nubes Guenée 101 nubiferalis Leech 64 nubila Butler 59 numisma Staudinger 80 nupta Linnaeus 57

Nycteola costalis 7 NYCTEOLA Hübner 72 NYCTYCIA Hampson 116 nymphula Rebel 39 NYSSOCNEMIS Lederer 135 O oberthueri Staudinger 103 obliqua Wileman 8, 39 obliquaria Marumo 33 obliquiorbis Moore 108 obliterata Rambur 100 obscena Alphéraky 58 obscura Bremer & Grey 56 obscurata Butler 36 obsoleta Hübner 128 obsoleta Marumo 33 occata Moore 47 occidua Prout 62 occulta Linnaeus 133 oceanica Kardakoff 135 ocellaris Borkhausen 113 ochreata Walker 76 ochreipennis Sugi 38 ochrogaster Guenée 129 ochroleucana Staudinger 71 OCHROPLEURA Hübner 130 octogenaria Goeze 102 oculata Draudt 99 oda deLattin 89 ODICE Hübner 31 odiosa Butler 121 ogasawarensis Matsumura 101 ogatai Kinoshita & Sasaki 54 OGLASA Walker 23, 38 ohtanensis Matsumura 133 okanoi Inoue 8, 68 oleagina Denis & Schiffermüller 92 oleracea Linnaeus 124 OLIGIA Hübner 110 OLIGONYX Sugi 25, 96 olista Swinhoe 55 olivacea Hampson 84 OLIVENEBULA Kishida & Yoshimoto 25, 103 Olulis Walker 37 OMOPTERINI Boisduval 55 ONCOCNEMIDINAE 22, 28, 91 ONCOCNEMIS Lederer 92 ononis Den. & Schiff. 93

opalina Butler 38 ophiogramma Esper 109 OPHIUSA Ochsenheimer 55 OPHIUSINI Guenée 27, 55 ORAESIA Guenée 52 orbiculosa Esper 106 ORBONA Hübner 115 orientalis Sugi 107 orientalis Warren 57 oriolus Guenée 61 ornata Bremer 78 ORTHOGONIA Felder & Felder 25, 104 orthogramma Boursin 99 ORTHOSIA Ochsenheimer 120 Orthosia satoi Sugi 121 ORTHOSIINI Guenée 28, 120 ORUZA Walker 23, 33 othello Fawcett 61 owadai Kononenko 7, 112 OXYODES Guenée 48 OXYTRIPIA Staudinger 106 OXYTRIPINA 20 OZARBA Walker 24, 83 P pabulatricula Brahm 107 PABULATRIX Sugi 107 PACHNOBIA Guenée 134 pacifica Boursin 131 pacifica Dufay 75 pacifica Filipjev 89 pacifica Kononenko 115 pacifica Sugi 110, 114 pacificus Owada 41 paki Kishida 58 paleacea Esper 117 pallens Linnaeus 126 pallescens Butler 131 pallidipennis Sugi 102 palliola Denis & Schiffermüller 66 palshkovi Filipjev 86 PANCHRYSIA Hübner 78 PANDESMINI Wiltshire 27, 62 PANGRAPTA Hübner 23, 35 PANGRAPTINI Grote 27, 35 PANOLIS Hübner 120 PANTHAUMA Staudinger 85 PANTHEA Hübner 85 PANTHEINAE 22, 25, 28, 85

457

PAPESTRA Sukhareva 124 PARACOLAX Hübner 40 PARACRAMA Moore 72 PARADIARSIA McDunnough 132 PARADRINA Boursin 101 PARAGABARA Hampson 23, 38 Paragona sp. 7 PARAGONA Staudinger 34 Parallelia 24 PARALLELIA Hübner 56 PARASTICHTIS Hübner 112 paritalis Walker 8, 45 parki Ronkay & Kononenko 93 paromoea Hampson 120 parvula Leech 36 PASIPEDA Walker 61 pastinum Treitschke 53 patala Felder & Rogenhofer 58 paupera Staudinger 86 PECHIPOGO Hübner 43 pecten Guenée 100 peponis Fabricius 76 perforate Bremer 91 perfractalis Bryk 43 PERIDROMA Hübner 128 PERIGRAPHA Lederer 122 PERINAENIA Butler 54 permuda Walker 101 persicariae Linnaeus 124 persimilis Butler 90 pertendens Walker 35, 62 petasitis Doubleday 106 petrina Butler 34 peustera Püngeler 85 pfennigschmidti Püngeler 121 PHLOGOPHORA Treitschke 105 PHLOGOPHORINI Hampson 28, 105 phoenissa Lederer 32 PHYLLOPHILA Guenée 100 PHYTOMETRA Haw 23, 48 PHYTOMETRINAE 22, 23, 27, 48 picta Butler 60 pilicornis Brahm 131 pirata Herz 59 placida Butler 126 placodoides Guenée 98 plagiola Hampson 68 PLATAPLECTA 87 PLATYPERIGEA Smith 102

458

plecta Linnaeus 130 plumbealis Matsumura 114 PLUSIA Ochsenheimer 79 PLUSIDIA Butler 78 PLUSIINA 28 PLUSIINAE 22, 24, 27, 75 PLUSIINI Boisduval 27, 76 Plusilia 25 PLUSILLA Staudinger 97 PLUSIODONTA Guenée 52 plusioides Butler 45 PLUSIOPALPA Holland 76 POECILONOLA Hampson 68 POLIA Ochsenheimer 123 Polia tincta Brahm 114 POLIOTHRIPA Hampson 66 polybapta Butler 37 POLYCHRYSIA Hübner 78 POLYDESMA Boisduval 62 POLYMIXIS Hübner 119 polyodon Clerck 105 POLYPHAENIS Boisduval 104 POLYPOGON Schrank 43 porrigens Walker 66 postalba Sugi 124 postflava Leech 104 postica Hampson 128 POTAMORPHA Guenée 63 praeclara Graeser 89 praecox Linnaeus 129 praecurrens Staudinger 129 praegnax Walker 59 prasina Den. & Schiff. 133 PREMUSIA Walker 24, 64 privata Walker 51 proboscidalis Linnaeus 47 procellosa Butler 57 PRODENIINI Forbes 28, 100 PROLOPHOTA Hampson 7, 34 PROMETOPUS Guenée 25, 97 prominens Moore 74 PROSPALTA Walker 25, 95 PROTODELTOTE Ueda 79 protogigas Inoue 70 PROTOLAMPRA McDunnough 135 PROTOMISELIA Sugi 122 PROTOSCHINIA Hardw 25, 94 proxeneta Alphéraky 59 proxima Leech 127 pruinosa Butler 114

pruinosa Guenée 87 pryeri Butler 40 pryeri Leech 119 PSAPHIDINAE 22, 28, 92 PSAPHIDINI Grote 28, 92 pseudalbonitens Sugi 118 PSEUDALETIA Franclemont 126 PSEUDEUSTROTIA Warren 25, 100 PSEUDEUSTROTIINI Beck 28, 100 PSEUDEUSTROTINAE 22 PSEUDOCOSMIA Kon 119 PSEUDOIPS Hübner 72 PSEUDOPANOLIS Inaba 122 pseudoturpis Kononenko & Ahn 111 pudicana Staudinger 73 pudorina Den. & Schiff. 126 puella Leech 58 pulchella Leech 8, 68 pulcherrima Butler 33 pulchra Butler 74, 81 pulverosa Hampson 87 punctalis Herz 44 punctata Aurivillius 133 punctigera Walker 83 punctum Fabricius 83 punicea Hübner 132 punkikonis Matsumura 103 PURIPLUSIA Chou & Lu 77 purissima Butler 77 pusilla Butler 34 pustulifera Walker 65 putnami Grote 79 putris Linnaeus 130 pygarga Hufnagel 79 pygmaea Wileman 32 PYGOPTERYX Staudinger 25, 116 pyralina Den. & Schiff. 117 pyraloides Sugi 30 pyramidea Linnaeus 92 pyropia Butler 76 PYRRHIA Hübner 94 PYRRHIDIVALVA Sugi 25, 96 Q quadra Graeser 40 quadrilinea Leech 115

R radiata Bremer 127 ramosula Staudinger 91 Raparna 23 RAPARNA Moore 49 raphael Oberthür 88 RAPHIA Hübner 85 RAPHIINAE 22, 25, 28, 85 ravida Den. & Schiff. 132 recta Bremer 53 rectalis Eversmann 41 rectifascia Sugi 7, 30 rectifasciata Ménétriés 50 rectilinea Esper 105 rectilineata Ueda 82 rectivittalis Moore 47 redunca Swinhoe 56 remota Hreblay & Ronkay 115 repetita Butler 111 repleta Walker 97 repugnalis Walker 41 restituta Walker 117 retorta Clerck 50 retusa Linnaeus 117 RHESALA Walker 49 RHIZEDRA Warren 107 RHYNCHAGLAEA Hampson 116 RHYNCHINA Guenée 23, 45 Rhynchodontodes sp. 45 RHYNCHOPALPUS Hampson 68 RISOBA Moore 74 RISOBINAE 22 RISOBINI 27, 74 RIVULA Guenée 29 rivularis Walker 98 rivuligera Butler 46 Rivuliinae 23 RIVULINAE 22, 27, 29 robiginosa Staudinger 44 ROESELIA Hübner 66 rosacea Leech 81 rosalia Staudinger 97 roseata Wileman & South 49 roseifera Butler 73 roseifera Sugi 74 roseiventris Gerstaecker 61 roseoviridis Sugi 7, 73 rosinae Püngeler 114 rosita Guenée 32 rostrata Fabricius 61

rotundipennis Owada 45 rotundipennis Sugi 85 rubricosa Den. & Schiff. 7, 131 rubrirena Treitschke 109 ruficauda Warren 131 ruficeps Walker 50 ruficirra Hampson 72 rufipalpis Walker 61 rufipennis Butler 126 rumicis Linnaeus 88 rurea Fabricius 109 ruta Eversmann 130 rutilifrons Walker 76 S SABLIA Sukhareva 127 sachalinensis Matsumura 134 sachalinensis rikovskensis Matsumura 134 SADARSA Moore 65 saga Butler 105 sagitta Fabricius 48 sagittata Butler 45 salaminia Cramer 52 salebrosa Butler 127 salicalis Den. & Schiff. 49 sanguinea Sugi 118 sanguinolenta Moore 72 SAPPORIA Sugi 111 SARBANISSA Walker 90 SARCOPOLIA Sugi 125 SARROTHRIPINA 27 SARROTHRIPINAE 22 SASUNAGA Moore 105 satoi Inoue 8, 69 satura Den. & Schiff. 119 sauberi Graeser 93 saucia Hübner 128 SCEDOPLA Butler 23, 38 SCHRANKIA Hübner 23, 30 schrenckii Ménétriés 92 scitula Butler 8, 116 SCLEROGENIA Ichinose 77 SCOLECOCAMPINAE 22 SCOLIOPTERIGINI Spuler 52 SCOLIOPTERYGINI 27 SCOLIOPTERYX Germ 52 scolopacina Esper 109 scopariae Dorfmeister 90 scotacra Filipjev 130

scrobiculata Fabricius 57 scutosa Goeze 94 secunda Remm 38 segetum Den. & Schiff. 130 segregata Butler 96 seifersi RANGNOW 127 SELCA Walker 66 semiherbida Walker 134 SEMIOPHORA Stephens 121 senex Butler 74 SENTA Stephens 128 separans Leech 58 separata Walker 126 separatalis Herz 30 sera Felder & Felder 104 sergia Oberthür 77 sericea Butler 85, 114 sericealis Scopoli 29 SERRODES Guenée 55 SESAMIA Guenée 111 SESAMIINA 28 shimekii Inoue 8, 70 sibirica Boisduval 129 SIDEMIA Staudinger 106 SIDERIDIS Hübner 124 sigillata Ménétriés 83 SIGLOPHORA Butler 72 sigma Den. & Schiff. 135 signata Butler 40 signifera Walker 81 silla Kononenko & Ahn 116 simplex Butler 8, 41 simplex Leech 127 SIMPLICIA Guenée 41 SIMYRA Ochsenheimer 89 SINARELLA Bryk 44 sincera Herrich-Schäffer 135 SINEUGRAPHE Boursin 135 SINNA Walker 73 SINOCHARINAE 22, 28, 90 SINOCHARIS Püngeler 90 smaragdina Walker 64 sobria Walker 62 sobrina Duponchel 135 socia Hfn. 114 sodalis Butler 8, 108 solidaginis Hübner 115 SOPHTA Walker 50 sordens Hufnagel 109 sordida Butler 96

459

sordidum Butler 39 SOROCOSTIA Rosenstock 66 spacoalis Walker 41 SPAELOTIS Boisduval 132 sparganii Esper 107 sparganoides Bang-Haas 107 speciosa Hübner 134 speculalis Swinhoe 47 SPEIREDONIINAE Swinhoe 62 SPHRAGIFERA Staudinger 24, 83 spilogramma Rambur 106 SPIRAMA Guenée 50 splendens Hübner 124 splendida Butler 78 splendida Staudinger 89 SPODOPTERA Guenée 100 squalida Butler 47 squalida Fabricius 63 squammigera Guenée 8, 65 staudingeri Leech 86 STAUROPHORA R. L. 106 stellata Butler 83 stellata Moore 103 STENBERGMANIA Bryk 48 STENHYPENA Hampson 44 stenochrysis Warren 77 STENOGRAPTA Sugi 23, 37 STENOLA Möschler 66 STENOLOBA Staudinger 25, 99 stenoptera Staudinger 122 stenoptera Sugi 37 stentzi Lederer 129 STICTOPTERINAE 22, 24, 27, 65 stigmaleuca Guenée 61 stigmosa Christoph 8, 122 STILBINA Staudinger 92 stola Strand 62 stolida Leech 127, 128 stolidoprocta Boursin 135 stramentacealis Bremer 44 streckeri Staudinger 59 STREPSIMANINAE 22 striata Haruta & Sugi 108 striata Leech 128 strigatus Fabricius 47 strigifera Butler 115 strigilata Linnaeus 43 strigilis Linnaeus 110 strigosa Den. & Schiff. 87 strigulosa Staudinger 69

460

stupenda Butler 133 stuposa Fabricius 56 stygia Butler 80 stygiana Butler 46 STYGIODRINA Boursin 102 suava Staudinger 116 suaveola Staudinger 36 suavis Staudinger 125 SUBACRONYCTA Kozh 89 subcostalis Walker 64 subcyanea Butler 47 subdistans Guenée 108 subfalcata Ménétriés 54 subflava Moore 90 subfulvida Warren 51 subgigas Inoue 8, 70 SUBLEUCONYCTA Kozh 86 submarginata Walker 8, 60 submira Sugi 33 submirella Ahn 33 submorosa Strand 41 subornata Leech 88 subpulchra Alphéraky 87 subrecta Sugi 53 subrosacea Ahn 81 subrosea Butler 50 subsatura Guenée 61 subtusa Den. & Schiff. 117 subviolacea Butler 64 subviridis Butler 104 suecia Aurivillius 133 suffusa Walker 35 SUGIA Ueda 80 sugii Kinoshita 88 sugii Kononenko 118 sugii Owada 40 sugii Ronkay 116 SUGITANIA Matsumura 117 sugitanii Boursin 99 SUPERSYPNOIDES Berio 60 suspecta Hübner 112 sviridovi Owada 43 sylpha Butler 72 SYNGRAPHA Hübner 79 Sypna infrapicta Strand 60 SYPNINI Holloway 27, 60 SYPNOIDES Hampson 24, 60 T tabida Butler

133,134

taeniata Snellen 8, 66 takao Sugi 37 TAMBA Walker 23, 34 tancrei Graeser 93 tanitalis 29 tapaishana Draudt 104 tarsicrinalis Knoch 44 tarsicrinata Bryk 43 tarsipennalis Treitschke 42 TELORTA Warren 113 tentacularia Linnaeus 43 tenuialis Rebel 7, 42, 43 TERATOGLAEA Sugi 114 textilis Leech 36 thalictri Borkhausen 51 THYAS Hübner 55 THYSANOPLUSIA Ichin 75 tigrina Kononenko 112 TILIACEA Tutt 112 timandra Alphéraky 36, 37 TINOLINI Moore 27, 61 TIRACOLA Moore 125 tirhaca Cramer 55 togata Esper 112 tokiensis Butler 103 tokionis Butler 130 TOXOCAMPINI Guenée 27, 53 trabealis Scopoli 84 TRACHEA Ochsnheimer 25, 103 transducta Walker 101 transversa Guenée 84 transversa Hufnagel 115 transversa Moore 49, 97 trapezina Linnaeus 118 TRIAENA Hübner 88 triangulalis Leech 8, 70 triangularis Moore 129 TRICHOCLEA Grote 122 TRICHOPLUSIA McDunnough 75 TRICHOSEA Grote 85 tricristata Draudt 8, 92 tridens Den. & Schiff. 88 trifolii Hufnagel 122 trifurca Eversmann 130 trigonalis Guenée 48 trigonifera Hampson 7, 34 trilatalis Walker 111 trilinea de Joannis 8, 84 trilinea Marumo 68 trilinealis Bremer 40

trilineata Leech 36 trimaculata Bremer 86 trinotata Walker 66 tripartita Butler 92 triphaenoides Walker 55 TRIPHAENOPSIS Butler 104 triplasia Linnaeus 75 triplex Leech 42 tripunctata Butler 116 TRISATELES Tams 23, 34 tristalis Fabricius 40 tristalis Lederer 47 tristalis Leech 84 tristis Butler 41 TRISULOIDES Butler 85 tsushimana Sugi 87 tunicata Graeser 113 turbida Esper 124 turca Linnaeus 126 turpis Butler 111 tutti Kostrowicky 77 tyrannus Guenée 52 TYTINAE 22 U UFEINAE 22 umbra Hufnagel 94 umbratica Linnaeus 91 umbrosa Butler 62 umbrosalis Staudinger 42, 43 unca Houlbert 86 unctalis Staudinger 29 uncula Clerck 80 undata Fabricius 55 undosa Leech 134 unduligera Butler 7, 64 unica Leech 124 unicolor Staudinger 117 uniformis Draudt 122 uniformis Sugi 74

USBECA Püngeler 89 ussurica Kononenko 118 ussuriensis Bremer 63 ussuriensis Dufay 75 ussuriensis Petersen 107 ussuriensis Rebel 39 ustulata Butler 114

vulnerata Butler 96 vulpecula Lederer 113 vulpina Grote 87

V

X XANTHIA Ochsenheimer 112 XANTHOCOSMIA Sugi 118 XANTHODES Guenée 24, 84 XANTHOGRAPTA Hampson 25, 111 xanthoma Prout 42 XANTHOMANTIS Warren 85 xena Staudinger 125 XENAPAMEA Sugi 110 XENOTRACHEA Sugi 25, 104 XESTIA Hübner 133 XYLENA Ochsenheimer 115 XYLENINA 28 XYLENINAE 22, 25, 28, 100 XYLENINI Guenée 28, 112 XYLOMOIA Staudinger 110 XYLOPOLIA Sugi 120

VALERIA Stephens 92 valida auct 132 variabilis Poole 78 variegata Oberthür 118 variolata Smith 125 vasava Butler 36 vecordialis Swinhoe 47 velutina Eversmann 126 venusta Leech 90, 114 venustula Hübner 101 versuta Püngeler 135 vespertilio Draudt 123 vespertilio Fabricius 50 veterina Lederer 109 vetustella Walker 66 viciae Hübner 53 vidua Staudinger 134 VILLOSA Koch 62 VIMINIA Chapman 88 violacea Butler 61 violacealis Staudinger 43 violetta Boursin 131 virens Butler 133 virescens Sugi 8, 80 VIRGO Staudinger 25, 111 virgo Treitschke 96 viridimacula Graeser 116 viridis Leech 93 v-minus Oberthür 79 vulcanea Butler 53 vulgaris Butler 106

W WESTERMANIINAE 22 WESTERMANNIINAE 22

Y yoshinensis Wileman & West

8, 67

Z zambesita Walker 61 ZANCLOGNATHA Lederer 42 ZEKELITA Walker 23, 45 zibellina Felder & Rogenhofer 64 zilla Butler 46 ZIRONA Walker 61 ZONOPLUSIA Chou & Lu 76 zosimi Hübner 77

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