stellate glandular scales - Naturalis repository

5 downloads 0 Views 736KB Size Report
Jul 12, 1999 - The plant is obviously a Mallotus Lour, (stellate hairs and ... marginally looped and closed, veins scalariform, veinlets densely reticulate.
BLUMEA

46

67-69

(2001)

Mallotus kongkandae (Euphorbiaceae), a new species

Peter+C. van

1

Welzen

from Thailand

I & Kanlaya

Phattarahirankanok

Summary

Mallotus

kongkandae

glandular scales endemic

Key

when

to northern

words

is

a

dry,

new

species

in

thick-walled

section

fruits,

Rottlera.

and the

Typical

seemingly

the ovate

are

caducous

leaves, greenish

columella. The

plant

is

Thailand.

: Euphorbiaceae,

Mallotus

kongkandae,

Thailand.

INTRODUCTION

Chayamarit

et

al.

collected

recently

Thailand (Kamphaeng Phet hairs and leaves

glandular scales)

comprises

several

species)

and

M.

montanus

often

when fresh)

most

locally

(Miill. Arg.) Airy Shaw,

of all,

the

glandular larger,

leaves, merits the description of a are

new

unknown. The

(still)

scales

thicker

kongkandae

species,

new

another, very locally endemic species

Mallotus

very

(red

or

species,

pilis

aggregata ovata

inermis, paries

2-3

Fructus

mm

stellatis

simplicibusque

subglobosus

crassus.

Chong Yen,

Nationaal

12

Herbarium

Netherlands;

2) Bangkok Thailand.

July 1999,

1340

Nederland,

when

mm

thick)

together

dry,

the

in the other and the

with the

the staminateand

ovate

pistillate

named after the collector, represents

—Typus:

et

nov.

elobatus K.



1

Infructescentiae

squamis glandulosis

Chayamarit

Fig.

squamis glandulosis viridulis.

infra tomentosa trinervata.

et al.

BKF, K), Thailand, Northern District, Kamphaeng Phet Prov.,

The

(2-3

Welzen & Phattarahirankanok, spec.

Arbor. Indumentum densum

solitariae.

are,

yellowish

in section Rottlera.

Folia alternata sed distaliter terminales

1)

Typical

whitish

though

repandus

species ( M. lepto-

pallidus (Airy Shaw) Airy

walled fruits

even

of

(stellate

Section Rottlera

Mull. Arg., M.

endemic

M.

provinces

Mallotus Lour,

Shaw (1968), because the

caducous fruit column. This character combination,

seemingly

flowers

Airy

a

(Airy Shaw, 1972).

is different in several characters.

specimen

greenish (orange

of the northern

one

obviously

widespread species (M. philippensis (Lam.)

stachyus Hook.f., new

is

plant

unarmed

are

(Willd.) Miill. Arg.), but also several,

Shaw). The

in

specimen The

of section Rottlera (Willd.)

alternate and the fruits

are

a

Province).

1551

Mae

dense obtectus

(holo L;

Wong

iso

National

AAU, Park,

m.

Universiteit

Leiden

branch,

P.O.

Box

9514,

2300 RA

Leiden,

e-mail: [email protected]

Forest Herbarium

(BKF), Royal Forestry

Department, Chatuchak, Bangkok 10900,

68

Fig.

BLUMEA

1. Mallotus

glandular one

areas

kongkandae adaxially;

attached seed

Tree,

c.

8

m

tomentose

glandular

c.

Welzen

fruit;

( K. Chayamarit

&

et al.

most

scales.

parts

terete,

cm

through

2-3

mm

when young, c.

tendency

long,

No.

1,

2001

fruit

a.

Habit;

showing

b.

thick

base

wall;

of leaf e.

with

columella

two

with

155J, L).

Stipules triangular,

1.5-5

46,

Phattarahirankanok.

section

high; flowering twigs

on

inside. Leaves alternate with

petiole

d.

Vol.

thick, young parts of

consisting 1.5 to

somewhat

by

0.6

mm,

terminal

pulvinate

on

Indumentum

and stellate hairs and

tomentose

groups at

tomentose.

simple

outside, glabrous

the branches,

both ends, with

simple;

longitudinal

P. C.

when

ridges

Welzen

van

dry,

&

K. Phattarahirankanok:

3.4-5.5

by

symmetric, drying greenish, older leaves seen), apex

seemingly glandless (only

Thailand

kongkandae from

when young; blade ovate, 8-18

tomentose

width ratio 2.4-3.3, chartaceous, entire and

Mallotus

cm,

length/

base rounded,

gradually

69

margin

acute to

acu-

minate, mucronulate, upper surface glabrous except for the basal part of the midrib, with 2, black,

basally

glandular

lower surface tomentose, white,

petiole attachment,

areas near

glandular

nerved, trinerved, slightly raised above, raised underneath,

marginally looped inflorescences 3.5-8

inside, bracts

tomentose; bracts and bracteoles 1

c.

terete, tomentose, flattened

and closed, veins scalariform, veinlets

and flowers unknown. Infructescences

long,

cm

by

0.8

mm,

dorsoventrally),

of orange

(green

when

bracteoles

0.5

c.

with abscission

subbasally

mm

dry) glandular scales,

column

apically;

6

c.

mm

11

c.

by

3.5

Distribution Mae

Wong

Ecology Note biaceae

Royal

single

mm,





not

branching,

outside, glabrous

Pedicels 2.5-3

mm

Forest

long,

capsular, subglobose (slightly

mm, not

armed, with dense covering

somewhat ribbed and

long, apically

10-12 per side,

wall thick,

broadened and

ridge

on

pustular

2

c.

mm

when

dry,

in middle

T-shaped,

to

caducous.

axillary side,

Endemic in Thailand, Northern Division,

in

Fruiting

Chong July.

Yen.

Only

Kamphaeng

known from the type

c.

5

by

Altitude 1340

Phet Province,

specimen.

m.

Named in honour of the main collector and initiator of the Thai

Project,

in size;

per locule, dark brown, naked.

National Park, —

tomentose

mm.

Fruits

Seeds ± ovoid but somewhat flattened and with central 4.2

mm

reticulate. Staminate

terminal, single,

0.5

by

tardily dehiscing loculicidally, partly septicidally; 3

0.7

by

nerves

densely

triangular,

zone.

lobed, 12-14 by

not

1

c.

scales many, domatia absent; venation penni-

Kongkanda

Mrs. Dr.

Chayamarit

of the Forest

Euphor-

Herbarium (BKF),

Department, Bangkok.

ACKNOWLEDGEMENTS

We like

a

thank

to

translation

Jan

joint Programme by

and

van

into Latin. We

Biotechnology)

Os

are

(L)

also

the Thai

for

for his

Research

funding

beautiful

grateful to

the

BRT

Fund

drawing

and

(Biodiversity

and the National

Euphorbiaceae of

Jan Frits

Veldkamp (L)

Research and

Thailand

Centre

for

the

Training Programme,

for Genetic

Engineering

Project.

REFERENCES

Airy Shaw,

H.K.

1968. Notes

H.K.

1972. The

on

Malesian

and

other Asiatic

Euphorbiaceae.

Kew

Bull.

392.

Airy Shaw,

Euphorbiaceae

of Siam.

Kew

Bull.

26:

292, 293,

299-301.

21:

391,