Department of Genetics, University of California, Davis, California 95616, U.S.A.. David J. Keil ... northern Mexico is indistinguishable from G. ca- nescens DC. of ...
New Combinations New CombinationsIn in Gnaphalium Inuleae) (Asteraceae:Inuleae) Gnaphalium(Asteraceae: G. Ledyard Stebbins G. Ledyard Stebbins Department Genetics, University California95616, 95616, U.S.A. Davis, California California, Davis, Department of Genetics, University of California,
David ]. J. Keil Luis Obispo, Biological Sciences Department, California Polytechnic Biological Obispo, Department, California University, San Luis Polytechnic State University, California 93407, U.S.A. 93407, U.S.A.
ABSTRACT. Because Because of extensive ABSTRACT. extensive morphological morphological inat the termediacy, plants formerly the spetermediacy, plants distinguishedat formerlydistinguished species level level as Gnaphalium G. microcemicrocebeneolens, G. Gnaphalium beneolens, phalum, G. thermale, treated thermale, and G. wrightii phalum, wrightii are treated as intergrading races of the the widespread as G. canescanesintergrading races widespread G. cens. The The new new combinations, cens. G. canescens subsp. combinations, G. subsp. G. canescens canescens subsp. beneolens, beneolens, G. subsp. microcephalum, microcephalum, and G. canescens subsp. thermale made. canescens subsp. thermale are made.
In preparing the treatment In preparing the treatment of Gnaphalium L. for for Gnaphalium L. the forthcoming flora The the flora of California, The Jepson ManCalifornia, forthcoming Jepson Manwe have have found found that that the the species ual, we recspecies currently currently recbeneolens A. Davidson, ognized thermale Davidson, G. thermale ognized as G. beneolens E. Nelson, G. microcephalum Nelson, G. microcephalum Nutt., Nutt., and G. wrighwrighA. Gray such aa degree with respect intergrade to such tii A. Gray intergrade degree with respect the characters used in to the in current charactersused current keys (Ferris, 1960; keys (Ferris, Munz, differentiatethemthemMunz, 1959, 1959, 1968, 1968, 1974) 1974) to differentiate decurrent versus decurrent versus nondecurrent bases, nature nondecurrentleaf bases, nature of character of capitulescence, tomentum, size and and tomentum, character capitulescence, size and acute obtuse shape of heads, and acute versus obtuse phyllary versus heads, shape phyllary tips-that be separated cannot consistently tips-that they they cannot consistently be separated from each each other. In addition, from wrightii of eastother. In G. wrightii eastaddition, G. ernmost Arizona, New Mexico, ernmost California, New and California, Arizona, Mexico, and is indistinguishable northern Mexico northern Mexico is from G. cafrom G. caindistinguishable central Mexico. DC. of central nescens DC. Mexico. These These entities entities are are therefore all all merged therefore polymorphic speinto aa single merged into single polymorphic speG. canescens. Recognizable cies, G. tenRecognizable geographic geographic tendencies in in variation dencies variation are are maintained as subspecies. maintained as subspecies. new combinations The proThe following combinationsare are therefore therefore profollowing new posed: posed: Gnaphalium beneolens canescens DC. DC. subsp. Gnaphalium canescens subsp. beneolens Stebbins& (A. Davidson) Stebbins & Keil, comb. nov. nov. GnaGnaKeil, comb. (A. Davidson) beneolens A. A. Davidson, phalium beneolens Davidson, Bull. Bull. So. So. Calif. Calif. phalium Acad. Sci. 17: 17. 17. 1918. 1918.
microceGnaphalium DC. subsp. canescens DC. subsp. microceGnaphalium canescens nov. comb. nov. phalum (Nutt.) & Keil, Stebbins & Keil, comb. (Nutt.) Stebbins phalum Gnaphalium microcephalum Nutt., Trans. Trans. Nutt., Gnaphalium microcephalum 404. 1841. 1841. Amer. 7: 404. Phil. Soc. n.s., 7: Amer. Phil. Gnaphalium DC. subsp. thermale (E. canescens DC. (E. subsp. thermale Gnaphalium canescens comb. nov. nov. GnaGnaNelson) Stebbins Stebbins & & Keil, Keil, comb. Nelson) E. Nelson, Bot. Gaz. Gaz. (Crawphalium thermale thermale E. Nelson, Bot. (Crawphalium fordsville) 1900. fordsville) 30: 121. 1900. Gnaphalium (insubsp. canescens (inGnaphalium canescens subsp. from the the southwestern southwestern cluding wrightii) occurs G. wrightii) occurs from cluding G. In the the desert desert mounUnited mounStates to central central Mexico. Mexico. In United States into subspecies tains grades into Californiait it grades eastern California tains of eastern subspecies Califorsouthern Califormicrocephalum, race chiefly chiefly of southern microcephalum, aa race beneolens into subspecies nia. in turn turn grades nia. This This in subspecies beneolens grades into Coast Ranges Ranges of California. California. Plants Plants of South Coast in the the South to are mostly the Nevada foothills assignable foothills are the Sierra Sierra Nevada mostly assignable to above beneolens, but above 1,000 m, plants subspecies but m, 1,000 beneolens, plants subspecies The latter latter agree thermale. The more with with subspecies subspecies thermale. agree more and northnorthMountainsand ranges eastwardto the the Rocky Rocky Mountains ranges eastward taxa these taxa ward BritishColumbia. Columbia.Descriptions ward to to British Descriptionsof these in our treatment and are included in our treatment of are included and their their ranges ranges in The The Jepson Gnaphalium Jepson Manual. Manual. Gnaphalium in Literature LiteratureCited Cited
L. in L. 98-613 in Ferris, R. S. 1960. Compositae. S. 1960. Ferris,R. Pp. 98-613 Compositae.Pp. of the the Pacific Pacific Floraof Abrams Flora Illustrated & R. R. S. S. Ferris, Abrams& Ferris,Illustrated Stanford. States, vol 4. Stanford StanfordUniv. Univ. Press, Press, Stanford. States,vol California Munz, Flora. Univ. Univ. California Flora. A California California P. A. A. 1959. 1959. A Munz,P. Press, Press, Berkeley. Berkeley. Flora.Univ. Univ. - - - . 1968. to Flora. California to A California 1968. Supplement Supplement California CaliforniaPress, Press, Berkeley. Berkeley. - - - . 1974. California.Univ. Univ. A Flora Floraof of Southern SouthernCalifornia. 1974. A California CaliforniaPress, Press, Berkeley. Berkeley.
437. 1992. NOVON 2: 437. 1992. NOVON
