zoologische mededelingen - Naturalis repository

ZOOLOGISCHE MEDEDELINGEN UITGEGEVEN DOOR

RIJKSMUSEUM

V A N

H E T

NATUURLIJKE HISTORIE

T E

LEIDEN

(MINISTERIE V A N C U L T U U R , R E C R E A T I EE N M A A T S C H A P P E L I J K

Deel 51 no. 13

WERK)

28 juni 1977

T H E A S I A N G E N U S PLEURONOTA KRAATZ AND ALLIED FORMS: A CLARIFICATION (COLEOPTERA: CETONIIDAE) by J. K R I K K E N Rijksmuseum van Natuurlijke Historie, Leiden W i t h 15 text-figures and one plate ABSTRACT Two

A s i a n genera

Kraatz (= Mikšić).

Macronotiola

Key

are g i v e n . T h e ( G e s t r o ) , elongata

alleni

Pleuronota

revised:

nov. ( =

f o r both g e n e r a and checklists o f

sensu

Pleuronota

the k n o w n species

are

sp. nov. ( B o r n e o ) is described a n d illustrated. S o m e new records

f o l l o w i n g new

combinations h a d to be made.

( G o r y & P e r c h e r o n ) , longiuscula

( F a i r m a i r e ) , setosa

Macronotiola), p atricia guttata

diagnosed a n d their nomenclature is

M i k š i ć , syn. nov.) a n d Macronotop s

frameworks

g i v e n . Pleuronota

fosquamosa

are

setosa

( M o s e r ) , setosa

imuganica

( G o r y & P e r c h e r o n ) ( f r o m Macronota)

( F a i r m a i r e ) , fulvop ilosa

( F a i r m a i r e ) , nigrop ubescens

r o w ) , sexmaculata

( K r a a t z ) , vuilleti

goin) (all f r o m

Pleuronota).

vuilleti

To

Pleuronota:

( M o s e r ) , monticola

aenea

(Gestro),

(Mikšić)

(all

ru-

from

; to Macronotop s:

fulvo-

( M i k š i ć ) , ovalicep s

(Ar

( B o u r g o i n ) , vuilleti

olivaceofusca

(Bour

Several species of Asian cetoniines placed in Macronota auctorum and Pleuronota Kraatz were recently rearranged, keyed and described by Mikšić (1972, 1976). H e proposed a new genus, Macronotiola, to accomodate some oddities up till then combined with Macronota. Because of a serious misinter pretation of the typespecies concept, however, a mixup took place, which is rectified below to prevent further confusion. The two genera involved are diagnosed, one is renamed, and all the species mentioned by M i k š i ć (l.c.), the type of Pleuronota excepted, are recombined. Furthermore, one new and one old species are added to Pleuronota, and new records are given, mainly based on material kept in Leiden. The key frameworks given below only provide a gross classification of the two genera. I have not attempted to devise exhaustive keys to all the species and subspecies recognized so far, for the following reasons: (1) my primary intention is to clarify the supraspecific classification; (2) the material studied

ZOOLOGISCHE M E D E D E L I N G E N

200

51 (1977)

so far, also with the present additions, is extremely poor; (3) some typespecimens were not available for study. The maps (figs. 14, 15) provide a reasonable approximation of the known distribution of these cetoniines. Pleuronota K r a a t z and Macronotops nov. Group diagnosis. — H a i r y but not bumblebee-like cetoniiform beetles with strongly lobate pronotal base; if glabrous, species with coniform pygidium in male sex. Derm lacking velvety shine, usually with whitish markings. Clypeus never deeply incised. Identification. — The two genera can be separated as follows: Pleuronota

Macronotops

Clypeal margin anteriorly

bisinuate, not raised ( f i g . 1)

2.

S

n o r m a l ( f i g . 9)

elongate ( f i g .

3.

$ pygidium

conical ( f i g . 7)

weakly convex

h u m p e d ( f i g . 10)

not h u m p e d ( f i g .

1.

4.

antennal

Basal

club

piece

of

phallus

straight o r rounded, raised ( f i g . 11) 12) 13)

dorsally

Affinities. — Pleuronota, Macronotops, Bombodes, Taeniodera, Euselates and some other genera are all closely allied inter se, and, in my opinion, only a thorough phylogenetic analysis could clarify their relationships. A t any rate, the bumblebee-like forms united in Bombodes Westwood are very close, and may eventually turn out to be closest to the two genera dealt with here. Pleuronota K r a a t z Pleuronota Macronotiola

K r a a t z , 1892:

312

M i k s i c , 1972:

(key to s p e c i e s ) ;

1976:

( m o n o t y p i c : Pleuronota 123

31

(type-sp. Macronota

( i n k e y ) , 152

octomaculata elongata

(diagnosis),

153

Kraatz).

Gory & Percheron), (key to species).

125

Syn. von.

Type-species. — Pleuronota octomaculata Kraatz, by monotypy. Miksic (1976) considers this species aberrant (sic), and terms its systematic position rather dubious. Considering the shape of the clypeus, however, P. octomaculata clearly belongs i n his Macronotiola, while the other three characters mentioned above are consistent as well, as proved by the study of the male sex. Because the type-species of Pleuronota and Macronotiola are here considered congeneric, the latter name falls as a junior synonym. Miksic's use of the term type-species implies that a type-species must be typical of the genus; there is, however, no such obligation in zoological nomenclature. Distribution and composition. — Oriental Region (map, fig. 15); 9 species, one with two subspecies.

KRIKKEN,

P L E U R O N O T A A N D A L L I E D FORMS

K e y framework for

201

Pleuronota

ι. P ronotal midline distinctly impressed, in some cases limited by paramedian costae. P ronotal sides more or less angulate and posterolateral angles more or less distinct 2 — Only pronotal base may be superficially impressed. P ronotal sides and posterolateral angles widely rounded off monticola + aenea + alleni (see table under alleni) 2. P ronotal impression constricted about halfway pronotal length. Small species (length less than 15 mm) elongata — P ronotal impression continuous 3 3. Dorsum shortly setose or glabrous longiuscula — Dorsum abundantly setose, setae long or very long 4 4. Head posteriorly with longitudinal smooth costa. P aramedian costae of pronotum distinct rufosquamosa + setosa — Head posteriorly lacking longitudinal smooth costa. P aramedian costae of pronotum indistinct 5 5. Colour more or less metallic. External denticles of fore tibia equally wide apart. Elytral apex lacking cretaceous marking . . . patricia — Colour generally black-brown. Distance between denticles 1-2 of fore tibia greater than between 2-3. Elytral apex with cretaceous marking octomaculata

Checklist of Pleuronota species ) 1

*aenea Gestro, 1879: 11 (Macronota), type in Genoa. — Sumatra (typeloc. Singalan). — S of monticola? — L . alleni K r i k k e n , present paper, holotype