Zootaxa, A catalog of the Tenuipalpidae (Acari)

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May 11, 2009 - A catalog of the Tenuipalpidae (Acari) of the World with a key to genera. NORA C. MESA, RONALD OCHOA, W. CALVIN WELBOURN, ...
ZOOTAXA 2098

A catalog of the Tenuipalpidae (Acari) of the World with a key to genera NORA C. MESA, RONALD OCHOA, W. CALVIN WELBOURN, GREGORY A. EVANS & GILBERTO J. DE MORAES

Magnolia Press Auckland, New Zealand

NORA C. MESA, RONALD OCHOA, W. CALVIN WELBOURN, GREGORY A. EVANS & GILBERTO J. DE MORAES A catalog of the Tenuipalpidae (Acari) of the World with a key to genera (Zootaxa 2098) 185 pp.; 30 cm. 11 May 2009 ISBN 978-1-86977-345-8 (paperback) ISBN 978-1-86977-346-5 (Online edition)

FIRST PUBLISHED IN 2009 BY Magnolia Press P.O. Box 41-383 Auckland 1346 New Zealand e-mail: [email protected] http://www.mapress.com/zootaxa/

© 2009 Magnolia Press All rights reserved.

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(Print edition)

ISSN 1175-5334

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2 · Zootaxa 2098 © 2009 Magnolia Press

MESA ET AL.

Zootaxa 2098: 1–185 (2009) www.mapress.com / zootaxa/

ISSN 1175-5326 (print edition)

Copyright © 2009 · Magnolia Press

ISSN 1175-5334 (online edition)

ZOOTAXA

A catalog of the Tenuipalpidae (Acari) of the World with a key to genera NORA C. MESA, RONALD OCHOA2, W. CALVIN WELBOURN3, GREGORY A. EVANS4 & GILBERTO J. DE MORAES5 Universidad Nacional de Colombia sede Palmira, Facultad de Ciencias Agropecuarias. A.A. 273, Palmira-Colombia [email protected] 2 US Department of Agriculture, ARS, Systematic Entomology Lab., Beltsville, MD, U.S.A. [email protected] 3 Florida Department of Agriculture & Consumer Services, Division of Plant Industry, Gainesville, Florida, U.S.A. [email protected] 4 US Department of Agriculture, APHIS, PPQ., Beltsville, MD, U.S.A. [email protected] 5 Depto. Entomologia, Fitopatologia e Zoologia Agrícola, Universidade de São Paulo, Escola Superior de Agricultura “Luiz de Queiroz”, 13418-900 Piracicaba-SP, Brazil.

TABLE OF CONTENTS ABSTRACT . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 4 ACKNOWLWDGEMENTS . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 4 INTRODUCTION & LITERATURE REVIEW . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 5 Zoogeography of the Genera and Species of Tenuipalpidae. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 7 Table 1. Tenuipalpid species described from each zoogeographic region . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 9 Format of catalog . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 9 FAMILY TENUIPALPIDAE BERLESE. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 11 Genus Acaricis Beard & Gerson . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . see addendum Genus Aegyptobia Sayed . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 11 Genus Afronychus Meyer . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 20 Genus Amblypalpus Mitrofanov & Strunkova . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 20 Genus Australopalpus Smiley & Gerson . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 20 Genus Brevipalpus Donnadieu . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 20 Genus Capedulia Meyer . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 49 Genus Cenopalpus Pritchard & Baker . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 50 Genus Chaudhripalpus Mesa, Welbourn & Evans, new genus . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 57 Genus Coleacarus Meyer . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 57 Genus Crossipalpus Smiley, Frost & Gerson . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 58 Genus Dolichotetranychus Sayed . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 58 Genus Krugeria Meyer . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 60 Genus Larvacarus Baker & Pritchard. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 61 Genus Lisaepalpus Smiley & Gerson . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 61 Genus Macfarlaniella Baker & Pritchard . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 61 Genus Magdalenapalpus, Mesa, Welbourn & Evans, new genus . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 61 Genus Obdulia Pritchard & Baker . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 62 Genus Obuloides Baker & Tuttle . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 62 Genus Pentamerismus McGregor . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 62 Genus Philippipalpus Corpuz-Raros . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 64 Genus Phyllotetranychus Sayed . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 64 Genus Phytoptipalpus Tragardh . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 65 Genus Priscapalpus De Leon . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 67 Genus Prolixus Beard, Fan & Walter . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 67 Genus Pseudoleptus Bruyant . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 67 Genus Raoiella Hirst. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 68 Genus Raoiellana Baker & Tuttle. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 70

Accepted by Z.-Q. Zhang: 1 Apr. 2009; published: 11 May 2009

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Genus Tegopalpus Womersley . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 70 Genus Tenuilichus Mohanasundaram . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 71 Genus Tenuipalpus Donnadieu . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 71 Genus Terminalichus Anwarullah & Khan1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 98 Genus Ultratenuipalpus Mitrofanov . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 99 Genus Urigersonus Mesa, Ochoa & Evans, new genus . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 102 Species incertae sedis . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 102 Addendum (Genus Acaricis Beard & Gerson) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 102 FINAL CONSIDERATIONS. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 103 MORPHOLOGY . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 105 Table 2. Body setae and shape, rostral shield and number of palpal segments. . . . . . . . . . . . . . . . . . . . . . . . . . . . . 107 KEY TO WORLD GENERA . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 110 APPENDIX A. Zoogeographic distribution of tenuipalpid species. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 131 REFERENCES. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 149 INDEX OF SPECIES. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 161 HOST INDEX . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 179

ABSTRACT The family Tenuipalpidae is worldwide in distribution and includes several economically important pest species. Species of the genus Brevipalpus have been identified as vectors of rhabdoviruses that cause diseases such as citrus leprosis, coffee ring spot virus, passion fruit green spot virus and others that reduce the production and the life span of the plants. An updated list of the Tenuipalpidae (Acari: Prostigmata), a key to the genera of the world and data on zoogeographical distribution of the species are presented. The catalogue provides information on 891 known species (including 3 species incertae sedis) belonging to 34 genera. The valid name of each species, the author, date and page of the corresponding original description, type locality and host, species designation(s), references to species redescription(s) and other complementary information are presented. The genera Rarosiella Rimando 1996 and Neoraoiella Mohanasundaram 1996 are synonymized with Raoiella Hirst 1924. The genus Meyeraepalpus Smiley, Frost & Gerson 1996 is synonymized with the genus Aegyptobia Sayed 1950. Three new genera, Magdalenapalpus, Chaudhripalpus and Urigersonus are erected. Indices of the plant host species and families associated with each species, the number and regional distribution of species in each genus, and a list of relevant publications of species from each zoogeographical region are included. Key words: Acari, Prostigmata, Tenuipalpidae, catalog, taxonomy, flat mites, false spider mites, virus, vector, fauna

ACKNOWLEDGEMENTS We are very grateful to Debra Creel (SEL-USDA), Alice Sanders (Division of Plant Industry, Gainesville, Florida), personnel of the Divisão de Biblioteca e Documentaçao (ESALQ, Piracicaba, Brazil) and National Agricultural Library, USDA (Beltsville, MD) for their assistance in the search, copying, retrieval and loan of literature used in this catalogue. We thank C.H.W. Flechtmann (ESALQ), Uri Gerson (Hebrew University of Jerusalem, Israel), Jenny Beard (Queensland Museum, Australia), Ibrahim Al-Gboory (University of Baghdad, Iraq), Ethan Kane and Eric McDonald (APHIS-PPQ-USDA) for their suggestions and for sharing several publications of interest to this work. The following authors also helped by sending publications included in this catalogue: Marisa Castagnoli (Laboratory of Acarology, Istituto Sperimentale per la Zoologia Agraria, Italy), Edward A. Ueckermann (ARC-Plant Protection Research Institute, South Africa) and Markus Knapp (International Centre of Insect Physiology and Ecology (ICIPE), Kenya). We thank Stuart McKamy, Douglass Miller and Gary Miller (SEL-USDA) for their review and comments regarding this manuscript. Our thanks to Elliot Kitajima (ESALQ), Jose Carlos Rodrigues (University of Puerto Rico) and Carl Childers (University of Florida) for their comments on the biology and ecology of flat mites. Our appreciation to Isabel Zuluaga Mesa, daughter of the senior author, for her help in sorting and organizing literature.

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A CATALOGUE OF THE TENUIPALPIDAE (ACARI) OF THE WORLD WITH A KEY TO GENERA The Tenuipalpidae (Acari: Prostigmata: Tetranychoidea), commonly known as flat mites or false spider mites, is worldwide in distribution and is comprised of 891 valid species belonging to 34 genera. All tenuipalpid species are phytophagous and damage plants by directly feeding on the epidermal cells of the stems, leaves and fruits. Several species are known to vector plant virus (Rodrigues et al. 2003; Childers and Derrick 2003; Kitajima et al. 2003) and some species have been collected carrying fungal spores associated with plants (Evans et al. 1991, Ochoa et al. 1994). The majority of the species that cause economic damage to cultivated plants belongs to the two largest genera, Brevipalpus Donnadieu and Tenuipalpus Donnadieu.

Literature Review—A Timeline This literature review is intended to cover only the taxonomic highlights (ie new genera, family names, etc.) in the Tenuipalpidae. 1. Dugès (1834) described the first tenuipalpid as Trombidium caudatus from France, placing it in the Trombidiidae Leach, 1815. 2. Donnadieu (1875) erected Tenuipalpus and Brevipalpus, placing both in the Tetranychidae and transfered Trombidium caudatus to Tenuipalpus. 3. Bruyant (1911) erected Pseudoleptus and placed it in the Tetranychidae. 4. Berlese (1913) published the family-group name Tenuipalpini. 5. Ewing (1922) erected the family Phytoptipalpidae for the aberrant Old World genus Phytoptipalpus Trägärdh (1904). This genus was not placed in the Tenuipalpidae until 1952 by Pritchard and Baker. 6. Hirst (1924) erected the genus Raoiella and placed it in the Tetranychidae. 7. Oudemans (1928b) erected the Pseudoleptidae to include Pseudoleptus and Raoiella, both later (Oudemans, 1938) moved to the new family Trichadenidae; ultimately, they were transferred to the Tenuipalpidae by Sayed (1950a). 8. Sayed (1938) erected Phyllotetranychus and Dolichotetranychus placing them in the Tetranychidae. 9. Womersley (1940) erected Tegopalpus from Australia and placed it in the Tetranychidae. 10. Sayed (1942c) included Tenuipalpus in the Pseudoleptidae and in a different paper published the same year (Sayed, 1942b) considered it to be a junior synonym of Brevipalpus. 11. Baker (1945) moved Phyllotetranchyus and Tegopalpus to the Trichadenidae. He also recognized Brevipalpus and Tenuipalpus as distinct genera in the Trichadenidae. 12. Baker (1949) revised Brevipalpus placing it in the Pseudoleptidae, with 39 species. 13. McGregor (1949) published the first synthesis of the Tenuipalpidae (then referred to as the Pseudoleptidae), reporting a total of 13 species in the genera Trichadenus (1 sp.), Tenuipalpus (3 spp.), Brevipalpus (6 spp.) and erected the genus Pentamerismus (3 spp.). 14. Sayed (1950a) was the first to use the name Tenuipalpidae, in which he included Brevipalpus, Tenuipalpus, Phyllotetranychus, Tegopalpus and Dolichotetranychus; however, he did not refer to Berlese (1913), who had previously published the family-group name Tenuipalpini. The ensuing confusion regarding the appropriate author and family name resulted in most researchers using Phytoptipalpidae. 15. Sayed (1950a) divided the Tenuipalpidae into the Phytoptipalpinae, which included only the Phytoptipalpus, and the Tenuipalpinae, which included, Aegyptobia, Brevipalpus, Dolichotetranychus, Phyllotetranychus, Pseudoleptus, Raoiella, Tegopalpus and Tenuipalpus. In an immediately later paper, Sayed (1950b) formally described the genus Aegyptobia. 16. In June, 1952 Baker & Wharton considered Aegyptobia to be a synonym of Pentamerismus. They also described Larvacarus with the type species Phytoptipalpus transistans Ewing, placing it in the Phytoptipalpidae. A month later, Reck (1952) published a description of Phytoptipalpoides, also designating P. transistans Ewing, as the type species. Since the description of Larvacarus predated the description of Phytoptipalpoides, the latter is an objective junior synonym of Larvacarus. A catalog of the Tenuipalpidae of the world with a key to genera

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17. Cunliffe (1955) re-established the name Tenuipalpidae, based on Berlese’s 1913 publication; the name Tenuipalpidae has been adopted by most subsequent authors on the family. 18. Pritchard & Baker (1958) published the first major revision of the family, comprising 143 species in 14 genera. They erected the genera Cenopalpus, Colopalpus and Obdulia, and re-establish and transfer the genus Pentamerismus to the family Tenuipalpidae. They did not support dividing the family into subfamilies or tribes. 19. De Leon (1961b) erected the genus Priscapalpus from Mexico. 20. Baker & Pritchard (1962) erected the genus Macfarlaniella from Australia. 21. Baker & Tuttle (1972) erected the genus Raoiellana from Turkey 22. Collyer (1973c) revised the genus Tenuipalpus, then with 105 species. 23. Anwarullah & Khan (1973) erected the genus Terminalichus from India. 24. Mitrofanov (1973a) erected the following genera and/or subgenera: Aegyptopalpus, Deleonipalpus, Gnathopalpus, Livschitzia, Stenotetranychus, Tuttlepalpus, Ultratenuipalpus and Zaheria. He divided the Tenuipalpidae into two subfamilies Brevipalpinae and Tenuipalpinae, and divided the Brevipalpinae into four tribes: Brevipalpini with Brevipalpus, Brachypalpus, Cenopalpus, Hystripalpus, Priscapalpus, Pritchardipalpus, Tauripalpus and Tegopalpus; Colopalpini with Colopalpus and Deleoniella; Pentamerismini with Aegyptobia, Oligomerismus and Pentamerismus; and Phyllotetranychini with Phyllotetranychus. Also in 1973, Mitrofanov (1973b) erected Brachypalpus, Deleoniella, Hystripalpus, Neophytoptipalpus, Oligomerismus, Pritchardipalpus and Tauripalpus. He then divided the Tenuipalpinae into four tribes: Dolichotetranychini with Dolichotetranychus and Stenotetranychus; Phytoptipalpini, with Larvacarus, Macfarlaniella, Neophytoptipalpus, Obdulia, Phytoptipalpus, Raoiella, Raoiellana and Zaheria; Pseudoleptini with Livschitzia and Pseudoleptus; and Tenuipalpini with Aegyptopalpus, Deleonipalpus, Extenuipalpus, Gnathopalpus, Tenuipalpus, Tuttlepalpus and Ultratenuipalpus. 25. Chaudhri et al. (1974) erected Amissus, later synonymized with Brevipalpus by Sepasgosarian (1983). 26. Baker & Tuttle (1975) erected the genus Obuloides. Baker et al. (1975) divided Brevipalpus into seven informal species groups, referring to them only by Roman numerals. Pegazzano (1975) erected Tauripalpoides, synonymized with Brevipalpus by Meyer (1979). 27. Corpus-Raros (1978) erected the genus Philippipalpus from the Philippines. 28. Mitrofanov & Strunkova (1978) erected the genus Amblypalpus from Tajikistan. 29. Mitrofanov & Strunkova (1979) published a revision of the Tenuipalpidae, reporting 365 species in 33 genera. Their classification differed from that of Pritchard & Baker (1958) and Meyer (1979), but used the same subfamilial and generic arrangement as Sayed (1950a). They included in the Phytoptipalpinae the genera proposed by Mitrofanov (1973b) in the tribes Dolichotetranychini and Phytoptipalpini. In Tenuipalpinae, they included the genera mentioned by Mitrofanov (1973a, b) under Brevipalpini, Pentamerismini, Phyllotetranychini, Pseudoleptini and Tenuipalpini, and considered Colopalpini to be a synonym of Tenuipalpini. Also in 1979, Meyer published a major revision of the Tenuipalpidae which included 504 species in 21 genera. She erected Afronychus, Krugeria, Capedulia and Coleacarus, listed Brachypalpus, Cenopalpus, Hystripalpus, Pritchardipalpus, Tauripalpus and Tauripalpoides as synonyms of Brevipalpus; Neophytoptipalpus and Zaheria as synonyms of Phytoptipalpus; Livschitzia and Oligomerismus as synonyms of Pentamerismus; Deleoniella as synonym of Priscapalpus; Stenotetranychus as synonym of Dolichotetranychus; and Colopalpus, Deleonipalpus, Gnathopalpus and Tuttlepalpus as synonyms of Tenuipalpus. She provided a key to the world genera and species and proposed a division of Brevipalpus into eight species groups, which she named absens, californicus, cuneatus, obovatus, phoenicis, pterinus, recki and spinosus. She considered Cenopalpus to be a synonym of Brevipalpus, and represented it as the spinosus species group. In addition, she divided other large tenuipalpid genera into species groups, as follows: Aegyptobia, groups trägardhi and macswaini; Dolichotetranychus, groups carnea, summersi and floridanus; and Tenuipalpus, groups albae, caudatus, elegans, granati, quadrisetodus and trisetosus. She did not follow Mitrofanov’s (1973a, b) family-group divisions.

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30. Sepasgosarian (1983) published the first species list of the tenuipalpids of the world, reporting 541 species in 21 genera and providing information on the name, author, date of publication and synonymies of each genus and species and a list of references. He listed Amissus as a synonym of Brevipalpus. Subsequently, Sepasgosarian (1984, 1985, 1987, 1990) published four addenda to his original list, increasing the number of species to 697, in 25 genera. 31. Ghai & Shenhmar (1984) published a detailed list of the 22 genera and 562 species of Tenuipalpidae, including a taxonomic key to the genera. 32. Baker & Tuttle (1987) reviewed the false spider mites of Mexico. They divide the species of the genus Brevipalpus into six species groups, stating that these groups corresponded to the division proposed by Baker et al. (1975). The species groups names used by Baker & Tuttle (1987) along with (in parentheses) the group numbers proposed by Baker et al. (1975) are: cuneatus (group I ); frankeniae (II); californicus (IV); obovatus (V); phoenicis (VI) and portalis (VII). The species in group III of Baker et al. (1975) were placed in either the obovatus or the portalis species group by Baker & Tuttle (1987). They did not support the synonymy of Cenopalpus with Brevipalpus proposed by Meyer (1979). Furthermore, they divided Tenuipalpus into the caudatus and proteae species groups, and then divided the caudatus group into the annonae, anoplus and bakeri subgroups. 33. Al-Gboory (1987) presented an excellent review on the false spider mite taxonomy, he discussed the phylogeny and ecology of the family, as well addressing the species present in central Iraq. 34. Mohanasundaram (1988) described Tenuilichus from India. 35. Meyer (1993) synonymized Aegyptopalpus and Amblypalpus under Tenuipalpus. She accepted the division of the genus Tenuipalpus into the groups proposed by Baker & Tuttle (1987), and created two new subgroups, pacificus and eriophyoides, in the caudatus group, and divided the proteae group into the keiensis, rhusi and xerocolus subgroups. 36. Smiley & Gerson (1995) described Australopalpus and Lisaepalpus from Australia. 37. Rimando (1996) described Rarosiella (synonymized with the Raoiella herein) from the Philippines. 38 Smiley et al. (1996) described Crossipalpus and Meyeraepalpus and erected the subfamily Tegopalpinae to include Tegopalpus Womersley, Philippipalpus Corpus-Raros and Australopalpus Smiley & Gerson, 1995. 39. Zhang & Liang (1997) provided an illustrated key to the tenuipalpid genera of the world. 40. Welbourn et al. (2003) considered Brevipalpus and Cenopalpus to be distinct and valid genera. 41. Rodrigues et al. (2004) performed molecular analyses of certain species of Tenuipalpidae. This study, along with cladistical analyses by Kane (unpublished thesis, 2003) and Quiros-Gonzalez (unpublished dissertation, 1985) and more detailed morphological observations with the LT-SEM (Welbourn et al. 2003), may help to elucidate the phylogeny of this important family of plant feeding mites. 42. Beard et al. (2005) described the genus Prolixus from Australia. 43. Beard & Gerson (2009) described the genus Acaricis from Australia.

Zoogeography of the Genera and Species of Tenuipalpidae Tenuipalpids have been described from all zoogeographic regions of the world with the exception of Antarctica. Table 1 shows the number of tenuipalpid species of each genus described from each of the following 8 zoogeographic regions - Nearctic (NA) Neotropical (NT); Western Palearctic (WPA); Afrotropical (AF); Eastern Palearctic (EPA); Oriental (O); Australian (AU) and Pacific Islands (PI). The classification used in this document is slightly different from that used by other authors. We considered all records from a given country to belong to a same zoogeographic region, independently of the position of the type locality within the country. Whereas other authors have placed northern Mexico in the Nearctic and the central and southern areas of Mexico in the Neotropical region, herein we place all records from Mexico in the Neotropical region. Similarly, northern China has been placed in the Palearctic and the central and southern A catalog of the Tenuipalpidae of the world with a key to genera

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areas of China in the Oriental region; herein, we place all records from China in the Eastern Palearctic. In addition, we divide the Palearctic region which traditionally covers the entire Eurasian continent into the Western Palearctic and Eastern Palearctic regions which are described in more detail below. Species distribution within each genus is given in Appendix A which shows the type locality (country) of each species within each genus among the zoogeographic regions. The country from which the species was described is in most cases referred to by the first 3 letters of the name of the country. Since the type locality of all but one of the species from the Nearctic region is from the United States, the state of that country is mentioned for the type localities of the corresponding species. The family name for the host plant from which each type specimen was collected is also included in the table corresponding to each genus. A plus sign (+) is used to denote the presence of the species in a zoogeographic region other than that of the type locality. Country and state abbreviations within each zoogeographic region is subsequently provided: 1. Nearctic (NA): Canada (CAN), Greenland (GRE), USA USA States: AK—Alaska; AL—Alabama; AR—Arkansas; AZ—Arizona; CA—California; CO—Colorado; CT—Connecticut; DC— District of Colombia; DE—Delaware; FL—Florida; GA—Georgia; HI—Hawaii; IA—Iowa; ID—Idaho; IL—Illinois; IN—Indiana; KS—Kansas; KY—Kentucky; LA—Louisanna; MA—Massachusetts; MD—Maryland; ME—Maine; MI—Michigan; MN—Minnesota; MO—Missouri;MS—Mississippi; MT—Montana; NC—North Carolina; ND—North Dakota; NE—Nebraska; NH—New Hampshire; NJ—New Jersey; NM—New Mexico; NV—Nevada; NY—New York; OH—Ohio; OK—Oklahoma; OR— Oregon; PA—Pennsylvania; RI—Rhode Island; SC—South Carolina; SD—South Dakota; TN—Tennessee; TX—Texas; UT—Utah; VA—Virginia; VT—Vermont; WA—Washington; WI—Wisconsin; WV—West Virginia; WY—Wyoming. 2. Neotropical (NT): Mexico (MEX), Central and South America and Caribbean. Argentina (ARG), Brazil (BRA), Chile (CHL), Costa Rica (CRi), Cuba (CUB), Dominican Republic (DRe), Honduras (HON), Jamaica (JAM), Panama (PAN), Puerto Rico (PRi). 3. Western Palearctic (WPA): European and Middle Eastern countries, as well as north African countries bordering the Mediterranean Sea, the Arabian Peninsula, Uzbekistasn, Kyrgzstan, Tajikistan, Afghanistan, Turkmenistan and the Azores, Madeira and Canary Islands. Armenia (ARM), Crete (CRE), Egypt (EGY), England (ENG), France (FRA), Georgia (GEO), Germany (GER), Greece (GRE), Iran (IRN), Iraq (IRQ), Israel (ISR), Italy (ITA), Jordan (JOR), Ukraine (UKR). 4. Afrotropical (AF): African countries south of those bordering the Mediterranean Sea. and Malagasian countries -Madagascar, Mauritius, Reunion, Comoros and Indian Ocean athols such as Aldabra Cameroon (CAM), Congo (CON), Ethiopia (ETH), Nigeria (NIG), South Africa (SAf), SUDAN (SUD). 5. Eastern Palearctic (EPA): Far East Asian countries, from China northwards to the Arctic Pole. China (CHN), Japan (JAP), Korea (KOR), Maritime Territory of Russia (MTE), 6. Oriental (O): continental Asian countries south of the Himalaya mountains, including the Indian subcontinent, southeast Asia as well as Austro-oriental countries—Malaysia (MAL), Indonesia (IDO), Papua New Guinea (PNG), Philippines (PHI) and the Solomon Islands (SoI). Bangladesh (BAN), Burma (BUR), Cambodia (CAM), Hong Kong (HoK), India (IND), Java (JAV), Laos (LAO), Nepal (NEP), Pakistan, (PAK), Taiwan (TAI), Thailand (THA), Vietnam (VIE). 7. Australian (AU): includes Australia (AUS), Lord Howe Island (LHI) and Tasmania (TAS). 8. Pacific Islands (PI): tropical Pacific Islands including Hawaiian Islands (HAI), New Caledonia (NCa), and New Zealand (NZ).

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TABLE 1. Number of tenuipalpid species described from each zoogeographic region. Genus Acaricis Aegyptobia Afronychus Amblypalpus Australopalpus Brevipalpus Capedulia Cenopalpus Chaudhripalpus Coleacarus Crossipalpus Dolichotetranychus Krugeria Larvacarus Lisaepalpus Macfarlaniella Magdalenapalpus Obdulia Obuloides Pentamerismus Philippipalpus Phyllotetranychus Phytoptipalpus Priscapalpus Prolixus Pseudoleptus Raoiella Raoiellana Tegopalpus Tenuilichus Tenuipalpus Terminalichus Ultratenuipalpus Urigersonus incertae sedis Total genera Total species

WW 2 93 4 2 1 282 4 60 1 1 2 23 1 1 1 1 1 2 2 17 1 2 17 3 2 10 12 1 1 2 306 6 22 1 3 34 891

NA 0 32 0 0 0 69 0 0 0 0 0 8 0 0 0 0 0 0 0 5 0 0 0 0 0 5 0 0 0 0 19 0 0 0 0 6 138

NT 0 11 0 0 0 129 0 0 0 0 0 0 0 0 0 0 0 0 0 2 0 0 3 2 0 1 0 0 0 0 64 0 5 0 0 8 217

WPA 0 26 0 1 0 23 1 39 0 0 0 3 0 0 0 0 0 2 0 5 0 1 2 0 0 2 1 1 0 0 29 0 0 0 0 14 136

AF 0 8 4 1 0 3 3 3 0 1 0 3 1 0 0 0 0 0 0 3 0 1 7 1 0 1 2 0 0 0 91 0 3 0 0 17 136

EPA 0 4 0 0 0 9 0 1 0 0 0 1 0 0 0 0 0 0 0 2 0 0 2 0 0 0 0 0 0 0 19 0 2 0 0 8 40

O 0 10 0 0 0 49 0 17 0 0 0 5 0 1 0 0 0 0 2 0 1 0 3 0 0 1 8 0 0 2 69 6 2 0 3 15 180

AU 2 1 0 0 1 0 0 0 1 0 2 1 0 0 1 1 1 0 0 0 0 0 0 0 2 0 1 0 1 0 4 0 1 1 0 14 21

PI 0 1 0 0 0 0 0 0 0 0 0 2 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 11 0 9 0 0 4 23

Worldwide (WW); Nearctic (NA) Neotropical (NT); Western Palearctic (WPA); AF (Afrotropical); Eastern Palearctic (EPA); Oriental (O); Australian (AU); Pacific Islands (PI).

Format used in catalog The following format is used to present information on each taxon reported in this work: Genus • Valid name, author, date and page of the corresponding original description.

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• •

Synonyms. Type species. Species • Catalog number. • Valid name of species, author, date and page of the corresponding original description, type locality and host. • Species designation(s): other names used for the species in the literature, each followed by the respective reference of its first citation. • Redescription(s): references of publications providing information on the morphology of the species, other than the original description. • Type depository: the institution where holotype is deposited. Abreviations consistant with the Insect and Spider Collections of the World list were used. This list is maintained by the Bishop Museum, The State Museum of Natural & Cultural History, Honolulu, Hawaii, USA (http://hbs.bishopmuseum.org/codens/ codensearch.html). Where no standard abbreviations could be found or were unknown, the institution was listed as designated in the original description. The following abbreviations were used: ASCU—Agricultural Scientific Collections Unit, Orange, New South Wales, Australia BITU—Biological Institute, Faculty of Education, Tottori University, Japan. BMNH [= NHM]—Natural History Museum, London, England, UK CEA-MHNS—Colecciòn de la Estacion Experimental Agronomica, Universidad de Chile (CEA); Museo de Historia Natural Santiago (MHNS), Chile CMNS—Museum of Natural History, Shanghai, China. CUE—Collection of Faculty of Agriculture, Cairo University, Egypt. DEES—Reference Collection of Departamento de Enotmologia, Fitopatologia e Zoologia Agricola, Escola Superior de Agricultura “Luiz de Queiroz”, Universidade de Sao Paulo, Piracicaba, SP, Brazil. GNHM—Collection of the Goulandris Museum of Natural History, Kiphissia, Athens, Greece. INPC—National Pusa Collection (NPC), Indian Agricultural Research Institute, New Delhi, India. ISZA—Istituto Sperimentale per la Zoologia Agraria di Firenza, Italy MCZ—Museum of Comparative Zoology, Cambridge, Massachusetts,USA. NSMT—National Science Museum (Natural History), Tokyo, Japan. PAUP—Collection of the Department of Zoology, Punjab Agricultural University, Ludhiana, India. QDPI—Queensland Department of Primary Industries Insect Collection, Indooroopilly, Queensland, Australia. QM—The Queensland Museum, South Brisbane, Queensland, Australia. SAM—South Australian Museum, Adelaide, South Australia, Australia. SANC—National Collection of Acari, Plant Protection Research Institute, Department of Agricultural Technical Services, Pretoria, South Africa. TNAU—Department of Agricultural Entomology Collection, Tamil Nadu Agricultural University, Coimbatore, India. TUPH—Acarology Research Section of Plant Pests & Diseases Research Institute, Iran. USNM—United States National Museum of Natural History at Beltsville, Maryland, USA. WINC—Waite Insect and Nematode Collection, Entomology Unit, South Australian Research and Development Institute, Adelaide (EUSARDI), South Australia, Australia.



Note(s): complementary information. Information on junior synonyms of a species is given after the valid name, followed by the reference in which the species was synonymized. The text for the junior synonyms is indented for easy recognition; each species is given the same catalog number as the corresponding senior synonym, followed by a sequential letter. An attempt was made to correct typographical errors in the scientific names of the species and/or collection localities. In addition, we have included the names of the plant families for each host, except for species where the plant family could not be determined. The correct spelling of scientific names of hosts and the determination of their respective families was done using “The International Plant Names Index” (IPNI) and “PLANTS National Database and Topics” available on-line at http://www.ipni.org/index.html and http:// plants.usda.gov/index.html, respectively.

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Family TENUIPALPIDAE Berlese Tenuipalpini Berlese, 1913: 17. Phytoptipalpidae Ewing, 1922: 107. Pseudoleptidae Oudemans, 1928b: 287. Tenuipalpidae Sayed, 1950a: 1015.

Aegyptobia Sayed 1950 Aegyptobia Sayed, 1950b: 1018; Type species: Aegyptobia tragardhi Sayed, 1950b: 1019. Pritchard & Baker, 1958: 179; Meyer & Van Dis, 1993: 310; Mitrofanov, 1973a: 508. Aegyptobia (Aegyptobia) Mitrofanov, 1973a: 508; synonymy according to Meyer 1979: 117. Aegyptobia (Aegyptobiosis) Mitrofanov, 1973a: 508; Type species: Pentamerismus macswaini Pritchard & Baker, 1952: 8; synonymy according to Meyer 1979: 117. Meyeraepalpus Smiley, Frost and Gerson 1996: 176. Type species: Meyeraepalpus delfinadae Smiley, Frost and Gerson; new synonym. Discussion: Smiley et al. (1996) distinguished the genus Meyeraepalpus from the genus Aegyptobia based upon the presence of an overhanging prodorsum, of which the anterior margin is invaginated, and by the absence of a rostral shield in the former. Our examination of the two species that they described and placed in the genus revealed that the type species, Meyeraepalpus delfinadae, has a rostral shield and the anterior margin of the prodorsum is not invaginated. However, these characters are apparent in Meyeraepalpus strandtmanni, the other species that they described in this genus. Since the characters found in the type species of the genus Meyeraepalpus do not differ significantly from those of the genus Aegyptobia, we are proposing that the genus Meyeraepalpus be considered a junior synonym of the genus Aegyptobia and that Meyeraepalpus strandtmanni be placed in its own genus based upon the characters that Smiley et al. used to define the genus Meyeraepalpus. However, since the type species of the genus belongs to the genus Aegyptgobia, the name Meyeraepalpus is no longer available. Therefore, we are proposing a new genus and combination for this species, Magdalenapalpus strandtmanni (Smiley, Frost and Gerson), in honor of Magdalena K. P. Smith-Meyer. 1. Aegyptobia abuzabiensis Meyer & Van Dis, 1993: 314; Abu-Zabi (Abu-Dhabi), United Arab Emirates, ex Cornulaca monacantha Delile (Chenopodiaceae). Type Depository: SANC. 2. Aegyptobia acacia Baker & Tuttle, 1972: 25; Quartzsite, Arizona, USA, ex Acacia greggii Gray (Mimosaceae). Species designation: Aegyptobia (Aegyptobia) acacia—Mitrofanov & Strunkova (1979). Redescriptions: Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: USNM, No. 3428. 3. Aegyptobia aletes (Pritchard & Baker, 1952): 9; Hume Lake, California, USA, ex Libocedrus decurrens Torrey (Cupressaceae). Species designations: Pentamerismus aletes—original designation, Aegyptobia aletes—Pritchard & Baker (1958), Aegyptobia (Aegyptobia) aletes—Mitrofanov et al (1975). Redescriptions: Baker & Pritchard (1953c), Gutierrez et al. (1989), Meyer (1979), Mitrofanov et al. (1975), Mitrofanov & Strunkova (1979), Pritchard & Baker (1958). Type Depository: USNM, No. 1907. 4. Aegyptobia aliartensis Hatzinikolis, 1987: 51; Aliartos, Viotia, Greece, ex Thuja sp. (Cupressaceae). Type Depository: Collection of the Acarology Laboratory of Agricultural Research Centre of Athens, Greece. 5. Aegyptobia allionia Baker, Tuttle & Abbatiello, 1975: 4; 16 km S Chiahuaha, Mexico, ex Allionia incarnata L. (Nyctaginaceae). Species designation: Aegyptobia allioniae [sic]—Baker & Tuttle (1987). Redescriptions: Baker & Tuttle (1987), Meyer (1979). Type Depository: USNM, No. 3622. 6. Aegyptobia alpinensis Baker & Tuttle, 1972: 22; Alpine, California, USA, ex Phoradendron bolleanum (Seemann) Eichler (Viscaceae).

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Redescription: Meyer (1979). Type Depository: USNM, No. 3420. 7. Aegyptobia ambrosiae Baker & Tuttle, 1987: 12; Culiacan, Sinaloa, Mexico, ex Ambrosia confertiflora (DC) Rydb. (Asteraceae). Type Depository: USNM. 8. Aegyptobia antenostoma Baker & Tuttle, 1972: 22; Alpine, California, USA, ex Adenostoma fasciculatum Hooker & Arnot (Rosaceae). Species designation: Aegyptobia (Aegyptobia) antenostomae [sic]—Mitrofanov & Strunkova (1979). Redescriptions: Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: USNM, No. 3422. Note: host originally listed as Antenostoma fasciculatum Hooker & Arnott. 9. Aegyptobia aplopappi Baker & Tuttle, 1964: 9; Arizona, USA, ex Haplopappus spinulosus (Pursh) DC. and H. acradenius (Greene) Blake (Asteraceae). Species designation: Aegyptobia (Aegyptobia) aplopappi—Mitrofanov & Strunkova (1979). Redescriptions: Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: USNM, No. 3016. Note: host originally listed as Aplopappus spinulosus (Pursh) DC. 10. Aegyptobia arbutusae Baker & Tuttle, 1987: 12; 152 km N Matehuala, San Luis Potosi, Mexico, ex Arbutus sp. (Ericaceae). Type Depository: USNM. 11. Aegyptobia arenaria Ehara, 1982: 175; Tottori sand dune, Tottori, Honshu, Japan, ex Artemisia capillaris Thumb. (Asteraceae). Type Depository: BITU. 12. Aegyptobia arizonensis Baker & Tuttle, 1964: 29; Dome Valley, Yuma, Arizona, USA, ex Atriplex canescens (Pursh) Nutt. (Chenopodiaceae). Species designation: Aegyptobia (Aegyptobia) arisonensis [sic]—Mitrofanov & Strunkova (1979). Redescriptions: Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: USNM, No. 3022. 13. Aegyptobia baccharis Baker & Tuttle, 1972: 21; Tucson, Arizona, USA, ex Baccharis glutinosa Persoon (Asteraceae). Species designation: Aegyptobia (Aegyptobia) baccharis—Mitrofanov & Strunkova (1979). Redescriptions: Mitrofanov & Strunkova (1979). Type Depository: USNM, No. 3419. 14. Aegyptobia baptus (Pritchard & Baker, 1952): 10; Tesla, California, USA, ex Hemizonia virgata Gray (Asteraceae). Species designations: Pentamerismus baptus—original designation, Aegyptobia baptus—Pritchard & Baker (1958), Aegyptobia (Aegyptobia) baptus—Mitrofanov & Strunkova (1979). Redescriptions: Baker & Pritchard (1953c), Baker & Tuttle (1964, 1987), Meyer (1979), Mitrofanov & Strunkova (1979), Pritchard & Baker (1958). Type Depository: USNM, No. 1906. 15. Aegyptobia beglarovi Livschitz & Mitrofanov, 1967: 40; Ukraine, ex Juniperus oxycedrus L. and Juniperus communis L. var. hibernica (Cupressaceae). Species designation: Aegyptobia (Aegyptobia) beglarovi—Mitrofanov et al. (1975). Redescriptions: Meyer (1979), Mitrofanov et al. (1975), Mitrofanov & Strunkova (1979). Type Depository: Unknown. 16. Aegyptobia bibbyi Baker & Tuttle, 1964: 27; Yuma, Arizona, USA, ex Atriplex canescens (Pursh) Nutt. (Chenopodiaceae). Species designation: Aegyptobia (Aegyptobia) bibbyi—Mitrofanov & Strunkova (1979). Redescriptions: Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: USNM, No. 3024.

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17. Aegyptobia cactaceae Baker & Tuttle, 1987: 14; Sonora, Mexico, intercepted at Nogales, Arizona quarantine station, Arizona, USA, ex cactus debris (Cactaceae). Type Depository: USNM. 18. Aegyptobia cactorum Baker & Tuttle, 1987: 9; Baja California, Mexico, intercepted at San Ysidro, California quarantine station, California, USA, ex an unidentified cactus (Cactaceae). Type Depository: USNM. 19. Aegyptobia campsis Baker & Tuttle, 1964: 17; Yuma, Arizona, USA, ex Campsis radicans (L.) Seem. ex Bureau (Bignoniaceae). Species designation: Aegyptobia (Aegyptobia) campis [sic]—Mitrofanov & Strunkova (1979). Redescriptions: Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: USNM, No. 3017. 20. Aegyptobia cassiae Baker & Tuttle, 1964: 12; Sells, Arizona, USA, ex Cassia covesii Gray (Caesalpineaceae). Species designation: Aegyptobia (Aegyptobia) cassiae—Mitrofanov & Strunkova (1979). Redescriptions: Baker & Tuttle (1987), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: USNM, No. 3018. 21. Aegyptobia cedermontana Meyer & Van Dis, 1993: 312; Dwars River, Ceder Mountains, Cape Province, South Africa, ex Anthospermum aethiopicum L. (Rubiaceae). Type Depository: SANC. 22. Aegyptobia changi Tseng, 1980: 153; Taiwan, ex Juniperus chinensis L. (Cupressaceae). Type Depository: Unknown. 23. Aegyptobia crotonae Baker & Tuttle, 1972: 21; Portal, Arizona, USA, ex Croton corymbulosus Engelm. ex C.F. Wheeler (Euphorbiaceae). Species designation: Aegyptobia (Aegyptobia) crotonae— Mitrofanov & Strunkova (1979). Redescriptions: Baker & Tuttle (1987), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: USNM, No. 3418. 24. Aegyptobia cupressus Baker & Tuttle, 1972: 18; Alpine, California, USA, ex Cupressus forbesii Jepson (Cupressaceae). Species designation: Aegyptobia (Aegyptobia) cupressus—Mitrofanov et al. (1975). Redescriptions: Meyer (1979), Mitrofanov et al. (1975), Mitrofanov & Strunkova (1979). Type Depository: USNM, No. 3416. 25. Aegyptobia curtipilis Baker & Tuttle, 1987: 9; 67.3 km east of Huauapan, Oaxaca, Mexico, ex Salvia sp. (Lamiaceae). Type Depository: USNM. 26. Aegyptobia daneshvari Parsi & Khosrowshahi, 1990: 38; Tehran Province, Iran, ex Populus sp. (Salicaceae). Redescription: Khosrowshahi & Arbabi (1997). Type Depository: TUPH. 27. Aegyptobia delfinadae (Smiley, Frost & Gerson), 1996: 178; —new combination; Saint Mary's Peak, Flinders Ranges, South Australia, Australia, ex Allocasuarina muelleriana (Miq.) L.A.S. Johnson (Casuarinaceae). Species designation: Meyeraepalpus delfinadae Smiley, Frost & Gerson, 1996: 178. Type Depository: WINC. Note: host originally listed as Allocasuariana muelleriana. 28. Aegyptobia desertorum Baker & Tuttle, 1964: 23; Dome Valley, Yuma County, Arizona, USA, ex Atriplex canescens (Pursh) Nutt. (Chenopodiaceae). Species designation: Aegyptobia (Aegyptobia) desertorum—Mitrofanov & Strunkova (1979). Redescriptions: Baker & Tuttle (1972, 1987), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: USNM, No. 3023.

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29. Aegyptobia edenvillensis Meyer, 1979: 119; Edenville, Orange Free State, South Africa, ex grass (Poaceae) and soil. Redescription: Meyer & Van Dis (1993). Type Depository: SANC. 30. Aegyptobia ephedrae Baker & Tuttle, 1964: 6; Yuma, Arizona, USA, ex Ephedra fasciculata A. Nels. (Ephedraceae). Species designation: Aegyptobia (Aegyptobiopsis) ephedrae—Mitrofanov & Strunkova (1979). Redescriptions: Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: USNM, No. 3015. 31. Aegyptobia eremia Meyer & Gerson, 1980: 77; Yeroham, Israel, ex Hammada scoparia (Pomel) (Chenopodiaceae). Redescription: Meyer & Van Dis (1993). Type Depository: Collection of the Department of Entomology, Faculty of Agriculture, Rehovot, Israel. Note: host originally listed as Hammada scoporia. 32. Aegyptobia ericae Meyer & Van Dis, 1993: 313; Dwars River, Ceder Mountains, Cape Province, South Africa, ex Erica walkeria Andr. (Ericaceae). Type Depository: SANC. 33. Aegyptobia eriogonum Baker & Tuttle, 1972: 24; Portal, Arizona, USA, ex Eriogonum wrightii Torrey ex Benth. (Polygonaceae). Redescriptions: Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: USNM, No. 3426. 34. Aegyptobia euphratica Al-Gboory, 1987: 109; Abu-Ghraib, Baghdad, Iraq, ex Cupressus sempervirens L. (Cupressaceae). Type Depository: Entomology Museum, College of Agriculture, Department of Plant Protection, Abu-Ghraib, Baghdad, Iraq. 35. Aegyptobia exarata Livschitz & Mitrofanov, 1967: 42; Ukraine, ex Artemisia sp. (Asteraceae). Species designation: Aegyptobia (Aegyptobia) exarata—Mitrofanov & Strunkova (1979). Redescriptions: Meyer (1979), Mitrofanov & Strunkova (1979), Strunkova (1978). Type Depository: Unknown. 36. Aegyptobia fallugia Baker & Tuttle, 1972: 20; Portal, Arizona, USA, ex Fallugia paradoxa (D. Don) Endlicher ex Torr. (Rosaceae). Species designation: Aegyptobia (Aegyptobiosis) fallugia—Mitrofanov & Strunkova (1979). Redescriptions: Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: USNM, No. 3417. 37. Aegyptobia flourensia Baker & Tuttle, 1972: 22; Portal, Arizona, USA, ex Flourensia cernua DC. (Asteraceae). Species designation: Aegyptobia (Aegyptobia) flourensiae [sic]—Mitrofanov & Strunkova (1979). Redescriptions: Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: USNM, No. 3423. 38. Aegyptobia forma Chaudhri, 1972a: 20; Hasan Abdal, Pakistan, ex Artemisia scoparia Waldst. & Kit. (Asteraceae). Redescriptions: Akbar & Mushtaq (1993), Chaudhri & Akbar (1985), Chaudhri et al. (1974), Hasan & Akbar (2002), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: Department of Entomology, West Pakistan Agricultural University Lyallpur, Pakistan. 39. Aegyptobia franseriae Baker & Tuttle, 1964: 18; Gila Mountains, Yuma County, Arizona, USA, ex Ambrosia ilicifolia (Gray) Payne (Asteraceae). Species designation: Aegyptobia (Aegyptobia) franseriae— Mitrofanov & Strunkova (1979). Redescriptions: Baker & Tuttle (1972), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: USNM, No. 3020. Note: host originally listed as Franseria ilicifolia Gray. 40. Aegyptobia glyptus Pritchard & Baker, 1958: 180; Guaymas, Sonora, Mexico intercepted at Nogales, Arizona, USA, ex Ferocactus sp. (Cactaceae). Species designation: Aegyptobia (Aegyptobiosis) glyptus—Mitrofanov & Strunkova (1979).

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Redescriptions: Baker & Tuttle (1987), De Leon (1962), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: USNM, No. 2293. 41. Aegyptobia hamus Chaudhri, 1972a: 20; Karachi, Sind, Pakistan, ex Heliotropium sp. (Boraginaceae). Redescriptions: Akbar & Mushtaq (1993), Chaudhri & Akbar (1985), Chaudhri et al. (1974), Hasan & Akbar (2002), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: Department of Entomology, West Pakistan Agricultural University Lyallpur, Pakistan. 42. Aegyptobia haplopappus Baker & Tuttle, 1972: 25; Pinedale, Arizona, USA, ex Haplopappus spinulosus (Pursh) DC. (Asteraceae). Species designation: Aegyptobia (Aegyptobia) haplopappus—Mitrofanov & Strunkova (1979). Redescriptions: Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: USNM, No. 3427. 43. Aegyptobia hefeiensis Ma & Yuan, 1981: 194; Hefei, China, ex Biota orientalis Endl. (Cupressaceae). Type Depository: SCAC. 44. Aegyptobia hymenocleae Baker & Tuttle, 1964: 25; Casa Grande, Gila Bend, Arizona, USA, ex Hymenoclea pentalepis Rydb. (Asteraceae). Species designation: Aegyptobia (Aegyptobia) hymenocleae—Mitrofanov & Strunkova (1979). Redescriptions: Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: USNM, No. 3021. 45. Aegyptobia incarnata Baker, Tuttle & Abbatiello, 1975: 4; Hermosillo, Sonora, México, ex Allionia incarnata L. (Nyctaginaceae). Species designation: Aegyptobia incarnatae [sic]— Baker & Tuttle (1987). Redescriptions: Baker & Tuttle (1987), Meyer (1979). Type Depository: USNM, No. 3623. 46. Aegyptobia karystensis Hatzinikolis, 1987: 50; Karystos, Evia, Greece, ex Cupressus sp. (Cupressaceae). Type Depository: ARCAG. 47. Aegyptobia kerkyrae Hatzinikolis & Papadoulis, 1997: 139; Ermones, Corfu Island, Eptanissa, Greece ex an unidentified herbaceous plant. Type Depository: BMNH. 48. Aegyptobia kharazii Mesa & Moraes, 2007; new replacement name for Aegyptobia meyerae Khosrowshahi & Arbabi (1997), name preoccuped by Aegyptobia meyerae Hatzinikolis & Panou (1996). (Article 58.7 4th Edition International Code of Zoological Nomenclature). The new species name is in honor of Dr. Kharazii, Prof. Entomology, Department of Plant Protection, College of Agriculture, Tehran University. Homonym: 48a. Aegyptobia meyerae Khosrowshahi & Arbabi, 1997: 10; Tehran Province, Iran, ex Juniperus communis L. (Cupressaceae). Type Depository: TUPH, No. T-1322. 49. Aegyptobia lacida Baker & Tuttle, 1972: 22; Prescott, Arizona, USA, ex Gutierrezia lucida Greene (Asteraceae). Species designation: Aegyptobia (Aegyptobia) lacida—Mitrofanov & Strunkova (1979). Redescriptions: Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: USNM, No. 3421. Note: host originally listed as Gutierrezia lacida Greene. 50. Aegyptobia leiahensis Chaudhri, Akbar & Rasool, 1974: 9; 3.2 km E Leiah, Punjab, Pakistan, ex Heliotropium sp. (Boraginaceae). Redescriptions: Akbar & Mushtaq (1993), Chaudhri & Akbar (1985), Hasan & Akbar (2002), Hatzinikolis (1987). Type Depository: Department of Entomology, University of Agriculture, Lyallpur, Pakistan. 51. Aegyptobia lineati Meyer & Van Dis, 1993: 313; Karoo National Park, near Beaufort-West, Cape Province, South Africa, ex Thesium lineatum L. (Santalaceae). Type Depository: SANC.

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52. Aegyptobia macswaini (Pritchard & Baker, 1952): 8; Tesla, California, USA, ex Gutierrezia californica (DC.) Torr. & Gray (Asteraceae). Species designations: Pentamerismus macswaini—original designation, Aegyptobia macswaini—Pritchard & Baker (1958), Aegyptobia (Aegyptobiosis) macswaini—Mitrofanov & Strunkova (1979). Redescriptions: Baker & Pritchard (1953c), Baker & Tuttle (1964, 1972, 1987), De Leon (1962), Hasan & Akbar (2002), Meyer (1979), Mitrofanov & Strunkova (1979), Pritchard & Baker (1958). Type Depository: USNM, No. 1909. 53. Aegyptobia mccormicki (Baker & Pritchard, 1953c): 363; Yakima, Washington, USA, ex arborvitae (Cupressaceae). Species designations: Pentamerismus mccormicki—original designation, Aegyptobia mccormicki—Pritchard & Baker (1958), Aegyptobia (Aegyptobia) mccormiki [sic]—Mitrofanov & Strunkova (1979). Redescriptions: Meyer (1979), Mitrofanov et al. (1975), Mitrofanov & Strunkova (1979), Pritchard & Baker (1958). Type Depository: USNM, No. No. 1987. 54. Aegyptobia menoni Maninder & Ghai, 1980: 311; Indian Agricultural Research Institute, New Delhi, India, ex Cupressus sp. (Cupressaceae). Redescriptions: Sadana (1985, 1997). Type Depository: INPC. 55. Aegyptobia mesopotamiensis Al-Gboory, 1987: 113; Abu-Ghraib, Baghdad, Iraq, ex Cupressus sp. (Cupressaceae). Type Depository: Entomology Museum, College of Agriculture, Department of Plant Protection, Abu-Ghraib, Baghdad, Iraq. 56. Aegyptobia meyerae Hatzinikolis & Panou, 1996b: 221; Oeti Mountain, Phthiotis, Greece, ex an unidentified herbaceous plant. Type Depository: BMNH. Note: Aegyptobia meyerae Khosrowshahi & Arbabi, 1997 has been replaced by A. kharazii Mesa & Moraes, 2007. 57. Aegyptobia monacanthae Meyer & Van Dis, 1993: 316; Abu-Zabi (Abu-Dhabi), United Arab Emirates, ex Cornulaca monacantha Delile (Chenopodiaceae). Type Depository: SANC. 58. Aegyptobia monitus Chaudhri, 1972a: 19; Karachi, Sind, Pakistan, ex Medicago sp. (Fabaceae). Redescriptions: Akbar & Mushtaq (1993), Chaudhri & Akbar (1985) Chaudhri et al. (1974), Hasan & Akbar (2002), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: Department of Entomology, West Pakistan Agricultural University Lyallpur, Pakistan. 59. Aegyptobia montana Bury & Krantz, 1977: 417; south side of Mount Hood, Clackamas, Oregon, USA, ex Penstemon davidsonii Greene (Scrophulariaceae). Type Depository: USNM. 60. Aegyptobia nasicornensis Meyer & Van Dis, 1993: 317; Pofadder, Cape Province, South Africa, ex Aptosimum spinescens (Thunb.) F. Weber (Scrophulariaceae). Type Depository: SANC. 61. Aegyptobia neobaptus Meyer, 1979: 125; Naudesfontein, Cape Province, South Africa, ex Populus alba L. (Salicaceae). Species designation: Aegyptobia neobapta [sic]—Meyer & Van Dis (1993). Redescriptions: Meyer & Ueckermann (1989), Meyer & Van Dis (1993). Type Depository: SANC. 62. Aegyptobia nina (Pritchard & Baker, 1952): 9; Hume Lake, California, USA, ex Libocedrus decurrens Torrey (Cupressaceae). Species designations: Pentamerismus nina—original designation, Aegyptobia nina Pritchard & Baker (1958), Aegyptobia (Aegyptobia) nina—Mitrofanov & Strunkova (1979). Redescriptions: Baker & Pritchard (1953c), Meyer (1979), Mitrofanov et al. (1975), Mitrofanov & Strunkova (1979), Pritchard & Baker (1958). Type Depository: USNM, No. 1908.

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63. Aegyptobia nomus (Baker & Pritchard, 1953c): 361; Mandan, North Dakota, USA, ex Bouteloua gracilis (Kunth) Lagasca ex. Griffiths (Poaceae). Species designations: Pentamerismus nomus—original designation, Aegyptobia nomus—Pritchard & Baker (1958), Aegyptobia (Aegyptobiopsis) nomus—Mitrofanov & Strunkova (1979). Redescriptions: Meyer (1979), Mitrofanov & Strunkova (1979), Pritchard & Baker (1958). Type Depository: USNM, No. 1988. Note: host originally listed as Bouteloa gracilis. 64. Aegyptobia nothus Pritchard & Baker, 1958: 182; Oklahoma City, Oklahoma, USA, ex Juniperus sp. (Cupressaceae). Species designation: Aegyptobia (Aegyptobia) nothus—Mitrofanov & Strunkova (1979). Redescriptions: De Leon (1961b), Evans et al. (1998), Meyer (1979), Mitrofanov et al. (1975), Mitrofanov & Strunkova (1979), Thewke & Enns (1970). Type Depository: USNM, No. 2207. Note: Also found in Bermuda (Evans et al. 1993). 65. Aegyptobia nummulus Chaudhri, 1972a: 21; Samli, Murree Hills, Pakistan, ex Cupressus sempervirens L. (Cupressaceae). Redescriptions: Akbar & Mushtaq (1993), Chaudhri & Akbar (1985), Chaudhri et al. (1974), Hatzinikolis & Panou (1996), Maninder & Ghai (1980), Hasan & Akbar (2002), Meyer (1979), Mitrofanov & Strunkova (1979), Sadana (1985, 1997). Type Depository: Department of Entomology, West Pakistan Agricultural University Lyallpur, Pakistan. Note: host originally listed Cupressus semipervirens L. 66. Aegyptobia odontopilis Meyer, in Meyer & Ueckermann, 1988: 19; Rooiputs, Kalahari Gemsbok National Park, Cape Province, South Africa, ex Aptosimum marlothii (Engler) Hiern (Scrophulariaceae). Redescription: Meyer & Van Dis (1993). Type Depository: SANC. 67. Aegyptobia parcus Akbar & Mushtaq, 1993: 153, Kabirwala, Pakistan, ex Mangifera indica L. (Anacardiaceae). Redescription: Hasan & Akbar (2002). Type Depository: Acarology Research Laboratory, Department of Agric. Entomology, University of Agriculture, Faisalabad, Pakistan. 68. Aegyptobia pavlovskii (Reck, 1951): 292; Georgia, ex Ephedra procera Fisch. & C.A. Mey. (Ephedraceae). Species designations: Brevipalpoides pavlovskii—original designation, Aegyptobia pavlovskii—Pritchard & Baker (1958), Aegyptobia (Aegyptobia) pavlovskii— Mitrofanov & Strunkova (1979). Redescriptions: Bagdasarian (1962), Livschitz & Mitrofanov (1967), Meyer (1979), Mitrofanov & Strunkova (1979), Pritchard & Baker (1958), Strunkova (1978). Type Depository: Unknown. 69. Aegyptobia pennatulae Baker & Tuttle, 1987: 16; Chapala, Jalisco, Mexico, ex Acacia pennatula (Schltdl. & Cham.) Benth. (Mimosaceae). Type Depository: USNM. 70. Aegyptobia perscia Khosrowshahi & Arbabi, 1997: 9; Tehran province, Iran, ex Plantago lanceolata L. (Plantaginaceae). Type Depository: TUPH, No. T-1222. 71. Aegyptobia physalis Baker & Tuttle, 1972: 23; Marana, Arizona, USA, ex Physalis wrightii A. Gray (Solanaceae). Species designation: Aegyptobia (Aegyptobia) physalis—Mitrofanov & Strunkova (1979). Redescriptions: Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: USNM, No. 3425. 72. Aegyptobia pomaderrisae Collyer, 1969: 815; Orere, Auckland Province, New Zealand, ex Pomaderris rugosa Cheeseman (Rhamnaceae). Species designation: Aegyptobia (Aegyptobia) pomaderrisae—Mitrofanov & Strunkova (1979). Redescriptions: Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: Collection of Entomology Division, Nelson, New Zealand.

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73. Aegyptobia populus Papaioannou-Souliotis, 1986: 24; Kiphissia, Attica, Greece, ex Populus sp. (Salicaceae). Type Depository: Acarology Laboratory at the Benaki Phytopathological Institute, Greece. 74. Aegyptobia prolixa Meyer, 1979: 119; Ngotsche, near Pongola, Natal, South Africa, ex grass (Poaceae) and soil. Redescription: Meyer & Van Dis (1993). Type Depository: SANC. 75. Aegyptobia pseudoleptoides (Baker & Pritchard, 1953c): 357; Modena, Utah, USA, ex Bouteloua gracilis (Kunth) Lagasca ex. Griffiths (Poaceae). Species designations: Pentamerismus pseudoleptoides—original designation, Aegyptobia pseudoleptoides—Pritchard & Baker (1958), Aegyptobia (Aegyptobiopsis) pseudoleptoides—Mitrofanov & Strunkova (1979). Redescriptions: Meyer (1979), Mitrofanov & Strunkova (1979), Pritchard & Baker (1958). Type Depository: USNM, No. 1986. Note: host originally listed as Bouteloa gracilis. 76. Aegyptobia pyramidi El-Enany & Soliman, 1987: 4; between Giza and Fayoum, Cairo, Egypt, ex a desert plant. Redescription: Meyer & Van Dis (1993), Zaher et al. (1984). Type Depository: CUE. 77. Aegyptobia salicicola Al-Gboory, 1987: 106; Abu-Ghraib, Baghdad, Iraq, ex Salix alba L. (Salicaceae). Type Depository: Entomology Museum, College of Agriculture, Department of Plant Protection, Abu-Ghraib, Baghdad, Iraq. 78. Aegyptobia salixi Zaher & Yousef, 1969: 273; Cairo, Giza, El-Sharkia, El-Kalubia and El-Menia, Egypt, ex Salix sp. (Salicaceae). Redescriptions: El-Enany & Soliman (1987), Meyer (1979), Meyer & Van Dis (1993), Yousef (1971), Zaher et al. (1984). Type Depository: CUE. 79. Aegyptobia salubris Chaudhri, 1972a: 22; 3.2 km E Multan, Pakistan, ex Cupressaceae. Species designation: Aegyptobia salubris [sic]—Sadana (1997). Redescriptions: Akbar & Mushtaq (1993), Chaudhri & Akbar (1985), Chaudhri et al. (1974), Hasan & Akbar (2002), Maninder & Ghai (1980), Meyer (1979), Mitrofanov & Strunkova (1979), Sadana (1985, 1997). Type Depository: Department of Entomology, West Pakistan Agricultural University Lyallpur, Pakistan. Note: host originally listed as Cupressa sp. 80. Aegyptobia sayedi Yousef, 1971: 135; Giza and Gaza Strip, Egypt, ex Cupressus sempervirens L. (Cupressaceae). Redescriptions: El-Enany & Soliman (1987), Meyer (1979), Meyer & Van Dis (1993), Mitrofanov & Strunkova (1979), Papaioannou—Souliotis (1986), Zaher et al. (1984). Type Depository: Collection of the National Research Centre, Egypt. 81. Aegyptobia semper Chaudhri & Akbar, 1985: 10; Garam Chashma, Chitral Valley, Pakistan, ex an unidentified plant. Redescriptions: Hasan & Akbar (2002). Type Depository: Department Entomology, University of Agriculture, Faisalabad, Pakistan. Note: suspected junior synonym Brevipalpus phoenicis (Geijskes). Synonym: 81a. Aegyptobia semper Akbar & Mushtaq, 1993. 155; Garam Chashma, Chitral Valley, Pakistan, ex an unidentified plant—new synonym. Note: We consider these species to be the same species, described with the same species name, but different authors names. The drawings of the original description of Aegyptobia semper Akbar & Mushtaq (1993) are exactly the same as those of the original description of Aegyptobia simper Chaudhri & Akbar (1985), the locality where the species was collected is similar, and the collector and dates of collection are the same. Redescription: Hasan & Akbar (2002). Type Depository: Acarology Research Laboratory, Department of Agric. Entomology, University of Agriculture, Faisalabad, Pakistan. Note: suspected junior synonym Brevipalpus phoenicis (Geijskes). 82. Aegyptobia sohanraensis Hasan & Akbar, 2002: 30; Lal Sohanra Forest, Bahawalpur, Pakistan, ex an unidentified plant.

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Type Depository: Acarology Research Laboratory, Department of Agric. Entomology, University of Agriculture, Faisalabad, Pakistan. 83. Aegyptobia solanum Baker & Tuttle, 1987: 17; Alamos, Sonora, Mexico, ex Solanum elaeagnifolium Cav. (Solanaceae). Type Depository: USNM. 84. Aegyptobia thujae Baker & Tuttle, 1964: 13; Yuma, Arizona, USA, ex Thuja occidentalis L. (Cupressaceae). Species designation: Aegyptobia (Aegyptobia) thujae—Mitrofanov & Strunkova (1979). Redescriptions: Baker & Tuttle (1972), Meyer (1979), Mitrofanov et al. (1975), Mitrofanov & Strunkova (1979). Type Depository: USNM, No. 3019. 85. Aegyptobia torreyi Baker & Tuttle, 1972: 23; McNary, Arizona, USA, ex Muhlenbergia torreyi (Kunth) A.S. Hitchcock & Bush (Poaceae). Species designation: Aegyptobia (Aegyptobiopsis) torreyi—Mitrofanov & Strunkova (1979). Redescriptions: Meyer (1979), Mitrofanov et al. (1975), Mitrofanov & Strunkova (1979). Type Depository: USNM, No. 3424. 86. Aegyptobia tragardhi Sayed, 1950b: 1019; Dokki, Giza, Egypt, ex Thuja orientalis L. (Cupressaceae). Species designations: Pentamerismus tragardhi—Baker & Pritchard (1953c), Aegyptobia (Aegyptobia) tragardhi—Mitrofanov et al. 1975). Redescriptions: Al-Gboory (1987), Baker & Pritchard (1953c), El-Enany & Soliman (1987), Gutierrez et al. (1989), Hasan & Akbar (2002), Livschitz & Mitrofanov (1967), Meyer (1979), Meyer & Van Dis (1993), Mitrofanov et al. (1975), Mitrofanov & Strunkova (1979), Pritchard & Baker (1958), Zaher & Yousef (1969), Zaher et al. (1984). Type Depository: Unknown. 87. Aegyptobia ueckermanni Khosrowshahi & Arbabi, 1997: 8; Tehran Province, Iran, ex Cupressus sp. (Cupressaceae). Type Depository: TUPH, No. T-1122. 88. Aegyptobia vannus Pritchard & Baker, 1958: 183; Mexico, intercepted at Nogales, Arizona, USA, ex Prosopis glandulosa Torr. (Mimosaceae). Species designations: Aegyptobia vanus [sic]—De Leon (1962), Aegyptobia (Aegyptobia) vannus—Mitrofanov & Strunkova (1979). Redescriptions: Baker & Tuttle (1964, 1987), De Leon (1962), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: USNM, No. 2294. 89. Aegyptobia villiensis Hatzinikolis & Emmanouel, 1990: 21; Villia Attiki, Greece, ex Pyrus amygdaliformis Vill. (Rosaceae). Type Depository: Collection of the Acarology Laboratory of Agricultural Research Centre of Athens, Greece. 90. Aegyptobia wainsteini (Bagdasarian, 1962): 49; Armenia, ex Juniperus sp. (Cupressaceae). Species designations: Pentamerismus wainsteini—original designation, Aegyptobia wainsteini Livschitz & Mitrofanov (1967), Aegyptobia (Aegyptobia) wainsteini—Mitrofanov & Strunkova (1979). Redescriptions: Livschitz & Mitrofanov (1967), Meyer (1979), Mitrofanov et al. (1975), Mitrofanov & Strunkova (1979). Type Depository: Unknown. 91. Aegyptobia xerophilus (Reck, 1953): 179; Georgia, ex Acantholimon lepturoides Boiss. (Plumbaginaceae) and Cerastium argenteum M. Bieb. (Caryophyllaceae). Species designations: Brevipalpoides xerophilus—original designation, Aegyptobia xerophilus—Pritchard & Baker (1958), Aegyptobia (Aegyptobia) xerophilus— Mitrofanov & Strunkova (1979). Redescriptions: Livschitz & Mitrofanov (1967), Meyer (1979), Mitrofanov & Strunkova (1979), Pritchard & Baker (1958). Type Depository: Unknown. 92. Aegyptobia xinjiangensis Li & Ma, 1982: 23; Aertai, Shihezi, Xianjiang, China, ex Kochia prostrata (L.) Schrad var. villosissima Bong. & Mev. (Chenopodiaceae). Type Depository: Shanghai College of Agriculture, China.

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93. Aegyptobia zaitzevi (Reck, 1951): 292; Georgia, ex Atraphaxis spinosa L. (Polygonaceae). Species designations: Brevipalpoides zaitzevi—original designation, Aegyptobia zaitzevi (1958), Pentamerismus zaitzevi—Bagdasarian (1962), Aegyptobia (Aegyptobia) zaitzevi—Mitrofanov & Strunkova (1979). Redescriptions: Bagdasarian (1962), Livschitz & Mitrofanov (1967), Meyer (1979), Mitrofanov & Strunkova (1979), Pritchard & Baker (1958). Type Depository: Unknown.

Afronychus Meyer 1979 Afronychus Meyer 1979: 103. Type species: Afronychus amnicus Meyer, 1979. 94. Afronychus amnicus Meyer, 1979: 105; Riviersonderend, Cape Province, South Africa, ex an unidentified wild plant. Type Depository: SANC. 95. Afronychus cliffortiae Meyer, 1979: 103; Mossel Bay, Cape Province, South Africa, ex Cliffortia ramosissima Schlecht (Rosaceae). Type Depository: SANC. 96. Afronychus diastellae Meyer, 1979: 105; Cape Point Nature Reserve, Cape Province, South Africa, ex Diastella divaricata (Berg.) J.P. Rourke (Proteaceae). Type Depository: SANC. 97. Afronychus mulibrinus Meyer, 1979: 107; Bontebok National Park, near Swellendam, Cape Province, South Africa, ex unidentified grass (Poaceae). Type Depository: SANC.

Amblypalpus Mitrofanov & Strunkova 1978 Amblypalpus Mitrofanov & Strunkova, 1978: 1095 Type species: – Amblypalpus narsikulovi Mitrofanov & Strunkova, 1978. 98. Amblypalpus masakii (Ehara & Ueckermann, 2003): 21—new combination; South Africa, intercepted at Narita airport, Chiba, Japan, ex Nebelia sp. (Bruniaceae). Species designations: Tenuipalpus masakii—original designation. Type Depository: SANC. 99. Amblypalpus narsikulovi Mitrofanov & Strunkova, 1978: 1097; Tajikistan, ex Aster alpinus L. (Asteraceae). Species designations: Tenuipalpus narsikulovi—Ghai & Shenhmar (1984). Redescriptions: Ghai & Shenhmar (1984), Mitrofanov & Strunkova (1979); Sepasgosarian (1983). Type Depository: State Nikitsky Botanical Gardens (Yalta) and Institute of Zoology and Parasitology, Academy of Sciences of the Tadjik SSR (Dushanbe). No. 2554/422.

Australopalpus Smiley & Gerson 1995 Australopalpus Smiley & Gerson, 1995: 34; Type species: Australopalpus alphitoniae Smiley & Gerson, 1995: 34; Smiley, Frost & Gerson, 1996: 170. 100. Australopalpus alphitoniae Smiley & Gerson, 1995: 34; Indooroopilly, Brisbane, Queensland, Australia, ex Alphitonia excelsa (Fenzl) Benth. (Rhamnaceae). Redescriptions: Smiley et al. (1996). Type Depository: USNM.

Brevipalpus Donnadieu 1875 Brevipalpus Donnadieu, 1875: 115. Type species: Brevipalpus obovatus Donnadieu 1875: 116; Akbar & Khalid, 1999: 155; André, 1953b: 563; Attiah, 1956: 433; Baker, 1949: 350; Baker & Pritchard, 1960: 562; Baker & Tuttle, 1964: 30; Baker & Tuttle, 1972: 29; Baker & Tuttle, 1987: 18; Chaudhri et al. 1974: 33; De Leon, 1960: 175; De Leon, 1965a: 65; Duzgunes, 1965: 124; Ehara, 1966b: 2; Evans et al. 1993: 129; Gonzalez, 1975: 81; Gupta, 1992: 92;

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Gupta, 1995: 25; Hatzinikolis, 1987: 55; Livschitz & Mitrofanov, 1967: 4; Lo, 1969b: 99; Lo, 1986: 276; Lo & Hsia, 1968: 2; McGregor, 1949: 9; Meyer, 1979: 74; Meyer & Gerson, 1980: 75; Mitrofanov, 1973a: 510; Mitrofanov & Strunkova, 1979: 77; Nassar & Ghai, 1981: 385; Pritchard & Baker, 1952: 13; Pritchard & Baker, 1958: 196; Rimando, 1962: 37; Smiley & Gerson, 1995: 36; Thewke & Enns, 1970: 80; Wainstein, 1960: 235; Welbourn et al. 2003: 109. Amissus Chaudhri, Akbar & Rasool, 1974: 31. Type species: Amissus ochaliensis Chaudhri, Akbar & Rasool, 1974; synonymy according to Sepasgosarian 1983. Brachypalpus Mitrofanov, 1973a: 511. Type species: Brevipalpus absens De Leon, 1965; synonymy according to Meyer 1979: 74, Sepasgosarian (1983), Baker & Tuttle (1987), Welbourn et al. (2003). Hystripalpus Mitrofanov, 1973a: 510. Type species: Caligonus cuneatus Canestrini & Fanzago, 1876; synonymy according to Meyer 1979: 74; Sepasgosarian (1983), Baker & Tuttle (1987), Welbourn et al. (2003). Tauripalpus Livschitz & Mitrofanov in Mitrofanov, 1973a: 511; synonymy according to Meyer 1979: 74, Sepasgosarian (1983), Welbourn et al. (2003). Tauripalpoides Pegazzano, 1975: 543. Type species: Tauripalpoides mitrofanovi Pegazzano; synonymy according to Meyer 1979: 74; Sepasgosarian (1983), Welbourn et al. (2003). Note: Meyer (1979) and Sepasgosarian (1983) considered Cenopalpus Pritchard & Baker to be a junior synonym of Brevipalpus, whereas Baker & Tuttle (1987), Hatzinikolis (1987) and Welbourn et al. (2003) considered them different genera as they are treated here. 101. Brevipalpus abiesae Baker & Tuttle, 1987: 46; Amecameca, Mexico, ex Abies religiosa Lindl. (Pinaceae). Type Depository: USNM. 102. Brevipalpus absens De Leon, 1965c: 195; El Yunque, Puerto Rico, ex Miconia foveolata Cogn. (Melastomataceae). Species designation: Brachypalpus absens—Mitrofanov & Strunkova (1979). Redescriptions: Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: MCZ. 103. Brevipalpus acatlanus Baker & Tuttle, 1987: 46; Acatlan, Jalisco, Mexico, ex Salix sp. (Salicaceae). Type Depository: USNM. 104. Brevipalpus acutangliae De Leon, 1965a: 73; Bartica Nature Reserve, Guyana (= British Guiana) ex Ocotea acutangula (Miq.) Mez. (Lauraceae). Redescriptions: Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: MCZ. Note: host originally listed as Ocotea acutanglia. 105. Brevipalpus adelos Ahmad & Akbar, 1984: 40; 4.8 km W Faisalabad, Pakistan, ex Hibiscus esculentus L. (Malvaceae). Type Depository: Acarology Research Laboratory, Department of Entomology, University of Agriculture, Faisalabad, Pakistan. Note: suspected junior synonym of Brevipalpus phoenicis (Geijskes). 106. Brevipalpus aeoloides Pritchard & Baker, 1958: 228; Phoenix, Arizona, USA, ex Franseria deltoidea Torr. (Asteraceae). Species designations: Hystripalpus aeoloides—Mitrofanov & Strunkova (1979), Brevipalpus aeloides [sic] —Baker & Yin (1988). Redescriptions: Baker & Tuttle (1964), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: USNM, No. 2361. Note: host originally listed as Franseria deltoides. 107. Brevipalpus aeolus Pritchard & Baker, 1952: 32; Indio, California, USA, ex Pluchea sericea (Nutt.) Coville (Asteraceae). Species designation: Hystripalpus aeolus—Mitrofanov & Strunkova (1979). Redescriptions: Baker & Tuttle (1964), Meyer (1979), Mitrofanov & Strunkova (1979), Pritchard & Baker (1958). Type Depository: USNM, No. 1924. 108. Brevipalpus aepi De Leon, 1961a: 43; Tuxtla Gutierrez, Chiapas, Mexico, ex Eupatorium hemiteropododum B.L. Rob. (Asteraceae).

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Species designation: Hystripalpus aepi—Mitrofanov & Strunkova (1979). Redescriptions: Baker & Tuttle (1987), Evans et al. (1993), Meyer (1979), Mitrofanov & Strunkova (1979), Ochoa & Salas (1989). Type Depository: MCZ. 109. Brevipalpus albus De Leon, 1960: 179; Santa María del Oro, Nayarit, Mexico, ex Quercus aristata Hook. & Arn. (Fagaceae). Species designation: Hystripalpus albus—Mitrofanov & Strunkova (1979). Redescriptions: Baker & Tuttle (1987), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: MCZ. 110. Brevipalpus allenrolfeae Baker & Tuttle, 1964: 49; Dome Valley, Arizona, USA, ex Allenrolfea occidentalis (S. Wats.) Kuntze (Chenopodiaceae). Species designations: Brevipalpus allenrolfia [sic]—Baker et al. (1975), Hystripalpus allenrolfeae— Mitrofanov & Strunkova (1979). Redescriptions: Baker & Tuttle (1987), Baker et al. (1975), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: USNM, No. 3027. 111. Brevipalpus alni De Leon, 1960: 183; near San Cristobal, Chiapas, Mexico, ex Alnus arguta (Schelechtendal) Spach. (Betulaceae). Species designation: Hystripalpus alni—Mitrofanov & Strunkova (1979). Redescriptions: Baker & Tuttle (1987), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: MCZ. 112. Brevipalpus alternatus De Leon, 1961a: 46; San Blas, Nayarit, Mexico, ex Conocarpus erecta L. (Combretaceae). Species designation: Hystripalpus alternatus—Mitrofanov & Strunkova (1979). Redescriptions: Baker & Tuttle (1987), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: MCZ. Synonym: 112a. Brevipalpus tetraflagellatus Livschitz in Livschitz & Salinas, 1968: 111; Santiago de las Vegas, Cuba, ex Swietenia macrophylla King (Meliaceae) and Trichilia hirta L. (Meliaceae)—Mitrofanov & Strunkova (1979). Note: first host originally listed as Swietenia macrophilla. Type Depository: Unknown. 113. Brevipalpus ambrosiae Baker & Tuttle, 1987: 31; 9.6 km north of Oaxaca, Mexico, ex Ambrosia sp. (Asteraceae). Type Depository: USNM. 114. Brevipalpus amecensis Baker & Tuttle, 1987: 49; near Amecameca, Estado de Mexico, Mexico, ex Abies religiosa Lindl. (Pinaceae). Type Depository: USNM. 115. Brevipalpus andrewsi Evans, in Evans et al. (1993): 137; Parque La Tigra, Francisco Morazan, Honduras, ex Solanum nudum Kunth & Dunal (Solanaceae). Type Depository: USNM. 116. Brevipalpus apalos Chaudhri & Akbar, 1985: 69; 3.2 km E Faisalabad, Pakistan, ex Cassia fistula L. (Caesalpiniaceae). Type Depository: Department Entomology, University of Agriculture, Faisalabad, Pakistan. 117. Brevipalpus araucanus Gonzalez, 1975: 90; transitional relict forest at Rari, near Panimavida, Linares, Chile, ex Lapageria rosea Ruiz & Pav. (Philesciaceae). Redescriptions: Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: Universidad de Chile. 118. Brevipalpus arcus Pritchard & Baker, 1958: 212; Davie, Florida, USA, ex Pluchea purpurascens (Sw.) DC. (Asteraceae). Species designation: Hystripalpus arcus—Mitrofanov & Strunkova (1979). Redescriptions: Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: USNM, No. 2349.

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119. Brevipalpus ardesiae De Leon, 1961a: 42; Mirador del Aguila, W Tepic, Nayarit, Mexico, ex Ardisia revoluta Kunth (Myrsinaceae). Species designations: Brevipalpus ardisiae [sic]—Baker & Tuttle (1987), Hystripalpus ardesiae—Mitrofanov & Strunkova (1979). Redescriptions: Baker & Tuttle (1987), Baker et al. (1975), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: MCZ. Note: host originally listed as Ardesia revoluta. 120. Brevipalpus arizonicae Baker & Tuttle, 1987: 49; Tlaxco, Tlaxcala, Mexico, ex Quercus arizonica Sarg. (Fagaceae). Type Depository: USNM. 121. Brevipalpus artemesiae Baker & Tuttle, 1964: 65; Flagstaff, Arizona, USA, ex Artemisia carruthii Wood & Carruth. (Asteraceae). Species designations: Brevipalpus artemisia [sic]—Baker et al. (1975), Hystripalpus artemisiae [sic]—Mitrofanov & Strunkova (1979), Brevipalpus artemisiae [sic] Baker & Tuttle (1987). Redescriptions: Baker & Tuttle (1987), Baker et al. (1975), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: USNM, No. 3028. 122. Brevipalpus arystis Hasan, Ashfaq, Li & Wakil, 2004: 178; Shujaabad, Faisalabad, Pakistan, ex Mangifera indica L. (Anacardiaceae). Type Depository: Acarology Research Laboratory, Department of Entomology, University of Agriculture, Faisalabad, Pakistan. Note: suspected junior synonym Brevipalpus colens Li, Hasan & Ashfaq. Leg chaetotaxy should be compared. 123. Brevipalpus asterae Baker & Tuttle, 1964: 69; Yuma, Arizona, USA, ex Aster exilis Ell. (Asteraceae). Species designations: Hystripalpus asterae—Mitrofanov & Strunkova (1979), Brevipalpus asteriae [sic]—Baker & Yin (1988). Redescriptions: Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: USNM, No. 3026. 124. Brevipalpus asthenes Pritchard & Baker, 1952: 23; Aguanga, California, USA, ex Eriodictyon crassifolium Benth (Hydrophyllaceae). Species designation: Hystripalpus asthenes—Mitrofanov & Strunkova (1979). Redescriptions: Meyer (1979), Mitrofanov & Strunkova (1979), Pritchard & Baker (1958). Type Depository: USNM, No. 1919. 125. Brevipalpus atalantae (Hatzinikolis, 1978): 46; Atalante, Pthiotis, Greece, ex Olea europaea L. (Oleaceae). Species designation: Hystripalpus atalantae—original designation, Brevipalpus (H.) atalantae—Hatzinikolis (1986b). Redescriptions: Hatzinikolis (1986b, 1986c, 1987), Hatzinikolis & Panou (1996). Type Depository: GNHM. 126. Brevipalpus baccharis Baker & Tuttle, 1987: 50; 30 km S Acatlan, Jalisco, Mexico, ex Baccharis heterophylla H.B. & K. (Asteraceae). Redescription: Evans et al. (1993). Type Depository: USNM. 127. Brevipalpus bakeri Cromroy, 1958: 78; Cayey-Salinas road, Puerto Rico, ex Guarea ramiflora Vent. (Meliaceae). Redescription: Meyer (1979). Type Depository: Collection at the Department of Entomology, North Carolina State College, USA. 128. Brevipalpus bauhiniae Manson, 1963a: 218; Lake Yojoa, Honduras, ex Bauhinia sp. aff. rubeleruziana Donn. Sm. (Caesalpiniaceae). Species designation: Hystripalpus bauhiniae—Mitrofanov & Strunkova (1979), Brevipalpus bauhinae [sic]—Evans et al. (1993). Redescriptions: Evans et al. (1993), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: USNM, No. 2888.

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129. Brevipalpus beeri Pritchard & Baker, 1952: 33; Bakersfield, California, USA, ex (Asteraceae). Species designation: Hystripalpus beeri—Mitrofanov & Strunkova (1979). Redescriptions: Meyer (1979), Mitrofanov & Strunkova (1979), Pritchard & Baker (1958). Type Depository: USNM, No. 1925. 130. Brevipalpus bicolpus Pritchard & Baker, 1958: 217; Glen Echo, Maryland, USA, ex Carica sp. (Caricaceae). Species designation: Hystripalpus bicolpus—Mitrofanov & Strunkova (1979). Redescriptions: Meyer (1979), Mitrofanov & Strunkova (1979), Ochoa & Salas (1989). Type Depository: USNM, No. 2353. 131. Brevipalpus borealis (Mitrofanov & Bitchevskis, 1975): 19; in: Mitrofanov et al. (1975), Ukraine, ex Pinus silvestris Thumb. (Pinaceae). Species designation: Hystripalpus borealis—original designation. Redescriptions: Meyer (1979), Mitrofanov et al. (1975), Mitrofanov & Strunkova (1979). Type Depository: Latvian Entomology Association. Latvian Institute Forestry Problems Scientific Research, No. 2130 132. Brevipalpus bouchea Baker, Tuttle & Abbatiello, 1975: 11; Guadalajara, Mexico, ex Bouchea prismatica (L.) Kuntze (Verbenaceae). Species designation: Brevipalpus boucheae [sic]—Baker & Tuttle (1987). Redescriptions: Baker & Tuttle (1987), Meyer (1979). Type Depository: USNM, No. 3630. 133. Brevipalpus bucerasae De Leon 1965c: 195; Juanadias, Puerto Rico, ex Bucida buceras Vell. (Combretaceae). Redescriptions: Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: MCZ. 134. Brevipalpus bumeliae De Leon, 1961c: 168; Key Largo, Florida, USA, ex Bumelia sp. (Sapotaceae). Species designation: Hystripalpus bumeliae—Mitrofanov & Strunkova (1979). Redescription: Mitrofanov & Strunkova (1979). Type Depository: MCZ. 135. Brevipalpus butcheri Pritchard & Baker, 1958: 210; S Miami, Florida, USA, ex Amaranthus sp. (Amaranthaceae). Species designation: Hystripalpus butcheri—Mitrofanov & Strunkova (1979). Redescriptions: Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: USNM, No. 2350. 136. Brevipalpus californicus (Banks, 1904): 55; Redlands, California, USA, ex orange (Rutaceae). Species designations: Tenuipalpus californicus—original designation, Brevipalpus californicus—McGregor (1949), Hystripalpus californicus—Mitrofanov & Strunkova (1979). Note: suspected senior synonym of Brevipalpus huananis Ma & Yuan and Brevipalpus junicus Ma & Yuan—this publication. Redescriptions: Ahmad & Akbar (1984), Baker (1949), Baker & Pritchard (1960), Baker & Tuttle (1964, 1987), Chandra & Channabasavanna (1974), Charanasri et al. (1986), De Leon (1961a, 1965a, 1965b, 1967), Evans et al. (1993), Gupta (1986), Gupta & Chatterjee (1997), Hatzinikolis (1986b, 1986c, 1987), Livschitz & Mitrofanov (1967), Lo (1969), McGregor (1949), Meyer (1979), Meyer & Ryke (1959), Mitrofanov & Strunkova (1979), Nassar & Ghai (1981), Ochoa & Salas (1989), Pritchard & Baker (1952, 1958), Rimando (1962), Sadana (1997), Siddiqui et al. (1979), Smiley & Gerson (1995), Wafa et al. (1968/1969), Welbourn et al. (2003), Womersley (1940) Zaher et al. (1984). Type Depository: USNM, No. 1827. Synonym: 136a. Brevipalpus browningi Baker, 1949: 382; Gaza Strip, Palestine, ex citrus (Rutaceae).—Pritchard & Baker (1958), Welbourn et al. (2003). Type Depository: BMNH. 136b. Brevipalpus confusus Baker, 1949: 380; College Park, Maryland, USA, ex orchid (Orchidaceae).—Pritchard & Baker (1958), Welbourn et al. (2003). Redescription: Andre (1953), Pritchard & Baker (1958). Type Depository: USNM, No. 1828.

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136c. Brevipalpus woglumi McGregor, 1949: 19; Spring Valley, San Diego, California, USA, ex lemon (Rutaceae).—Pritchard & Baker (1958), Welbourn et al. (2003). Redescription: Baker (1949). Type Depository: USNM, No. 1528. 136d. Tenuipalpus australis Tucker, 1926: 3; Groot Drakenstein, Western Cape Province; Durban, Natal; Pretoria and district Tomango, Tzaneen and Rustenburg, Transvaal; Bathurst, Eastern Cape Province all in South Africa, and Mazoe, Zimbabwe (= Southern Rhodesia), ex citrus (Rutaceae).—Pritchard & Baker (1958), Welbourn et al. (2003). Species designation: Brevipalpus australis—Baker (1949). Redescription: Attiah (1956), Baker (1949), Cromroy (1958), Pritchard & Baker (1952). Type Depository: Unknown. 136e. Tenuipalpus vitis Womersley, 1940: 241; Western Australia, ex lemon (Rutaceae).—Pritchard & Baker (1958), Welbourn et al. (2003). Type Depository: Unknown. 137. Brevipalpus cardinalis (Banks, 1912): 96; Phoenix, Arizona, USA, ex bark of ash tree [Fraxinus sp.] Species designations: Tenuipalpus cardinalis—original designation, Brevipalpus cardinalis—McGregor (1949), Hystripalpus cardinalis Mitrofanov & Strunkova (1979). Redescriptions: Baker (1949), Baker & Tuttle (1964), McGregor (1949), Meyer (1979), Mitrofanov & Strunkova (1979), Pritchard & Baker (1952, 1958). Type Depository: Unknown. 138. Brevipalpus cassia Baker, Tuttle & Abbatiello, 1975: 12; Fresnillo, Zacatecas, Mexico, ex Cassia crotolarioides Kunth (Caesalpiniaceae). Species designation: Brevipalpus cassiae [sic]—Baker & Tuttle (1987). Redescriptions: Baker & Tuttle (1987), Meyer (1979). Type Depository: USNM, No. 3632. 139. Brevipalpus castillejae Baker & Tuttle, 1987: 52; Tlaxco, Tlaxcala, Mexico, ex Castilleja sp. (Scrophulariaceae). Type Depository: USNM. 140. Brevipalpus ceanothus Baker & Tuttle, 1972: 29; Prescott, Arizona, USA, ex Ceanothus fendleri A. Gray (Rhamnaceae). Species designation: Hystripalpus ceanothus—Mitrofanov & Strunkova (1979). Redescriptions: Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: USNM, No. 3433. 141. Brevipalpus celtis Baker, Tuttle & Abbatiello, 1975: 11; Torreon, Coahuila, Mexico, ex Celtis pallida Torrey (Ulmaceae). Redescriptions: Baker & Tuttle (1987), Meyer (1979). Type Depository: USNM, No. 3629. 142. Brevipalpus cercidium Baker, Tuttle & Abbatiello, 1975: 18; Mazatlan, Sinaloa, Mexico, ex Cercidium floridum Benth. & A. Gray (Caesalpiniaceae). Redescriptions: Baker & Tuttle (1987), Meyer (1979). Type Depository: USNM, No. 3637. 143. Brevipalpus chalkidicus (Hatzinikolis, 1985): 37; Ormylia, Chalkidiki, Greece, ex olive (Oleaceae). Species designation: Hystripalpus chalkidicus—original designation. Redescriptions: Hatzinikolis (1986b, 1986c, 1987), Hatzinikolis & Panou (1996). Type Depository: Unknown. 144. Brevipalpus chamaedorea Baker, Tuttle & Abbatiello, 1975: 8; Mexico, intercepted at San Antonio Texas, USA, ex Chamaedorea sp. (Arecaceae). Species designation: Brevipalpus chamaedoreae [sic]—Baker & Tuttle (1987). Redescriptions: Baker & Tuttle (1987), Meyer (1979). Type Depository: USNM, No. 3627.

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145. Brevipalpus chilensis Baker, 1949: 358; Chile, intercepted at New York, USA, ex lemon (Rutaceae). Redescriptions: Gonzalez (1958), Meyer (1979), Mitrofanov & Strunkova (1979), Pritchard & Baker (1958), Sadana (1997). Type Depository: USNM, No. 1814. 146. Brevipalpus chucumayi De Leon, 1960: 179; Trinidad, Oaxaca, Mexico, ex Styrax argenteus C. Presl. (Styracaceae). Species designations: Hystripalpus chucumayi—Mitrofanov & Strunkova (1979), Brevipalpus chucamayi [sic]—Baker & Tutle (1987). Redescriptions: Baker & Tuttle (1987), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: MCZ. Note: host originally listed as Styrax argentata. 147. Brevipalpus clypealis Siddiqui, Chaudhri & Akbar, 1979: 14; 3.2 km E Peshawer, Pakistan, ex Punica granatum L. (Punicaceae). Redescription: Ahmad & Akbar (1984). Type Depository: Department of Entomology, West Pakistan Agricultural University Lyallpur, Pakistan. 148. Brevipalpus cnidosculos Baker & Tuttle, 1987: 55; 9 km S Iguala, Guerrero, Mexico, ex Cnidosculos sp. (Euphorbiaceae). Type Depository: USNM. 149. Brevipalpus cochlospermi De Leon, 1961a: 43; San Blas, Nayarit, Mexico, ex Cochlospermum sp. (Cochlospermaceae). Species designation: Hystripalpus cochlospermi—Mitrofanov & Strunkova (1979). Redescriptions: Baker & Tuttle (1987), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: MCZ. 150. Brevipalpus coffeae Meyer, 1979: 85; Jimma, Ethiopia, ex Coffea arabica L. (Rubiaceae). Type Depository: SANC. 151 Brevipalpus coldenia Baker, Tuttle & Abbatiello, 1975: 21; Chihuahua, Chihuahua, Mexico, ex Coldenia greggii A. Gray (Boraginaceae). Species designation: Brevipalpus coldeniae [sic]—Baker & Tuttle (1987). Redescriptions: Baker & Tuttle (1987), Meyer (1979). Type Depository: USNM, No. 3640. Note: host originally listed as Coldenia greggi (Torr. & Gray) Gray. 152. Brevipalpus colens Li, Hasan & Ashfaq, 2003: 645; Shujaabad, Pakistan, ex debris Mangifera indica L. (Anacardiaceae). Type Depository: Acarology Research Laboratory, Department of Entomology, University of Agriculture, Faisalabad, Pakistan. Note: suspected junior synonym of Brevipapus phoenicis (Geijskes). Leg chaetotaxy should be compared. 153. Brevipalpus colpodes Pritchard & Baker, 1958: 228; Miami, Florida, USA, ex Baccharis sp. (Asteraceae). Species designation: Hystripalpus colpodes—Mitrofanov & Strunkova (1979). Redescriptions: De Leon (1961b), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: USNM, No. 2346. 154. Brevipalpus columbiensis Thewke, 1967: 585; University of Missouri, Columbia, Missouri, USA, ex Platanus occidentalis L. (Platanaceae). Redescriptions: Meyer (1979), Thewke & Enns (1970). Type Depository: USNM. 155. Brevipalpus combreti De Leon, 1961a: 47; Cuitlahuac, Veracruz, Mexico, ex Combretum farinosum H.B. & K. (Combretaceae). Species designation: Hystripalpus combreti—Mitrofanov & Strunkova (1979). Redescriptions: Baker & Tuttle (1987), Meyer (1979), Mitrofanov & Strunkova (1979), Thewke & Enns (1970). Type Depository: MCZ.

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156. Brevipalpus conocarpi De Leon, 1961c: 168; Coral Gables, Florida, USA, ex Conocarpus erecta Griseb. (Combretaceae). Species designation: Hystripalpus conocarpi—Mitrofanov & Strunkova (1979). Redescriptions: De Leon (1965b), Mitrofanov & Strunkova (1979). Type Depository: MCZ. 157. Brevipalpus cordiae De Leon, 1961a: 42; Reynosa, Tamaulipas, Mexico, ex Cordia boissieri DC. (Boraginaceae). Species designation: Hystripalpus cordiae—Mitrofanov & Strunkova (1979). Redescriptions: Baker & Tuttle (1987), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: MCZ. 158. Brevipalpus cortesi Gonzalez, 1975: 86; Quebrada Dupliza, road to Mamiña, Cordillera de Tarapaca, Chile, ex Escallonia sp. (Escalloniaceae). Redescriptions: Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: Universidad de Chile. 159. Brevipalpus crataegus Baker & Tuttle, 1987: 35; 22.4 km east of Morelia, Michoacan, Mexico, ex Crataegus sp. (Rosaceae). Type Depository: USNM. 160. Brevipalpus creber Chaudhri, 1972b: 58; West Pakistan Agricultural University, Lyallpur, Faisalabab, Pakistan, ex Corchorus capsularis L. (Tiliaceae). Redescriptions: Ahmad & Akbar (1984), Chaudhri et al. (1974), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: Department of Entomology, West Pakistan Agricultural University Lyallpur, Pakistan. 161. Brevipalpus cromroyi Evans, in Evans et al. (1993): 139; Parque La Tigra, Francisco Morazan, Honduras, ex Ficus carica L. (Moraceae). Type Depository: USNM. 162. Brevipalpus crotonellae Baker & Tuttle, 1987: 55; Santa Catarina, San Luis Potosi, Mexico, ex Croton ciliatoglandulifer Ortega (Euphorbiaceae). Type Depository: USNM. Note: host originally listed as Croton ciliatoglandulosus Ortega. 163. Brevipalpus crotoni De Leon, 1960: 177; Ciudad Victoria, Tamaulipas, Mexico, ex Croton ciliatoglandulosus Ortega (Euphorbiaceae). Species designation: Hystripalpus crotoni—Mitrofanov & Strunkova (1979). Redescriptions: Baker & Tuttle (1987), Baker et al. (1975), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: MCZ. Synonym: 163a. Brevipalpus solanum Baker, Tuttle & Abbatiello, 1975: 7; Cavara, Alamos, Mexico, ex Solanum elaeagnifolium Cav. (Solanaceae)—Baker & Tuttle (1987). Type Depository: USNM, No. 3624. 164. Brevipalpus cucurbitae Mohanasundaram, 1982: 427; Coimbatore, Tamil Nadu, India, ex Cucurbita maxima Wall. (Cucurbitaceae). Redescriptions: Sadana (1997), Sadana & Balpreet (1995), Sadana & Kaur (1992). Type Depository: TNAU. 165. Brevipalpus cuneatus (Canestrini & Fanzago, 1876): 136; Trevignano, Treviso, Veneto, Italy, ex a hedge plant. Species designations: Caligonus cuneatus—original designation, Tenuipalpus cuneatus—Berlese (1887), Brevipalpus cuneatus—Baker (1949), Hystripalpus cuneatus—Mitrofanov & Strunkova (1979). Redescriptions: Berlese (1887), Baker (1949), Canestrini & Fanzago (1878), Geijskes (1939), Hatzinikolis (1986c, 1987), Livschitz & Mitrofanov (1967), Meyer (1979), Mitrofanov & Strunkova (1979), Pritchard & Baker (1958). Type Depository: Unknown. 166. Brevipalpus daqingis Ma & Yuan, 1982: 72; Hainan Island, Guangzhou, China, ex Clerodendrum cyrtophyllum Turcz (Verbenaceae) and Lantana camara L. (Verbenaceae).

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Type Depository: Shanghai College of Agriculture, China. Note: suspected junior synonym of Brevipalpus phoenicis (Geijskes). 167. Brevipalpus delus Pritchard & Baker, 1952: 21; Camp Nelson, California, USA, ex Libocedrus decurrens Torr. (Cupressaceae). Species designation: Hystripalpus delus—Mitrofanov et al. (1975). Redescriptions: Meyer (1979), Mitrofanov et al. (1975), Mitrofanov & Strunkova (1979), Pritchard & Baker (1958). Type Depository: USNM, No. 1915. 168. Brevipalpus dentatae Baker & Tuttle, 1987: 36; Tuxtla Gutierrez, Chiapas, Mexico, ex Cordia alba Roem. & Schult. (Boraginaceae). Redescription: Evans et al. (1993). Type Depository: USNM. 169. Brevipalpus desmodium Baker & Tuttle, 1987: 57; Puerto Palmas, Oaxaca, Mexico, ex Desmodium sp. (Fabaceae). Type Depository: USNM. 170. Brevipalpus dilaccus Pritchard & Baker, 1952: 31; Camp Nelson, California, USA, ex mountain mahogany [=Cercocarpus sp. (Rosaceae)] Species designations: Hystripalpus dilaccus—Mitrofanov & Strunkova (1979), Brevipalpus delaccus [sic]—Baker & Yin (1988). Redescriptions: Meyer (1979), Mitrofanov & Strunkova (1979), Pritchard & Baker (1958). Type Depository: USNM, No. 1923. 171. Brevipalpus dipholisi De Leon, 1961c: 167; Key Largo, Florida, USA, ex Dipholis salicifolia D.C. (Sapotaceae). Redescriptions: Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: MCZ. 172. Brevipalpus disparis Chaudhri & Akbar, 1985: 55; Gatwala forests, Faisalabad, Pakistan, ex Withania somnifera (L.) Dunal (Solanaceae). Type Depository: Department Entomology, University of Agriculture, Faisalabad, Pakistan. 173. Brevipalpus docimas Pritchard & Baker, 1958: 225; Durham, North Carolina, USA, ex hickory [= Carya sp. (Juglandaceae)]. Species designation: Hystripalpus docimas—Mitrofanov & Strunkova (1979). Redescriptions: Meyer (1979), Mitrofanov & Strunkova (1979), Thewke & Enns (1970). Type Depository: USNM, No. 2345. 174. Brevipalpus dosis Chaudhri, Akbar & Rasool, 1974: 37; University of Agriculture, Lyallpur, Faisalabad, Pakistan, ex Olea europaea L. (Oleaceae). Species designation: Brevipalpus (H.) dosis—Hatnikolis (1986b) Redescriptions: Ahmad & Akbar (1984), Hatnikolis (1986b), Hatnikolis & Panou (1996). Type Depository: Department of Entomology, University of Agriculture, Lyallpur, Pakistan. Note: suspected junior synonym of Brevipalpus oleae Baker. Chaetotaxy of gena should be compared. 175. Brevipalpus edax De Leon, 1961a: 48; Tuxtla Gutierrez, Chiapas, Mexico, ex Cordia elaeagnoides D.C. (Boraginaceae). Suspected senior synonym of: Brevipalpus rapii Hasan, Bashir, Wakil & Afzal, 2003. Based on descriptions figures these species could not be separated. Redescriptions: Baker & Tuttle (1987), Meyer (1979), Mitrofanov & Strunkova (1979). Note: host originally listed as Cordia eleagnoides. Type Depository: MCZ. 176. Brevipalpus edurensis Thewke & Enns, 1971: 246; 0.1—0.2 km from Catskill Creek, East Durham, Greene, New York, USA, ex Tilia americana L. (Tiliaceae). Species designations: Hystripalpus edurensis—Mitrofanov & Strunkova (1979), Brevipalpus endurensis [sic]—Meyer (1979). Redescriptions: Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: USNM.

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177. Brevipalpus ehsanii Hasan, Bashir, Wakil & Afzal, 2003: 355; 1 km W Narwala Bangla, Faisalabad, Punjab, Pakistan, ex Solanum tuberosum L. (Solanaceae). Type Depository: Acarology Research Laboratory, Department of Agric. Entomology, University of Agriculture, Faisalabad, Pakistan. 178. Brevipalpus edwinae Baker, 1949: 356; km 67 of Cuernavaca Highway, Morelos, Mexico, ex Eupatorium glabratum Kunth (Asteraceae). Species designation: Hystripalpus edwinae—Mitrofanov & Strunkova (1979). Redescriptions: Baker & Tuttle (1987), De Leon (1961a), Meyer (1979), Mitrofanov & Strunkova (1979), Pritchard & Baker (1958). Type Depository: USNM, No. 1812. 179. Brevipalpus emarginatae Baker & Tuttle, 1987: 57; Los Mochis, Sinaloa, Mexico, ex Atamisquea emarginata Miers (Capparaceae). Type Depository: USNM. 180. Brevipalpus encelia Baker, Tuttle & Abbatiello, 1975: 21; Hermosillo, Sonora, Mexico, ex Encelia farinosa Gray ex Torr. (Asteraceae). Species designation: Brevipalpus enceliae [sic]—Baker & Tuttle (1987). Redescriptions: Baker & Tuttle (1987), Meyer (1979). Type Depository: USNM, No. 3641. 181. Brevipalpus encinarius De Leon, 1961a: 42; E Morelia, Michoacan, km post 292 on route 15, Mexico, ex Quercus sp.(near castanea) Nee (Fagaceae). Species designation: Hystripalpus encinarius—Mitrofanov & Strunkova (1979). Redescriptions: Baker & Tuttle (1987), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: MCZ. 182. Brevipalpus ennsi Thewke, 1967: 590; 9.6 km E Ashland, Missouri, USA, ex Antennaria plantaginifolia Hook. (Asteraceae). Redescriptions: Meyer (1979), Thewke & Enns (1970). Type Depository: USNM. 183. Brevipalpus erectus Baker & Tuttle, 1987: 103; S of Cucula, Jalisco, Mexico, ex Hamelia erecta Jacq. (Rubiaceae). Type Depository: USNM. 184. Brevipalpus ericae Meyer, 1979: 90; Kirstenbosch Botanical Garden, Cape Province, South Africa, ex Erica sp. (Ericaceae). Type Depository: SANC. 185. Brevipalpus essigi Baker, 1949: 367; Berkeley, California, USA, ex Acuba sp. (Cornaceae). Species designation: Hystripalpus essigi—Mitrofanov & Strunkova (1979). Redescriptions: Baker & Tuttle (1987), Baker et al. (1975), De Leon (1961c), Gupta (1992), Hatzinikolis (1986c, 1987), Meyer (1979), Mitrofanov & Strunkova (1979), Pritchard & Baker (1952, 1958), Sadana (1997). Type Depository: USNM, No. 1819. 186. Brevipalpus euphorbiae Mohanasundaram, 1982: 427; Coimbatore, Tamil Nadu, India, ex Croton sp. (Euphorbiaceae). Redescriptions: Sadana (1997), Sadana & Kaur (1992). Type Depository: TNAU. 187. Brevipalpus ewpristori De Leon, 1960: 180; Jalapa, Veracruz, Mexico, ex Baccharis conferta H.B. & K. (Asteraceae). Species designation: Hystripalpus ewpristori—Mitrofanov & Strunkova (1979). Redescriptions: Baker & Tuttle (1987), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: MCZ. 188. Brevipalpus favus Chaudhri, 1972b: 54; 3.2 km E Taxilla, Punjab, Pakistan, ex Gymnosporia royleana M. Laws. (Celastraceae).

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Species designation: Hystripalpus favus—Mitrofanov & Strunkova (1979). Redescriptions: Chaudhri & Akbar (1985), Chaudhri et al. (1974), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: Department of Entomology, West Pakistan Agricultural University Lyallpur, Pakistan. Note: host originally listed as Gyamnosporia royleana. 189. Brevipalpus fenghuangis Ma & Yuan, 1982: 71; Jianfeng, Hainan Island, China, ex Delonix regia (Bojer) Raf. (Caesalpiniaceae). Type Depository: Shanghai College of Agriculture, China. 190. Brevipalpus filifer De Leon, 1960: 182; near San Cristobal, Chiapas, Mexico, ex Persea schiedeana Nees (Lauraceae). Species designation: Hystripalpus filifer—Mitrofanov & Strunkova (1979). Redescriptions: Baker & Tuttle (1987), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: MCZ. Note: host originally listed as Persea schideana. 191. Brevipalpus filifolia Baker, Tuttle & Abbatiello, 1975: 19; Agua, south of Ciudad Juarez, Mexico, ex Artemisia filifolia Torr. (Asteraceae). Species designation: Brevipalpus filifoliae [sic]—Baker & Tuttle (1987). Redescriptions: Baker & Tuttle (1987), Meyer (1979). Type Depository: USNM, No. 3638. 192. Brevipalpus fleschneri Pritchard & Baker, 1958: 203; Juancito Mountain, Honduras, ex Persea popenoei L.O. Williams (Lauraceae). Species designation: Hystripalpus fleschneri—Mitrofanov & Strunkova (1979). Redescriptions: Evans et al. (1993), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: USNM, No. 2356. Note: host originally listed as Persea popenoi. 193. Brevipalpus floridianus De Leon, 1956b: 113; Coral Gables, Florida, USA, ex Persea (Tamala) borbonia Spreng. (Lauraceae). Species designations: Hystripalpus floridianus—Mitrofanov & Strunkova (1979), Brevipalpus floridanus [sic]—Meyer (1979). Redescriptions: Meyer (1979), Mitrofanov & Strunkova (1979), Pritchard & Baker (1958). Type Depository: USNM. 194. Brevipalpus formosus De Leon, 1960: 177; Aticama, Nayarit, Mexico, ex Licaria sp. (Lauraceae). Species designation: Hystripalpus formosus—Mitrofanov & Strunkova (1979). Redescriptions: Baker &Tuttle (1987), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: MCZ: 195. Brevipalpus frankenia Baker, Tuttle & Abbatiello, 1975: 14; Hermosillo, Sonora, Mexico, ex Frankenia palmeri S. Watson (Frankeniaceae). Species designation: Brevipalpus frankeniae [sic]—Baker &Tuttle (1987). Redescriptions: Baker & Tuttle (1987), Meyer (1979). Type Depository: USNM, No. 3633. 196. Brevipalpus fraxini De Leon, 1961c: 173; Columbus, Georgia, USA, ex Fraxinus profunda Bush & Britton (Oleaceae). Species designation: Hystripalpus fraxini—Mitrofanov & Strunkova (1979). Redescriptions: Baker & Tuttle (1987), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: MCZ. 197. Brevipalpus galliprodiens Ochoa & Salas, 1987: 227; El Pelón de la Bajura, Guanacaste, Costa Rica, ex Cochlospermum vitifolium Spreng. (Cochlospermaceae). Redescription: Ochoa & Salas (1989). Type Depository: Colección de Referencia, Laboratorio de Acarologia, Facultad de Agronomia, Universidad de Costa Rica, Costa Rica.

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198. Brevipalpus garmani Baker, 1949: 363; Hamden, Connecticut, USA, ex Sambucus sp. elder (Caprifoliaceae). Species designation: Hystripalpus garmani—Mitrofanov & Strunkova (1979). Redescriptions: De Leon (1961c), Meyer (1979), Mitrofanov & Strunkova (1979), Pritchard & Baker (1958), Thewke & Enns (1970). Type Depository: USNM, No. 1818. 199. Brevipalpus gauhatiensis Sadana & Gupta, 1983: 3; Gauhati, Kamrup, Assan, India, ex Jasminum grandiflorum L. (Oleaceae). Redescription: Sadana (1997). Type Depository: Unknown. Note: host originally listed as Jasminum grandiflora. 200. Brevipalpus geranium Baker & Tuttle, 1987: 61; Mexico, intercepted at Roma, Texas quarantine station, Texas, USA, ex Geranium sp. (Geraniaceae). Type Depository: USNM. 201. Brevipalpus gliricidiae De Leon, 1961a: 43; Tuxtla Gutierrez, Chiapas, Mexico, ex Gliricidia sepium (Jacq.) Kunth & Walp. (Fabaceae). Species designation: Hystripalpus gliricidiae—Mitrofanov & Strunkova (1979). Redescriptions: Baker & Tuttle (1987), Meyer (1979), Mitrofanov & Strunkova (1979), Ochoa & Salas (1989). Type Depository: MCZ. 202. Brevipalpus glomeratus Pritchard & Baker, 1958: 222; McLean, Virginia, USA, ex oak (Quercus sp.—Fagaceae). Species designation: Hystripalpus glomeratus—Mitrofanov & Strunkova (1979). Redescriptions: De Leon (1961c), Meyer (1979), Mitrofanov & Strunkova (1979), Thewke & Enns (1970). Type Depository: USNM, No. 2344. 203. Brevipalpus glymma Pritchard & Baker, 1958: 208; Homestead, Florida, USA, ex wild lantana [=Lantana sp., Verbenaceae] Species designation: Hystripalpus glymma—Mitrofanov & Strunkova (1979). Redescriptions: Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: USNM, No. 2358. 204. Brevipalpus gortiniensis Hazinikolis & Panou, 1996a: 15; Monastiraki, Gortinia, Arkadia, Peleponessus, Greece, ex Olea europaea L. (Oleaceae). Type Depository: Collection of the Acarology Laboratory of Agricultural Research Centre of Athens, Greece. 205. Brevipalpus grandis Mitchell, 1974: 4; Monroe, Benton, Oregon, USA, ex Abies grandis (Dougl. & D. Don) Lindl. (Pinaceae). Redescription: Meyer (1979). Type Depository: USNM. 206. Brevipalpus grewiae (Rübsaamen, 1899): 255; Mangalor, India, ex Grewia microcos L. (Tiliaceae). Species designations: Pediculoides grewiae—original designation, Tenuipalpus grewiae—Oudemans (1938), Brevipalpus grewiae—Baker (1949). Redescriptions: Baker (1949), Oudemans (1938). Type Depository: Unknown. 207. Brevipalpus guihuanis Ma & Yuan, 1982: 72; Kunming, Yunnan, China, ex Osmanthus fragrans Lour. (Oleaceae). Type Depository: Shanghai College of Agriculture, China. 208. Brevipalpus hafizii Chaudri & Akbar, 1985: 74; 19.2 km N Karachi, Sind, Pakistan, ex Azadirachta indica A. Juss. (Meliaceae). Type Depository: Department Entomology, University of Agriculture, Faisalabad, Pakistan. Note: suspected junior synonym Brevipalpus phoenicis (Geijskes). Synonym: 208a Brevipalpus hafizii Akbar & Khalid, 1999: 159; 12 km N Karachi, Sind, Paquistão, ex Azadirachta indica A. Juss. (Meliaceae)—new synonomy. Note: The drawings of the original description of B. hafizzi Akbar & Khalid (1999) are exactly the same as those of

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the original description of B. hafizzi Chaudhri & Akbar (1985). The site of Collection of the former species (12 km north of Karachi) is also similar to that of the latter (12 miles north of Karachi). Despite of the minor differences in the collectors and collection dates, they appear to be the same species and therefore B. hafizii Akbar & Khalid, 1999 is a primary homonym and junior synonym. Type Depository: Acarology Research Laboratory, Department of Agric. Entomology, University of Agriculture, Faisalabad, Pakistan. Note: suspected junior synonym Brevipalpus phoenicis (Geijskes). 209. Brevipalpus hainanensis Ma & Yuan, 1982: 72; Hainan Island, China, ex an unidentified plant. Type Depository: Shanghai College of Agriculture, China. Note: suspected junior synonym Brevipalpus phoenicis (Geijskes). 210. Brevipalpus hamelrectae Baker & Tuttle, 1987: 61; Fortin, Veracruz, Mexico, ex Hamelia erecta Jacq. (Rubiaceae). Type Depository: USNM. 211. Brevipalpus hamzaii Hasan, Wakil, Bashir & Kwon, 2005: 15; 3.2 km E Gujranwala, Punjab, Pakistan, ex Helianthus annuus L. (Asteraceae). Type Depository: Acarology Research Laboratory, Department of Agricultural Entomology, University of Agriculture, Faisalabad, Pakistan. 212. Brevipalpus hashmii Chaudhri & Akbar, 1985: 53; 12.8 km N Faisalabad, Pakistan, ex Cucurbita pepo L. (Cucurbitaceae). Type Depository: Department Entomology, University of Agriculture, Faisalabad, Pakistan. Note: host originally listed as Cucurbita peppo. 213. Brevipalpus hellenicus (Hatzinikolis & Kolovos, 1981): 205; Sperhogaea, Messinia and Aghios Theologos, Phthiotis, Greece, ex Olea europaea L. (Oleaceae). Species designation: Hystripalpus hellenicus—original designation. Redescriptions: Hatzinikolis (1986b, 1986c, 1987), Hatzinikolis & Panou (1996). Type Depository: BMNH. 214. Brevipalpus hernandiae De Leon, 1965b: 519; Bath Fountain, Saint Thomas Parish, Jamaica, ex Hernandia catalpifolia Britton & Harris (Hernandiaceae). Redescription: Meyer (1979). Type Depository: MCZ. Note: host originally listed as Hernandia catalpafolia. 215. Brevipalpus homalus Pritchard & Baker, 1952: 25; Mojave, California, USA, ex (Asteraceae). Species designation: Hystripalpus homalus—Mitrofanov & Strunkova (1979). Redescriptions: Baker & Tuttle (1964), Meyer (1979), Mitrofanov & Strunkova (1979), Pritchard & Baker (1958). Type Depository: USNM, No. 1920. 216. Brevipalpus hondurani Evans, in Evans et al. (1993): 141; Parque La Tigra, Francisco Morazan, Honduras, ex Heterocentron subtriplinervium A. Braun & Bouche (Melastomataceae). Redescription: Evans et al. (1998). Type Depository: USNM. 217. Brevipalpus huananis Ma & Yuan, 1982:72; Guangzhou and Hainan Islands, Guangdong and Nanning Islands, Guangxi, China, ex Bauhinia purpurea L. (Caesalpiniaceae), Camellia sinensis (L.) O. Kuntze (Theaceae), Urena lobata L. (Malvaceae), Photinia sp. (Rosaceae), Chrysalidocarpus lutescens H.Wendl. (Arecaceae), Caryota ochlandra Hance (Arecaceae), Celtis sinensis Pers., (Ulmaceae), Ipomoea cairica (L.) Sweet (Convolvulaceae), Ruellia repens Ruiz (Acanthaceae), Syringa sp. (Oleaceae), Hevea brasiliensis Mull. Arg. (Euphorbiaceae), Plumeria rubra L. (Apocynaceae), Allamanda cathartica L., (Apocynaceae), Ipomoea sp. (Convolvulaceae), Theobroma cacao L. (Sterculiaceae), Pharbitis nil Gardn. (Convolvulaceae). Type Depository: Shanghai College of Agriculture, China. Note: host originally listed as Phabitis nil.

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218. Brevipalpus hybus Pritchard & Baker, 1958: 212; Indiantown, Florida, USA, ex an unidentified weed. Species designation: Hystripalpus hubus [sic]—Mitrofanov& Strunkova (1979). Redescription: Mitrofanov & Strunkova (1979). Type Depository: USNM, No. 2359. 219. Brevipalpus hypti De Leon, 1960: 175; Ixtlan del Rio, Nayarit, Mexico, ex Hyptis albida H. B. & K. (Lamiaceae). Species designation: Hystripalpus hypti—Mitrofanov & Strunkova (1979). Redescriptions: Baker & Tuttle (1987), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: MCZ. 220. Brevipalpus incarnatae Baker & Tuttle, 1987: 63; south of Cucula, Jalisco, Mexico, ex Allionia incarnata L. (Nyctaginaceae). Type Depository: USNM. 221. Brevipalpus inermis Akbar & Aheer, 1994: 340; 0.5 km SW Hafizabad, Pakistan, ex Momordica charantia L. (Cucurbitaceae). Type Depository: Acarology Research Laboratory, Deparment of Agric. Entomology, University of Agriculture, Faisalabad, Pakistan. 222. Brevipalpus insinuatus De Leon, 1960: 180; Tuxtla Gutierrez, Chiapas, Mexico, ex Quercus sp. (Fagaceae). Species designation: Hystripalpus insinuatus—Mitrofanov & Strunkova (1979). Redescriptions: Baker & Tuttle (1987), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: MCZ. 223. Brevipalpus ipomoeae Baker & Tuttle, 1964: 58; McNary, Arizona, USA, ex Ipomoea plummerae A. Gray (Convolvulaceae). Species designations: Brevipalpus ipmoeae and Brevipalpus ipomoeae—original designation, Hystripalpus ipomoeae—Mitrofanov & Strunkova (1979). Redescriptions: Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: USNM, No. 3033. 224. Brevipalpus iraquiensis Al-Gboory, 1987: 117; Abu-Ghraib, Baghdad, Iraq, ex Codiaeum variegatum Muell (Euphorbiaceae). Type Depository: Entomology Museum, College of Agriculture, Department of Plant Protection, Abu-Ghraib, Baghdad, Iraq. Note: host originally listed as Croton variegatus L. 225 Brevipalpus jambhiri Sadana & Balpreet, 1995: 75; Punjab Agricultural University, Ludhiana, Punjab, India, ex Citrus jambhiri Lush. (Rutaceae). Type Depository: PAUP. Note: suspected junior synonym of Brevipapus phoenicis (Geijskes). 226. Brevipalpus janeae De Leon, 1961c: 171; Columbus, Georgia, USA, ex Aureolaria flava Farw. subsp. reticulata (Raf.) Pennell (Scrophulariaceae). Species designation: Hystripalpus janeae—Mitrofanov & Strunkova (1979). Redescriptions: Evans et al. (1993); Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: MCZ. 227. Brevipalpus johnstoni Baker & Tuttle, 1987: 64; Mexico, intercepted at San Antonio, Texas quarantine station, Texas, USA, ex Chamaedorea sp. (Arecaceae). Type Depository: USNM. 228. Brevipalpus jordani Dosse, 1972: 7; Zitrone, Jerico, Jordan, ex Citrus sp. (Rutaceae). Redescription: Mitrofanov & Strunkova (1979). Type Depository: Universitat Hohenhein, Stuttgart-Hohenheim, Germany. Note: Dosse (1972) recorded the holotype as a larve and paratype a nymph. 229. Brevipalpus juncus Chaudhri & Akbar, 1985: 57; 4.8 km S Karachi, Sind, Pakistan, ex Ocimum basilicum L. (Lamiaceae).

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Type Depository: Department Entomology, University of Agriculture, Faisalabad, Pakistan. Synonym: 229a. Brevipalpus juncus Akbar & Khalid, 1999: 156; 3 km S Karachi, Sind, Pakistan, ex Ocimum basilicum L. (Labiatae)—new synonomy. Note: The drawings of the original description of B. juncus Akbar & Khalid (1999) are exactly the same as those of the original description of B. juncus Chaudhri & Akbar (1985). The site of Collection of the former species (3 km south of Karachi) is also similar to that of the latter (3 miles south of Karachi). Despite of the minor differences in the collectors and collection dates, they appear to be the same species and therefore B.juncusi Akbar & Khalid, 1999 is a primary homonym and junior synonym Type Depository: Acarology Research Laboratory, Department of Agric. Entomology, University of Agriculture, Faisalabad, Pakistan. 230. Brevipalpus junicus Ma & Yuan, 1982: 72; Xinglong, Hainan Island, China, ex Citrus reticulata Blanco (Rutaceae). Type Depository: Shanghai College of Agriculture, China. Note: suspected junior synonym of Brevipalpus californicus (Banks). 231. Brevipalpus juniperus Baker & Tuttle, 1987: 64; Tlaxco, Tlaxcala, Mexico, ex Juniperus deppeana Steud. (Cupressaceae). Type Depository: USNM. Note: host originally listed as Juniperus mexicanus. 232. Brevipalpus khalidae Hasan, Wakil, Bashir & Kwon, 2005: 13; 3 km N Gujrat, Punjab, Pakistan, ex Trianthema portulacastrum L. (Aizoaceae). Type Depository: Acarology Research Laboratory, Department of Agricultural Entomology, University of Agriculture, Faisalabad, Pakistan. Note: host originally listed as Trianthema partulacastrum L. 233. Brevipalpus karackiensis Chaudhri, Akbar & Rasool, 1974: 43; 16 km N Karachi, Sind, Pakistan, ex Convolvulus sp. (Convolvulaceae). Species designations: Brevipalpus karackiensis and B. karachiensis—original designation. Redescriptions: Ahmad & Akbar (1984), Sadana (1997). Type Depository: Department of Entomology, University of Agriculture, Lyallpur, Pakistan. 234. Brevipalpus kasuriensis Chaudhri & Akbar, 1985: 78; 1.6 km E Kasur, Pakistan, ex an unidentified plant. Type Depository: Department Entomology, University of Agriculture, Faisalabad, Pakistan. 235. Brevipalpus keiferi Pritchard & Baker, 1958: 205; Nimbus, California, USA, ex Baccharis pilularis DC. (Asteraceae). Species designation: Hystripalpus keiferi—Mitrofanov & Strunkova (1979). Redescriptions: Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: USNM, No. 2357. 236. Brevipalpus lagasceae De Leon, 1960: 176; Arenal, Jalisco, Mexico, ex Lagascea angustifolia DC. (Asteraceae). Species designation: Hystripalpus lagasceae—Mitrofanov & Strunkova (1979). Redescriptions: Baker & Tuttle (1987), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: MCZ. 237. Brevipalpus lantanae Baker & Tuttle, 1987: 66; Zempoala, Veracruz, Mexico, ex Lantana sp. (Verbenaceae). Type Depository: USNM. 238. Brevipalpus lepidium Baker & Tuttle, 1987: 38; La Placita, 145 km N Pachuca, Hidalgo, Mexico, ex Lepidium sp. (Brassicaceae). Type Depository: USNM. 239. Brevipalpus levis De Leon, 1960: 183; Cuidad Veracruz, Veracruz, Mexico, ex Cordia glabra Cham. (Boraginaceae). Species designation: Hystripalpus levis—Mitrofanov & Strunkova (1979). Redescriptions: Baker & Tuttle (1987), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: MCZ.

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240. Brevipalpus lewisi McGregor, 1949: 17; Porterville, California, USA, ex lemon (Rutaceae). Species designation: Hystripalpus lewisi—Mitrofanov & Strunkova (1979). Redescriptions: Attiah (1956), Baker (1949), Baker & Tuttle (1964, 1987), Ehara (1956), Hatzinikolis (1982, 1986c, 1987), Khosrowshahi & Arbabi (1997), Livschitz & Mitrofanov (1967), Ma & Yuan (1977), Meyer (1979), Mitrofanov & Strunkova (1979), Pritchard & Baker (1952, 1958), Sadana (1997), Smiley & Gerson (1995), Tseng (1974). Type Depository: USNM, No. 1527. 241. Brevipalpus lilium Baker, 1949: 369; Mabton, Washington, USA, ex Lilium regale E.H. Wilson (Liliaceae). Species designation: Hystripalpus lilium—Mitrofanov & Strunkova (1979). Redescriptions: De Leon (1965a), Hatzinikolis (1986c, 1987), Meyer (1979), Mitrofanov & Strunkova (1979), Pritchard & Baker (1952, 1958). Type Depository: USNM, No. 1821. 242. Brevipalpus linki Baker, 1949: 370; Orlando, Florida, USA, ex Quercus sp. (Fagaceae). Species designation: Hystripalpus linki—Mitrofanov & Strunkova (1979). Redescriptions: Meyer (1979), Mitrofanov & Strunkova (1979), Pritchard & Baker (1958). Type Depository: USNM, No. 1822. 243. Brevipalpus lippiae Baker & Tuttle, 1987: 68; Puerto Palma, Oaxaca, Mexico, ex Lippia sp. (Verbenaceae). Type Depository: USNM. 244. Brevipalpus lochmius Pritchard & Baker, 1958: 230; Camp Nelson, California, USA, ex chaparral. Redescriptions: Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: USNM, No. 2347. 245. Brevipalpus longisetosus Baker, 1949: 377; Yauco, Puerto Rico, ex an unidentified plant. Species designation: Hystripalpus longisetosus—Mitrofanov & Strunkova (1979). Redescriptions: Baker & Tuttle (1987), De Leon (1961a), Meyer (1979), Mitrofanov & Strunkova (1979), Pritchard & Baker (1958). Type Depository: USNM, No. 1826. 246. Brevipalpus lotus Baker & Tuttle, 1987: 39; 1.6 km north of Oaxaca, Oaxaca State, Mexico, ex Lotus sp. (Fabaceae). Type Depository: USNM. 247. Brevipalpus lupinus Baker & Tuttle, 1987: 96; 33.6 km E Huajuapan, Oaxaca, Mexico, ex Lupinus sp. (Fabaceae). Type Depository: USNM. 248. Brevipalpus lyallpuriensis Ahmad & Akbar, 1984; Faisalabad, Pakistan, ex Hibiscus esculentus L. (Malvaceae). Type Depository: Unknown. Note: original description of species could not be located. The information provided here was taken from Akbar & Aheer (1994). 249. Brevipalpus lysilomae De Leon, 1961c: 172; Key Largo, Florida, USA, ex Lysiloma bahamensis Benth. (Fabaceae). Species designations: Hystripalpus lysilomae—Mitrofanov & Strunkova (1979), Brevipalpus lysiloma [sic]—Baker & Yin (1988). Redescriptions: Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: MCZ. 250. Brevipalpus macedonicus (Hatzinikolis, 1983): 71; Vrasna, Thessaloniki, Macedonia, Greece, ex olive (Oleaceae). Species designation: Hystripalpus macedonicus—original designation. Redescriptions: Hatzinikolis (1986b, 1986c, 1987), Hatzinikolis & Panou (1996). Type Depository: female holotypes and male allotype with a number of females, males, nymphs and a larva paratypes are deposited in the author’s Collection. 251. Brevipalpus mallorquensis Pritchard & Baker, 1958: 214; Palma Mallorca, Spain, ex Adenostoma sp. (Rosaceae). Species designation: Hystripalpus mallorquensis—Mitrofanov & Strunkova (1979). Redescriptions: Hatzinikolis (1986c, 1987), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: USNM, No. 2352.

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252. Brevipalpus mcgregori Baker, 1949: 357; Santa Paula, California, USA, ex lemon Citrus limon L. (Rutaceae). Redescriptions: Khosrowshahi & Arbabi (1997), Meyer (1979), Mitrofanov & Strunkova (1979), Pritchard & Baker (1952, 1958). Type Depository: USNM, No. 1813. 253. Brevipalpus melichrus Pritchard & Baker, 1952: 40; Modesto, California, USA, ex Acer negundo L. var. californicum (Torr. & A. Gray) Sarg. (Aceraceae). Redescriptions: Baker & Tuttle (1964, 1972), Lo (1969), Meyer (1979), Mitrofanov & Strunkova (1979), Pritchard & Baker (1958). Type Depository: USNM, No. 1927. 254. Brevipalpus mexicanus Baker & Tuttle, 1987: 69; Tlaxco, Tlaxcala, Mexico, ex Juniperus deppeana Steud. (Cupressaceae). Redescription: Evans et al. (1993). Type Depository: USNM. Note: host originally listed as Juniperus mexicanus Spreng. 255. Brevipalpus mingoraensis Chaudhri & Akbar, 1985: 59; Mingora, Swat Valley, Pakistan, ex an unidentified plant. Type Depository: Department Entomology, University of Agriculture, Faisalabad, Pakistan. 256. Brevipalpus mitrofanovi (Pegazzano, 1975): 549; Selvapiana, (Rufina—Firenze), Italy, ex Quercus cerris L. (Fagaceae). Species designation: Tauripalpoides mitrofanovi—original designation, Brevipalpus mitrofanovi—Meyer (1979). Redescription: Meyer (1979). Type Depository: ISZA. 257. Brevipalpus moreliensis Baker & Tuttle, 1987: 69; 22.4 km E Morelia, Michoacan, Mexico, ex Quercus sp. (Fagaceae). Type Depository: USNM. 258. Brevipalpus mori De Leon, 1960: 184; Trinidad, Oaxaca, Mexico, ex Morus alba L. (Moraceae). Species designation: Hystripalpus mori—Mitrofanov & Strunkova (1979). Redescriptions: Baker & Tuttle (1987), De Leon (1965b), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: MCZ. Synonym: 258a. Brevipalpus sexflagellatus Livschitz in Livschitz & Salinas, 1968: 108; Jardin Botanico, Universidad de La Habana, Cuba, ex Guazuma tomentosa H.B. & K. (Sterculiaceae) Salinas—Mitrofanov & Strunkova (1979). Type Depository: Unknown. 259. Brevipalpus mumai Baker & Tuttle, 1964: 47; Grand Canyon, Arizona, USA, ex Petrophytum caespitosum (Nutt.) Rybd. (Rosaceae). Species designation: Hystripalpus mumai—Mitrofanov & Strunkova (1979). Redescriptions: Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: USNM, No. 3032. 260. Brevipalpus nangalensis Sadana & Kaur, 1992: 95, Nangal, Rupnagar (= Ropar), Punjab, India, ex Jasminum sambac (L.) Ait. (Oleaceae). Type Depository: Unknown. 261. Brevipalpus nannus Pritchard & Baker, 1952: 34; Aguanga, California, USA, ex Arctostaphylos bicolor A. Gray Nutt. (Ericaceae). Species designation: Hystripalpus nannus—Mitrofanov & Strunkova (1979). Redescriptions: Meyer (1979), Mitrofanov & Strunkova (1979), Pritchard & Baker (1958). Type Depository: USNM, No. 1922. 262. Brevipalpus neoardisiae Baker & Tuttle, 1987: 40; 19.2 km east of Morelia, Michoacan, Mexico, ex Baccharis heterophylla H.B. & K. (Asteraceae). Type Depository: USNM.

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263. Brevipalpus neobicolpus Baker & Tuttle, 1987: 41; 6.4 km E Pachuca, El Manzana, Hidalgo, Mexico, ex Lupinus sp. (Fabaceae). Type Depository: USNM. 264. Brevipalpus neohyptis Baker, Tuttle & Abbatiello, 1975: 8; Zapotlanejo, Jalisco, Mexico, ex Hyptis sp. (Lamiaceae). Redescriptions: Baker & Tuttle (1987), Meyer (1979). Type Depository: USNM, No. 3625. 265. Brevipalpus neoreligiosae Baker & Tuttle, 1987: 71; near Mexico city, Federal District, Mexico, ex Abies religiosa Lindl. (Pinaceae). Type Depository: USNM. 266. Brevipalpus nocivus Siddiqui, Chaudhri & Akbar, 1979: 12; University of Faisalabad, Punjab, Pakistan, ex Cordia rothii Roem. & Schult. (Boraginaceae). Redescription: Ahmad & Akbar (1984). Type Depository: Department of Entomology, West Pakistan Agricultural University Lyallpur, Pakistan. Note: suspected junior synonym of Brevipalpus phoenicis (Geijskes). 267. Brevipalpus nodiflorae Baker & Tuttle, 1987: 72; south of Los Mochis, Sinaloa, Mexico, ex Phyla nodiflora (L.) Greene (Verbenaceae). Redescription: Evans et al. (1993). Type Depository: USNM. 268. Brevipalpus oaxacensis De Leon, 1960: 182; Oaxaca, Oaxaca State, Mexico, ex Quercus conzattii Trel. (Fagaceae). Species designation: Hystripalpus oaxacensis—Mitrofanov & Strunkova (1979). Redescriptions: Baker & Tuttle (1987), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: MCZ. Note: host originally listed as Quercus conzatii. 269. Brevipalpus obovatus Donnadieu, 1875: 116; Lyon, France, ex Phytolacca sp. (Phytolaccaceae). Suspected senior synonym of: Brevipalpus quianniunis Ma & Yuan; Brevipalpus tinsukiaensis Sadana & Gupta—this publication. Redescriptions: Ahmad & Akbar (1984), Al-Gboory (1987), Attiah (1956), Baker (1945, 1949), Baker & Pritchard (1960), Baker & Tuttle (1964, 1987), Banks (1912), Blanchard (1940), Chaudhri et al. (1974), De Leon (1961a, 1965b, 1967), Duzgunes (1965), Ehara (1966), Evans et al. (1993, 1998), Gupta (1986, 1992), Gupta & Chatterjee (1997), Hatzinikolis (1986c, 1987), Khosrowshahi & Arbabi (1997), Livschitz & Mitrofanov (1967), Livschitz & Salinas (1968), Ma & Yuan (1977), McGregor (1914, 1949), Meyer (1979), Meyer & Ryke (1959), Mitrofanov & Strunkova (1979), Nassar & Ghai (1981), Ochoa & Salas (1989), Pritchard & Baker (1952, 1958), Reck (1951, 1952), Rimando (1962), Sadana (1997), Smiley & Gerson (1995), Thewke & Enns (1970), Tseng (1974), Wafa et al. (1968/1969), Wainstein (1960), Welbourn et al. (2003), Zaher et al. (1984). Type Depository: Faculté des Sciences de Lyon, France. Synonym: 269a Brevipalpus amicus Chaudhri, 1972b: 65; Charrapani, Murree Hills, Pakistan, ex Andrachne sp. (Euphorbiaceae)—new synonym. Redescriptions: Ahmad & Akbar (1984), Chaudhri et al. (1974), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: Department of Entomology, West Pakistan Agricultural University Lyallpur, Pakistan. 269b. Brevipalpus assamensis Sadana & Gupta, 1983: 1; New Bongaigaon, Assam, India, ex an unidentified plant—new synonym. Redescriptions: Sadana (1997). Type Depository: Unknown. 269c. Brevipalpus origanum Baker, Tuttle & Abbatiello, 1975: 18; Mexico, intercepted at San Antonio, Texas, USA, ex Origanum sp. (Lamiaceae)—Pritchard & Baker (1952), Welbourn et al. (2003). Redescription: Baker & Tuttle (1987), Meyer (1979). Type Depository: USNM, No. 3636.

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269d. Brevipalpus pereger Donnadieu, 1875: 117; Lyon, France, ex Rubus sp. (Rosaceae) and rose.—Pritchard & Baker (1952), Welbourn et al. (2003). Type Depository: Faculté des Sciences de Lyon, France. 269e. Tenuipalpus bioculatus McGregor, 1914: 354; Batesburg, South Carolina, USA, ex Ligustrum amurense Carriere (Oleaceae).—Pritchard & Baker (1952), Welbourn et al. (2003). Species designation: Brevipalpus bioculatus—McGregor (1916). Redescriptions: Reck (1951, 1952). Type Depository: USNM, No. 19090. 269f. Tenuipalpus inornatus Banks, 1912: 97; Batesburg, California, USA, ex golden rod (Asteraceae)—Pritchard & Baker (1952), Welbourn et al. (2003). Species designation: Tenuipalpus inornatus—original designation, Brevipalpus inornatus—Baker (1945). Redescriptions: Attiah (1956), Baker (1945, 1949), Chaudhri (1972b), Cromroy (1958), McGregor (1949), Pritchard & Baker (1952). Type Depository: USNM, No. 1815. 269g.Tenuipalpus pseudocuneatus Blanchard, 1940: 11; Corrientes, Entre Ríos and Santa Fe, Argentina, ex ?Citrus sp. (Rutaceae)—Pritchard & Baker (1952), Welbourn et al. (2003). Species designation: Brevipalpus pseudocuneatus—Baker (1949). Redescription: Baker (1949). Type Depository: types are deposited in the author’s Collection. 270. Brevipalpus obovoides De Leon, 1961c: 167; Key Largo, Florida, USA, ex Forestiera porulosa Poir. (Oleaceae). Redescriptions: Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: MCZ. 271. Brevipalpus ochaliensis (Chaudhri, Akbar & Rasool, 1974): 31; 1.6 km W Ochali, Pakistan, ex Adhatoda vasica Nees (Acanthaceae). Species designation: Amissus ochaliensis—original designation, Brevipalpus ochaliensis—Ghai & Shenhmar (1984). Catalogue: Ghai & Shenhmar (1984). Type Depository: Department Entomology, University of Agriculture Lyallpur, Pakistan. Note: host originally listed as Adhatoda vesica. 272. Brevipalpus ochoai Evans, in Evans et al. (1993): 141; El Zamorano, Francisco Morazan, Honduras, ex Lasianthaea fruticosa (L.) K. M. Becker (Asteraceae). Type Depository: USNM. 273. Brevipalpus ocoteae De Leon, 1961c: 173; Key Largo, Florida, USA, ex Ocotea coriacea Britton (Lauraceae). Species designation: Hystripalpus ocotea—Mitrofanov & Strunkova (1979). Redescriptions: Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: MCZ. 274. Brevipalpus ogcotus Pritchard & Baker, 1952: 21; Lemon Cove, California, USA, ex Eriodictyon californicum Greene (Hydrophyllaceae). Species designation: Hystripalpus ogcotus—Mitrofanov & Strunkova (1979). Redescriptions: Meyer (1979), Mitrofanov & Strunkova (1979), Pritchard & Baker (1958). Type Depository: USNM, No. 1918. 275. Brevipalpus ogmellus Pritchard & Baker, 1958: 225; Key Largo, Florida, USA, ex oak [Quercus sp., Fagaceae] Species designation: Hystripalpus ogmellus—Mitrofanov & Strunkova (1979). Redescriptions: Baker & Tuttle (1964), De Leon (1961b), Evans et al. (1993), Meyer (1979), Mitrofanov & Strunkova (1979), Thewke & Enns (1970). Type Depository: USNM, No. 2360. 276. Brevipalpus ogmus Pritchard & Baker, 1952: 27; Palo Alto, California, USA, ex Quercus lobata Nee (Fagaceae). Species designation: Hystripalpus ogmus—Mitrofanov & Strunkova (1979). Redescriptions: Meyer (1979), Mitrofanov & Strunkova (1979), Pritchard & Baker (1958). Type Depository: USNM, No. 1914.

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277. Brevipalpus oleae Baker, 1949: 361; Morocco?, ex olive (Oleaceae). Species designation: Hystripalpus oleae—Castagnoli & Pegazzano (1979), Brevipalpus (H.) oleae—Hatzinikolis (1986b). Suspected senior synonym of: Brevipalpus dosis Chaudhri, Akbar & Rasool, Brevipalpus olearius Sayed, Brevipalpus olivicola Pegazzano & Castagnoli—this publication. Redescriptions: Baker & Pritchard (1960), Castagnoli & Pegazzano (1979), Hatzinikolis (1978, 1986b, 1986c, 1987), Hatzinikolis & Panou (1996), Meyer (1979), Mitrofanov & Strunkova (1979), Pritchard & Baker (1958). Type Depository: USNM, No. 1816. 278. Brevipalpus olearius Sayed, 1950b: 1018; Dokki, Giza, Egypt, ex Olea europaea L. (Oleaceae). Species designation: Hystripalpus olearius—Castanoli & Pegazzano (1979), Brevipalpus (H.) olearius—Hatzinikolis (1986b). Redescriptions: Attiah (1956), Baker & Pritchard (1960), Castanoli & Pegazzano (1979), Duzgunes (1965), Hatzinikolis (1986b, 1986c, 1987), Hatzinikolis & Panou (1996), Khosrowshahi & Arbabi (1997), Livschitz & Mitrofanov (1967), Meyer (1979), Mitrofanov & Strunkova (1979), Pritchard & Baker (1958), Wafa et al. (1968/1969), Zaher et al. (1984). Type Depository: Unknown. Note: suspected junior synonym of Brevipalpus oleae Baker. 279. Brevipalpus oleasteri Hatzinikolis & Panou, 1996a: 20, Heraeo, Korinthia, Greece, ex Olea europaea L. var. oleaster Hoffm. (Oleaceae). Type Depository: Collection of the Acarology Laboratory of Agricultural Research Centre of Athens, Greece. 280. Brevipalpus olivicola Pegazzano & Castagnoli, 1972: 139; Alberese, Grosseto, Tuscany, Italy, ex Olea europaea L. (Oleaceae). Species designation: Hystripalpus olivicola—Castagnoli & Pegazzano (1979), Brevipalpus (H.) olivicola—Hatzinikolis (1986b). Redescriptions: Castagnoli & Pegazzano (1979), Hatzinikolis (1986b, 1986c,1987), Hatzinikolis & Panou (1996), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: Collezione di Acari dell’Istituto Sperimentale per la Zoologia Agraria di Firenze, Italy. Note: suspected junior synonym of Brevipalpus oleae Baker. 281. Brevipalpus oncidii Baker, 1949: 361; Jarvisbrook, England, ex Oncidium sp. (Orchidaceae). Species designation: Hystripalpus oncidii—Mitrofanov & Strunkova (1979). Redescriptions: Meyer (1979), Mitrofanov & Strunkova (1979), Pritchard & Baker (1952, 1958). Type Depository: USNM, No. 1817. Suspected senior synonym of: Brevipalpus psychotriae De Leon—Mitrofanov & Strunkova (1979). 282. Brevipalpus oreopanacis De Leon, 1960: 180; Santa María del Oro, Nayarit, Mexico, ex Oreopanax peltatum Linden (Araliaceae). Species designation: Hystripalpus oreopanacis—Mitrofanov & Strunkova (1979). Redescriptions: Baker & Tuttle (1987), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: MCZ. Note: host originally listed as Oreopanax peltata. 283. Brevipalpus ornantis Chaudhri & Akbar, 1985: 60; Mingora, Swat Valley, Pakistan, ex an unidentified plant. Type Depository: Department Entomology, University of Agriculture, Faisalabad, Pakistan. 284. Brevipalpus ornatus Baker & Tuttle, 1987: 41; 19 km S Morelia, Michoacan, Mexico, ex Atriplex sp. (Chenopodiaceae). Type Depository: USNM. 285. Brevipalpus orobos Hasan, Akbar & Ashfaq, 2002: 55; Sheikhupura, Pakistan, ex Helianthus annuus L. (Asteraceae). Type Depository: Acarology Research Laboratory, Department of Entomology, University of Agriculture, Faisalabad, Pakistan. 286. Brevipalpus ortizi Ochoa & Salas, 1989: 25; Las Nubes de Coronado, San Jose, Costa Rica, ex Chamaedorea sp. (Arecaceae).

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Type Depository: Colección de Referencia del Laboratorio de Acarologia, Facultad de Agronomia, Universidad de Costa Rica, Costa Rica. 287. Brevipalpus pachucensis Baker & Tuttle, 1987: 74; 6.4 km east of El Manzana, Pachuca, Hidalgo, Mexico, ex Lupinus sp. (Fabaceae). Type Depository: USNM. 288. Brevipalpus parthenium Baker & Tuttle, 1972: 29; Portal, Arizona, USA, ex Parthenium incanum H.B. & K. (Asteraceae). Species designation: Hystripalpus parthenium—Mitrofanov & Strunkova (1979). Redescriptions: Baker & Tuttle (1987), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: USNM, No. 3434. 289. Brevipalpus patulus Chaudhri & Akbar, 1985: 80; 1.6 km E Bhera, Punjab, Pakistan, ex Whitania somnifera Dun. (Solanaceae). Type Depository: Department Entomology, University of Agriculture, Faisalabad, Pakistan. 290. Brevipalpus perseae De Leon, 1960: 182; Tepic, Nayarit, Mexico, ex Persea hintonii C.K. Allen (Lauraceae). Species designation: Hystripalpus perseae—Mitrofanov & Strunkova (1979). Redescriptions: Baker & Tuttle (1987), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: MCZ. Note: host originally listed as Persea hintoni. 291. Brevipalpus persicanus Chaudhri, Akbar & Rasool, 1974: 51; 1.6 km E Kasur, Punjab, Pakistan, ex Prunus persica (L.) Batsch (Rosaceae). Redescription: Ahmad & Akbar (1984). Type Depository: Department of Entomology, University of Agriculture, Lyallpur, Pakistan. 292. Brevipalpus phoenicis (Geijskes, 1939): 23; The Hague, The Netherlands, ex Phoenix sp. (Arecaceae). Species designations: Tenuipalpus phoenicis—original designation, Brevipalpus phoenicis—Sayed (1946), Brevipalpus pseudocuneatus—Baker (1949), Brevipalpus phoenisis [sic]—Duzgunes (1965). Suspected senior synonym of: Brevipalpus adelos Ahmad & Akbar; Brevipalpus colens Li, Hasan & Ashfaq; Brevipalpus daqingis Ma & Yuan; Brevipalpus hafizii Chaudri & Akbar; Brevipalpus hainanensis Ma & Yuan; Brevipalpus jambhiri Sadana & Balpreet; Brevipalpus nocivus Siddiqui, Chaudhri & Akbar; Brevipalpus portheo Chaudhri & Akbar; Brevipalpus recula Chaudhri—this publication. Redescriptions: Ahmad & Akbar (1984), Attiah (1956), Baker (1949), Baker & Pritchard (1960), Baker & Tuttle (1987), Charanasri et al. (1986), Cromroy (1958), De Leon (1961a, 1965a, 1965b, 1967), Duzgunes (1965), Ehara (1966), Evans et al. (1993, 1998), Gonzalez (1975), Gupta (1986, 1992, 1995), Gupta & Chatterjee (1997), Hatzinikolis (1986b, 1986c, 1987), Hatzinikolis & Panou (1996), Khosrowshahi & Arbabi (1997), Livschitz & Mitrofanov (1967), Livschitz & Salinas (1968), Lo (1969), Lo & Hsia (1968), Manson (1963), Meyer (1970, 1979), Meyer & Ueckermann (1988), Mitrofanov & Strunkova (1979), Nassar & Ghai (1981), Ochoa & Salas (1989), Pritchard & Baker (1952, 1958), Rimando (1962), Sadana (1997), Sayed (1946), Smiley & Gerson (1995), Wafa et al. (1968/ 1969), Welbourn et al. (2003), Womersley (1940), Zaher et al. (1984). Type Depository: Unknown. Synonym: 292a. Brevipalpus deleoni Pritchard & Baker, 1958: 234; Miami, Florida, USA, ex Petrea sp. (Verbenaceae)—De Leon (1961), Gonzalez (1975), Welbourn et al. (2003). Species designations: Brevipalpus delconi [sic]—Gupta (1992). Redescriptions: Gupta (1992), Sadana (1997). Type Depository: USNM, No. 2348. 292b. Brevipalpus mcbridei Baker, 1949: 374; Orlando, Florida, USA, ex English walnut (Juglandaceae).—Pritchard & Baker (1952), Welbourn et al. (2003). Type Depository: USNM, No. 1824. 292c. Brevipalpus papayensis Baker, 1949: 375; Kailua, Oahu, Hawaii, ex papaya (Caricaceae)—Pritchard & Baker (1952), Welbourn et al. (2003). Type Depository: USNM, No. 1825.

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292d. Brevipalpus phoenicoides Gonzalez, 1975: 86, Thailand, ex Citrus reticulata Blanco (Rutaceae)—Evans et al. (1993), Welbourn et al. (2003). Redescription: Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: University of de Chile, Chile. 292e. Brevipalpus yothersi Baker, 1949: 373; Orlando, Florida, USA, ex privet [Ligustrum sp] (Oleaceae)—Pritchard & Baker (1952), Welbourn et al. (2003). Type Depository: USNM, No. 1823. 293. Brevipalpus physali De Leon, 1961a: 47; Veracruz, Veracruz State, Mexico, ex Tridax procumbens L. (Asteraceae). Species designations: Hystripalpus physali—Mitrofanov & Strunkova (1979), Brevipalpus physalis [sic]—Baker & Tuttle (1987). Redescriptions: Baker & Tuttle (1987), De Leon (1967), Evans et al. (1993), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: MCZ. 294. Brevipalpus pini Baker, 1949: 383; Glendale, California, USA, ex ‘‘Monterey pine’’ Pinus radiata (D. Don). (Pinaceae). Species designations Hystripalpus pini—Mitrofanov & Strunkova (1979). Redescriptions: Baker & Tuttle (1964, 1987), Evans et al. (1993), Hatzinikolis (1986c, 1987), Meyer (1979), Mitrofanov et al. (1975), Mitrofanov & Strunkova (1979), Pritchard & Baker (1952, 1958). Type Depository: USNM, No. 1831. Synonym: 294a. Brevipalpus tlaxcensis Baker & Tuttle, 1987: 90; Tlaxco, Tlaxcala, Mexico, ex Pinus sp. (Pinaceae)—Evans et al. (1993). Type Depository: USNM. 295. Brevipalpus piniceltis Baker & Tuttle, 1987: 76; Saltillo, Coahuila, Mexico, ex Celtis reticulata Torr. (Ulmaceae). Type Depository: USNM. 296. Brevipalpus pinicola Pritchard & Baker, 1958: 220; Saint Petersburg, Florida, USA, ex pine (Pinaceae). Species designations: Hystripalpus pinicola—Mitrofanov & Strunkova (1979), Brevipalpus pinicolus [sic]—Meyer (1979). Redescriptions: Meyer (1979), Mitrofanov et al. (1975), Mitrofanov & Strunkova (1979). Type Depository: USNM, No. 2354. 297. Brevipalpus piniwaltheriae Baker & Tuttle, 1987: 76; San Vicente, 208 km N of Pachuca, Hidalgo, Mexico, ex Waltheria sp. (Sterculiaceae). Type Depository: USNM. 298. Brevipalpus plexus Chaudhri, Akbar & Rasool, 1974: 41; 46.4 km S Rawalpindi, Punjab, Pakistan, ex Morus alba L. (Moraceae). Redescription: Ahmad & Akbar (1984). Type Depository: Department of Entomology, University of Agriculture, Lyallpur, Pakistan. 299. Brevipalpus pluchea Baker, Tuttle & Abbatiello, 1975: 12; Topolobampo, Los Mochis, Sinaloa, Mexico, ex Pluchea odorata (L.) Cass. (Asteraceae). Species designation: Brevipalpus plucheae [sic]—Baker & Tuttle (1987). Redescriptions: Baker & Tuttle (1987), Meyer (1979). Type Depository: USNM, No. 3632. 300. Brevipalpus pocillator De Leon, 1961a: 47; Jacotepec, Jalisco, Mexico, ex Verbesina sp. (Asteraceae). Redescriptions: Baker & Tuttle (1987), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: MCZ. 301. Brevipalpus porca Pritchard & Baker, 1958: 208; Bryce Canyon, Utah, USA, ex Phoradendron sp. (Viscaceae) Species designation: Hystripalpus porca—Mitrofanov & Strunkova (1979). Redescriptions: Baker & Tuttle (1964), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: USNM, No. 2351.

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Note: According to Baker & Tuttle (1964) the type host in the original description is incorrect and should be Arceuthobium sp. (Viscaceae). 302. Brevipalpus portalis Baker & Tuttle, 1972: 30; Portal, Arizona, USA, ex Parthenium incanum H.B. & K. (Asteraceae). Species designation: Hystripalpus portalis—Mitrofanov & Strunkova (1979). Redescriptions: Baker & Tuttle (1987), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: USNM, No. 3435. Synonym: 302a. Brevipalpus incanum Baker, Tuttle & Abbatiello, 1975: 19; Fresnillo, Zacatecas, Mexico, ex Parthenium incanum Kunth (Asteraceae)—Baker & Tuttle (1987). Redescription: Meyer (1979). Type Depository: USNM, No. 3639. 303. Brevipalpus portheo Chaudhri & Akbar, 1985: 76; 3.2 km S Karachi, Sind, Pakistan, ex Thuja orientalis L. (Cupressaceae). Type Depository: Department Entomology, University of Agriculture, Faisalabad, Pakistan. Note: suspected junior synonym Brevipapus phoenicis (Geijskes). Synonym: 303a. Brevipalpus portheo Akbar & Khalid, 1999: 161; 2 km S Karachi, Sind, Pakistan, ex Thuja orientalis L. (Cupressaceae).—new synonym. Note: The drawings of the original description of B. portheo Akbar & Khalid (1999) are exactly the same as those of the original description of B. portheo Chaudhri & Akbar (1985). The site of collection of the former species (2 km south of Karachi) is also similar to that of the latter (2 miles south of Karachi). Despite of the minor differences in the collectors and collection dates, they appear to be the same species and therefore B. portheo Akbar & Khalid, 1999 is a primary homonym and junior synonym. Type Depository: Acarology Research Laboratory, Department of Agric. Entomology, University of Agriculture, Faisalabad, Pakistan. Note: suspected junior synonym Brevipapus phoenicis (Geijskes). 304. Brevipalpus potentillae Baker & Tuttle, 1964: 38; Flagstaff, Arizona, USA, ex Potentilla hippiana Lehm. (Rosaceae). Species designation: Hystripalpus potentillae—Mitrofanov & Strunkova (1979). Redescriptions: Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: USNM, No. 3025. 305. Brevipalpus pritchardi Baker & Tuttle, 1964: 34; Palm Canyon, Yuma, Arizona, USA, ex Ditaxis lanceolata (Benth.) Pax & Hofmm (Euphorbiaceae). Species designation: Hystripalpus pritchardi—Mitrofanov & Strunkova (1979). Redescriptions: Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: USNM, No. 3031. 306. Brevipalpus proboscidius De Leon, 1961a: 41; Tuxtla Gutierrez, Chiapas, Mexico, ex Liabum glabrum Hemsl. var. hypoleucum Greene. (Asteraceae). Species designation: Hystripalpus proboscidius—Mitrofanov & Strunkova (1979). Redescriptions: Baker & Tuttle (1987), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: MCZ. 307. Brevipalpus psephoides Pritchard & Baker, 1952: 19; Cedar Grove, California, USA, ex Garrya flavescens S. Watson (Garryaceae). Species designation: Hystripalpus psephoides—Mitrofanov & Strunkova (1979). Redescriptions: Meyer (1979), Mitrofanov & Strunkova (1979), Pritchard & Baker (1958). Type Depository: USNM, No. 1916. 308. Brevipalpus pseudoleptoides De Leon, 1960: 176; Oaxaca, near km 681, route 190, Mexico, ex Dodonaea viscosa Jacq. (Sapindaceae). Species designation: Hystripalpus pseudoleptoides—Mitrofanov & Strunkova (1979). Redescriptions: Baker & Tuttle (1987), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: MCZ.

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309. Brevipalpus pseudolilium Livschitz in Livschitz & Salinas, 1968: 113; Santiago de las Vegas, Cuba, ex Codiaeum variegatum (L.) A. Juss. (Euphorbiaceae). Type Depository: Unknown. 310. Brevipalpus pseudophoenicis Baker, Tuttle & Abbatiello, 1975: 11; Zapotlanejo, Jalisco, Mexico, ex Sida sp. (Malvaceae). Redescriptions: Baker & Tuttle (1987), Meyer (1979). Type Depository: USNM, No. 3628. Note: Locality mentioned in the original description as Zapotleneja. 311. Brevipalpus pseudopini Baker & Tuttle, 1987: 42; 22.4 km E Morelia, Michoacan, Mexico, ex Quercus sp. (Fagaceae). Type Depository: USNM. 312. Brevipalpus pseudopinicolus Baker & Tuttle, 1987: 79; Tlaxco, Tlaxcala, Mexico, ex Juniperus deppeana Steud. (Cupressaceae). Type Depository: USNM. Note: host originally listed as Juniperus mexicanus Spreng. 313. Brevipalpus pseudostriatus Ochoa & Salas, 1989: 26; Universidad de Costa Rica, San Jose, Costa Rica, ex Erythrina sp. (Fabaceae). Type Depository: Colección del Laboratorio de Acarologia, Facultad de Agronomia, Universidad de Costa Rica, Costa Rica. 314. Brevipalpus psilostropheae Pritchard & Baker, 1964: 61; Sells, Arizona, USA, ex Psilostrophe cooperi (Gray) Greene (Asteraceae). Species designations: Hystripalpus psilostropheae—Mitrofanov & Strunkova (1979), Brevipalpus psilostrophes [sic]—Baker & Yin (1988). Redescription: Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: USNM, No. 3034. 315. Brevipalpus psychotriae De Leon, 1961c: 171; Key Largo, Florida, USA, ex Psychotria sp. (Rubiaceae).—Mitrofanov & Strunkova (1979), Evans unpublished. Redescrition: Meyer (1979). Type Depository: MCZ. Note: suspected senior synonym of Brevipalpus oncidii Baker according to Mitrofanov & Strunkova (1979) and Evans unpublished. 316. Brevipalpus punicans Pritchard & Baker, 1952: 24; Tesla, California, USA, ex Gutierrezia californica Torr. & A. Gray (Asteraceae). Species designation: Hystripalpus punicans—Mitrofanov & Strunkova (1979). Redescriptions: Meyer (1979), Mitrofanov & Strunkova (1979), Pritchard & Baker (1958). Type Depository: USNM, No. 1917. 317. Brevipalpus pycnanthemi De Leon, 1961c: 169; Pensacola, North Carolina, USA, ex Pycnanthemum pycnanthemoides Fernald (Lamiaceae). Species designation: Hystripalpus pycnanthemi—Mitrofanov & Strunkova (1979). Redescription: Mitrofanov & Strunkova (1979). Type Depository: MCZ. 318. Brevipalpus qianniunis Ma & Yuan, 1982: 72; Hainan Island, China, ex Pharbitis nil (L.) Choisy (Convolvulaceae). Type Depository: Shanghai College of Agriculture, China. Note: suspected synonym Brevipalpus obovatus Donnadieu. 319. Brevipalpus quercicolus De Leon, 1960: 175; near km 134, of route 71, Mexico, ex Quercus sp. (Fagaceae). Species designation: Hystripalpus quercicolus—Mitrofanov & Strunkova (1979). Redescriptions: Baker & Tuttle (1987), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: MCZ.

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320. Brevipalpus querensis Baker & Tuttle, 1987: 80; 43 km N Oaxaca, Oaxaca State, Mexico, ex Quercus sp. (Fagaceae). Type Depository: USNM. 321. Brevipalpus rapii Hasan, Bashir, Wakil & Afzal, 2003: 353; 10 km N Faisalabad, Punjab, Pakistan, ex Brassica rapa L. (Brassicaceae). Type Depository: Acarology Research Laboratory, Department of Agric. Entomology, University of Agriculture, Faisalabad, Pakistan. Note: suspected junior synonym Brevipalpis edax Baker & Tuttle. These species can not be separated based on descriptions or figures. 322. Brevipalpus recki Livschitz & Mitrofanov, 1967: 14; Ukraine, ex Quercus sp. (Fagaceae), Rubus (Rosaceae) and Inula vulgaris Trevis. (Asteraceae). Species designation: Tauripalpus recki—Mitrofanov (1973). Redescriptions: Hatzinikolis (1987), Meyer (1979), Mitrofanov (1973), Mitrofanov & Strunkova (1979). Type Depository: Unknown. 322a. Brevipalpus recki Livschitz & Mitrofanov, 1968: 416; 2 km from Massandra, along the road to the Crimean Reservation , Cimea, ex Quercus pubescens Will. (Fagaceae). Type Depository: Collections of the Nikita State Botanical Garden, Yalta, Crimea. 323. Brevipalpus recula Chaudhri, 1972b: 63; Multan, Punjab, Pakistan, ex Rat-ka-Raja. Redescriptions: Ahmad & Akbar (1984), Chaudhri et al. (1974), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: Department of Entomology, West Pakistan Agricultural University Lyallpur, Pakistan. Note: suspected junior synonym of Brevipalpus phoenicis (Geijskes), trochanter leg IV should be rexamined. According to Chaudhri et al. (1974) the host plant is Buddleia asiatica Lour. (Buddleiaceae). 324. Brevipalpus religiosae De Leon, 1960: 186; E Morelia near km 270, route 15, Michoacan, Mexico, ex Abies religiosa Lindl. (Pinaceae). Species designation: Hystripalpus religiosae—Mitrofanov & Strunkova (1979). Redescriptions: Baker & Tuttle (1987), Meyer (1979), Mitrofanov et al. (1975), Mitrofanov & Strunkova (1979). Type Depository: MCZ. 325. Brevipalpus rhanis Hasan, Akbar & Ashfaq, 2002: 57; Sheikhupura, Punjab, Pakistan, ex Helianthus annuus L. (Asteraceae) Type Depository: Acarology Research Laboratory, Department of Entomology, University of Agriculture, Faisalabad, Pakistan. 326. Brevipalpus rica Chaudhri, 1972b: 56; 1.6 km E Muzaffargarh, Punjab, Pakistan, ex Tagetes tenuifolia H.B. & K. (Asteraceae). Species designation: Hystripalpus rica—Mitrofanov & Strunkova (1979). Redescriptions: Ahmad & Akbar (1984), Chaudhri et al. (1974), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: Department of Entomology, West Pakistan Agricultural University Lyallpur, Pakistan. Note: host originally listed as Tegetes tenuifolia. 327. Brevipalpus rostratus De Leon, 1961a: 41; San Cristobal, Chiapas, Mexico, ex Myrica mexicana Humb. Bonpl. & Willd. (Myricaceae). Species designation: Hystripalpus rostratus—Mitrofanov & Strunkova (1979). Redescriptions: Baker & Tuttle (1987), Meyer (1979), Papaioannou-Souliotis (1986). Type Depository: MCZ. 328. Brevipalpus rotai (Castagnoli & Pegazzano, 1979): 285; Massafra, Taranto, Italy, ex Olea europaea L. (Oleaceae). Species designation: Hystripalpus rotai—original designation, Brevipalpus rotai—Meyer (1979). Redescriptions: Hatzinikolis (1986b), Hatzinikolis & Panou (1996), Meyer (1979). Type Depository: Collezione dell’Istituto Sperimentale per la Zoologia Agraria di Firenze, Italy. 329. Brevipalpus rubus Baker & Tuttle, 1987: 82; 19 km E Morelia, Michoacan, Mexico, ex Rubus cordifolius Weihe & Nees (Rosaceae). Type Depository: USNM.

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Note: host originally listed as Rubus cortifolius. 330. Brevipalpus ruelliae Baker & Tuttle, 1987: 107; Torreon, Coahuila, Mexico, ex Ruellia nudiflora (Engelm. & A. Gray) Urb. (Acanthaceae). Type Depository: USNM. 331. Brevipalpus rugosus De Leon, 1960: 184; Nochitlan, Oaxaca, Mexico, ex Quercus magnoliaefolia Nee (Fagaceae). Species designation: Hystripalpus rugosus—Mitrofanov & Strunkova (1979). Redescriptions: Baker & Tuttle (1987), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: MCZ. 332. Brevipalpus rugulosus Chaudhri, Akbar & Rasool, 1974: 45; Jahangir Tomb, Lahore, Punjab, Pakistan, ex an unidentified plant. Redescriptions: Ahmad & Akbar (1984), Sadana (1997). Type Depository: Department of Entomology, University of Agriculture, Lyallpur, Pakistan. 333. Brevipalpus russulus (Boisduval, 1867): 89; France, ex Mammillaria sp., Echinocactus sp. (Cactaceae) and related genera. Species designations: Acarus russulus—original designation, Brevipalpus russulus—Oudemans (1938), Hystripalpus russulus—Mitrofanov & Strunkova (1979). Redescriptions: Baker (1949), Ehara (1968), Geijskes (1939), Hatzinikolis (1986c, 1987), Livschitz & Mitrofanov (1967), Meyer (1979), Mitrofanov & Strunkova (1979), Ochoa & Salas (1989), Oudemans (1929, 1938), Pritchard & Baker (1952, 1958), Sayed (1946). Type Depository: Unknown. Synonym: 333a. Tenuipalpus cactorum Oudemans, 1929: 483; Germany, ex cacti (Cactaceae)—Pritchard & Baker (1952). Species designation: Brevipalpus cactorum—Sayed (1946). Redescriptions: Andre (1953), Geijskes (1939), Rech (1951), Sayed (1946). Type Depository: Unknown. 334. Brevipalpus sahii Hasan, Wakil & Bashir, 2004: 104; 1 km E Bhera, Punjab, Pakistan, ex Capsicum frutescens L. (Solanaceae). Type Depository: Acarology Research Laboratory, Department of Agricultural Entomology, University of Agriculture, Faisalabad, Pakistan. Note: this especies need to be redescribed befor it can be properly placed. Host originally listed as Capcicum frutiscens. 335. Brevipalpus salasi Ochoa, 1986: 155; Guadarrama de Desamparados, San Jose, Costa Rica, ex Spondias purpurea L. (Anacardiaceae). Redescription: Ochoa & Salas (1989). Type Depository: Coleccion de Referencia, Laboratorio Acarologia, Facultad de Agronomia, Universidad de Costa Rica. 336. Brevipalpus salicis Baker & Tuttle, 1964: 43; Yuma, Arizona, USA, ex Salix gooddingii Ball. (Salicaceae). Species designation: Hystripalpus salicis—Mitrofanov & Strunkova (1979). Redescriptions: Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: USNM, No. 3029. 337. Brevipalpus salix Baker & Tuttle, 1987: 83; 9.6 km N Oaxaca, Oaxaca State, Mexico, ex Salix sp. (Salicaceae). Type Depository: USNM. 338. Brevipalpus salignae Meyer, 1979: 94; 2 km from Thomas Baines Nature Reserve, Grahamstown, Cape Province, South Africa, ex Buddleja saligna Willd. (Buddlejaceae). Type Depository: SANC. Note: host originally listed as Buddleia saligna. 339. Brevipalpus salpizo Hasan, Akbar & Ashfaq, 2002: 53; 4 km E Sheikhupura, Pakistan, ex Helianthus annuus L. (Asteraceae). Type Depository: Acarology Research Laboratory, Department of Entomology, University of Agriculture, Faisalabad, Pakistan.

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340. Brevipalpus salviae McGregor, 1949: 21; Santa Paula, California, USA, ex Salvia sp. (Lamiaceae). Species designation: Hystripalpus salviae—Mitrofanov & Strunkova (1979). Redescriptions: Baker (1949), Meyer (1979), Mitrofanov & Strunkova (1979), Pritchard & Baker (1952, 1958). Type Depository: USNM, No. 1529. 341. Brevipalpus salviella Pritchard & Baker, 1952: 35; Elsinore, California, USA, ex Salvia mellifera Greene (Lamiaceae). Redescriptions: Meyer (1979), Mitrofanov & Strunkova (1979), Pritchard & Baker (1958). Type Depository: USNM, No. 1926 342. Brevipalpus sayedi Baker, 1949: 367; Monticello, Florida, USA, ex hickory (Carya sp., Juglandaceae). Species designation: Hystripalpus sayedi—Mitrofanov & Strunkova (1979). Redescriptions: Hatzinikolis (1986c, 1987), Meyer (1979), Mitrofanov & Strunkova (1979), Pritchard & Baker (1958), Thewke & Enns (1970). Type Depository: USNM, No. 1820. 343. Brevipalpus selas Pritchard & Baker, 1952: 34; Cordelia (Green Valley), California, USA, ex Photinia arbutifolia Lindl. (Rosaceae). Species designation: Hystripalpus selas—Mitrofanov & Strunkova (1979). Redescriptions: Meyer (1979), Mitrofanov & Strunkova (1979), Pritchard & Baker (1958). Type Depository: USNM, No. 1921. 344. Brevipalpus serratus De Leon, 1960: 176; Tuxtla Gutierrez, Chiapas, Mexico, ex an unidentified tree. Species designation: Hystripalpus serratus—Mitrofanov & Strunkova (1979). Redescriptions: Baker & Tuttle (1987), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: MCZ. 345. Brevipalpus shafii Ahmad & Akbar, 1984: 42; 6.4 km N Faisalabad, Pakistan, ex Solanum melongena L. (Solanaceae). Type Depository: Acarology Research Laboratory, Department of Entomology, University of Agriculture, Faisalabad, Pakistan. 346. Brevipalpus sida Baker, Tuttle & Abbatiello, 1975: 8; Tequila, Jalisco, Mexico, ex Sida sp. (Malvaceae). Species designation: Brevipalpus sidae [sic]—Baker & Tuttle (1987). Redescriptions: Baker & Tuttle (1987), Meyer (1979). Type Depository: USNM, No. 3626. 347. Brevipalpus similis Baker & Tuttle, 1987: 85; Zempoala, Mexico, ex Abies religiosa Lindl. (Pinaceae). Type Depository: USNM. Note: There are at least two places in Mexico called Zempoala, one in the Hidalgo and other in the Veracruz, but the original description does not indicate the state. 348. Brevipalpus simlensis De Leon, 1967: 39; Asa Wright Nature Center (= Simla), Trinidad, Trinidad and Tobago, ex Convolvulus umbellatus Sessé & Moc. (Convolvulaceae). Redescription: Meyer (1979). Type Depository: MCZ. Note: host originally listed as Convolvulus umbellata. 349. Brevipalpus sipho Akbar & Aheer, 1990: 257; 2 km E Garah More, Pakistan, ex Luffa acutangula (L.) Roxb. (Cucurbitaceae). Type Depository: Acarology Research Laboratory, Department of Agric. Entomology, University of Agriculture, Faisalabad, Pakistan. 350. Brevipalpus solidus Chaudhri & Akbar, 1985: 65; 4.8 km N Karachi, Sind, Pakistan, ex an unidentified plant. Type Depository: Department Entomology, University of Agriculture, Faisalabad, Pakistan. Synonym: 350a. Brevipalpus solidus Akbar & Khalid, 1999: 156; 3 km N Karachi, Sind, Paquistão, ex an unidentified plant—new synonomy. Note: The drawings of the original description of B. solidus Akbar & Khalid (1999) are exactly the same as those of the original description of B. solidus Chaudhri & Akbar (1985). The site of Collection of the former species (3 km

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north of Karachi) is also similar to that of the latter (3 miles north of Karachi). Despite of the minor differences in the collectors and collection dates, they appear to be the same species and therefore B. solidus Akbar & Khalid, 1999 is a primary homonym and junior synonym. Type Depository: Acarology Research Laboratory, Department of Agric. Entomology, University of Agriculture, Faisalabad, Pakistan. 351. Brevipalpus spaticus Hatzinikolis & Emmanouel, 1991: 17; Spata, Attiki, Greece, ex Olea europaea L. (Oleaceae). Redescription: Hatzinikolis & Panou (1996). Type Depository: Collection of the Acarology Laboratory of the Agricultural Research Centre of Athens, Greece. 352. Brevipalpus spatulatus Baker & Tuttle, 1987: 107; 91 km N Matehuala, San Luis Potosi, Mexico, ex Parthenium incanum Kunth (Asteraceae). Type Depository: USNM. 353. Brevipalpus spitzeri Baker & Tuttle, 1987: 85; Mexico, intercepted at Nogales, Arizona quarantine station, Arizona, USA, ex cactus (Cactaceae). Type Depository: USNM. 354. Brevipalpus spurcus Chaudhri, 1972b: 61; 3.2 km W Shakargarh, Punjab, Pakistan, ex Ocimum basilicum L. (Lamiaceae). Redescriptions: Ahmad & Akbar (1984), Chaudhri et al. (1974), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: Department of Entomology, West Pakistan Agricultural University Lyallpur, Pakistan. 355. Brevipalpus stenolobae Baker & Tuttle, 1987: 86; 91 km N Matehuala, San Luis Potosi, Mexico, ex Viguiera stenoloba S.F. Blake (Asteraceae). Type Depository: USNM. Note: host originally listed as Viquiera stenoloba Blake. 356. Brevipalpus stipae Baker & Tuttle, 1987: 87; Tlaxcala, Tlaxcala State, Mexico, ex Stipa ichu (Ruíz & Pav.) Kunth (Poaceae). Type Depository: USNM. 357. Brevipalpus striatus De Leon, 1960: 183; Trinidad, Chiapas, Mexico, ex "Mata gusano" (a large tree). Species designation: Hystripalpus striatus—Mitrofanov & Strunkova (1979). Redescriptions: Baker &Tuttle (1987), Evans et al. (1993), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: MCZ. 358. Brevipalpus styxus De Leon, 1961c: 173; Delray Beach, Florida, USA, ex Solanum bahamense L. (Solanaceae). Species designation: Hystripalpus styxus—Mitrofanov & Strunkova (1979). Redescriptions: De Leon (1967), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: MCZ. 359. Brevipalpus tagetinae Baker & Tuttle, 1987: 108; replacement to Brevipalpus psilostrophe Baker, Tuttle & Abbatiello (1975), name pre-occuped by B. psilostropheae Pritchard & Baker (1964). (Article 58.7 4th Edition International Code of Zoological Nomenclature). Synonym: 359a. Brevipalpus psilostrophe Baker, Tuttle & Abbatiello, 1975:14; Chihuahua, Chihuahua State, Mexico, ex Psilostrophe tagetina Greene (Asteraceae)—Baker & Tuttlle (1987). Species designations: Brevipalpus psilotrophe and Brevipalpus psilostrophe—original designation. Redescription: Meyer (1979). Type Depository: USNM, No. 3634. 360. Brevipalpus tarus Gonzalez, 1975: 88; Pica oasis, Tarapaca, Chile, ex an unidentified ornamental plant. Redescription: Meyer (1979). Type Depository: Acarological Collection Universidad de Chile, Chile. 361. Brevipalpus tepicbutilonae Baker & Tuttle, 1987: 88; Tepic, Nayarit, Mexico, ex Abutilon sp. (Malvaceae). Type Depository: USNM.

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362. Brevipalpus tepicensis Baker & Tuttle, 1987: 88; Tepic, Nayarit, Mexico, ex Hyptis sp. (Lamiaceae). Type Depository: USNM 363. Brevipalpus testudinalis De Leon, 1960: 186; San Blas, Nayarit, Mexico, ex an unidentified tree. Species designation: Hystripalpus testudinalis—Mitrofanov & Strunkova (1979). Redescriptions: Baker & Tuttle (1987), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: MCZ. 364. Brevipalpus theae Ma & Yuan, 1977: 121; Guangzhou, Guangdong, China, ex Thea sinensis L. (Theaceae). Type Depository: Unknown. 365. Brevipalpus thelycraniae Livschitz & Mitrofanov, 1967: 6; Ukraine, ex Cornus sanguinea L. (Cornaceae). Species designation: Cenopalpus (Cenopalpus) thelicraniae [sic]—Mitrofanov & Strunkova (1979). Redescription: Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: Unknown. 366. Brevipalpus tiliae De Leon, 1961c: 172; Erwin, Tennessee, USA, ex Tilia heterophylla Vent. (Tiliaceae). Species designation: Hystripalpus tiliae—Mitrofanov & Strunkova (1979). Redescriptions: Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: MCZ. 367. Brevipalpus tinsukiaensis Sadana & Gupta, 1983: 4; Tinsukia, Assam, India, ex Ocimum sanctum L. (Lamiaceae). Redescriptions: Sadana (1997). Type Depository: Unknown. Note: suspected junior synonym Brevipalpus obovatus Donnadieu. 368. Brevipalpus tiwanensis Hasan, Wakil & Bashir, 2004: 105; Mitha-Tiwana Forest, Punjab, Pakistan, ex an unidentified plant. Type Depository: Acarology Research Laboratory, Department of Agric. Entomology, University of Agriculture, Faisalabad, Pakistan. 369. Brevipalpus trinidadensis Baker, 1949: 381; Saint Augustine, Trinidad, ex Lantana sp. (Verbenaceae). Species designation: Hystripalpus trinidadensis—Mitrofanov & Strunkova (1979). Redescriptions: Baker & Tuttle (1987), De Leon (1961a, 1965a, 1965b, 1967), Evans et al. (1993), Meyer (1979), Mitrofanov & Strunkova (1979), Ochoa & Salas (1989), Pritchard & Baker (1958). Type Depository: USNM, No. 1829. Note: De Leon (1967) suspected Brevipalpus trinidadensis to be a junior synonym of Brevipalpus californicus (Banks), but we agree with Ochoa & Salas (1989) and Evans et al. (1993) who considered it to be a valid species. 370. Brevipalpus tuberellus De Leon, 1960: 177; near Antiguo Morelos, Tamaulipas, Mexico, ex Phoebe tampicensis Mez (Lauraceae). Species designations: Brevipalpus tubercellus [sic]—Baker et al. (1975); Hystripalpus tuberellus—Mitrofanov & Strunkova (1979). Redescriptions: Baker & Tuttle (1987), Baker et al. (1975), De Leon (1967), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: MCZ. 371. Brevipalpus turrialbensis Manson, 1963a: 220; Santa Cruz de Turrialba, Cartago, Costa Rica, ex an unidentified plant. Species designation: Hystripalpus turrialbensis—Mitrofanov & Strunkova (1979). Redescriptions: Meyer (1979), Mitrofanov & Strunkova (1979), Ochoa & Salas (1989). Type Depository: USNM, No. 2889. 372. Brevipalpus utriculus Chaudhri & Akbar, 1985: 70; 3.2 km S Sargodha, Pakistan, ex Malvastrum sp. (Malvaceae). Type Depository: Department Entomology, University of Agriculture, Faisalabad, Pakistan. 373. Brevipalpus variolatus De Leon, 1960: 184; Trinidad, Chiapas, Mexico, ex Ceiba acuminata Rose (Malvaceae). Species designation: Hystripalpus variolatus—Mitrofanov & Strunkova (1979).

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Redescriptions: Baker & Tuttle (1987), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: MCZ. 374. Brevipalpus ventus Chaudhri & Akbar, 1985: 67; Haripur, Pakistan, ex an unidentified plant. Type Depository: Department Entomology, University of Agriculture, Faisalabad, Pakistan. 375. Brevipalpus venutus Baker & Tuttle, 1987: 92; 20 km N Pachuca, Hidalgo, Mexico, ex Haplopappus venutus (H.B. & K.) Blake (Asteraceae). Type Depository: USNM 376. Brevipalpus verbenae Baker & Tuttle, 1987: 109; Chihuahua, Chihuahua State, Mexico, ex Verbena sp. (Verbenaceae). Type Depository: USNM. 377. Brevipalpus viguiera Baker, Tuttle & Abbatiello, 1975: 16; Los Mochis, Sinaloa, Mexico, ex Viguiera sp. (Asteraceae). Species designations: Brevipalpus viquierae [sic]—Baker & Tuttle (1987), Brevipalpus viquiera [sic]—Evans et al. (1993, 1998). Redescriptions: Baker & Tuttle (1987), Evans et al. (1993, 1998), Meyer (1979). Type Depository: USNM, No. 3635. 378. Brevipalpus xystus Pritchard & Baker, 1958: 220; Miami, Florida, USA, ex Ichthyomethia piscipula Hitchc. (Fabaceae). Species designation: Hystripalpus xystus—Mitrofanov & Strunkova (1979). Redescriptions: Meyer (1979), Mitrofanov & Strunkova (1979), Thewke & Enns (1970). Type Depository: USNM, No. 2355. 379. Brevipalpus zempoalensis Baker & Tuttle, 1987: 93; Zempoala, Mexico, ex Abies religiosa Lindl. (Pinaceae). Type Depository: USNM. Note: There are at least two places in Mexico called Zempoala, one in Hidalgo and other in Veracruz, but the original description, does not indicate the state. 380. Brevipalpus zhenzhumeis Ma & Yaun, 1982: 3; Chongqing, Sichuan, China, ex Sorbaria arborea Schneider (Rosaceae). Type Depository: Shanghai College of Agriculture, China. 381. Brevipalpus ziaii Chaudhri & Akbar, 1985: 73; 4.8 km N Faisalabad, Pakistan, ex Gossypium hirsutum L. (Malvaceae). Type Depository: Department Entomology, University of Agriculture, Faisalabad, Pakistan. 382. Brevipalpus zinniae Baker & Tuttle, 1987: 109; El Sueco, Rancho Grande, Chihuahua, México, ex Zinnia acerosa (DC.) Gray (Asteraceae). Type Depository: USNM.

Capedulia Meyer 1979 Capedulia Meyer, 1979: 109. Type species: Capedulia calendulae Meyer, 1979: 109; Gerson & Meyer, 1980: 9; Ueckermann & Tiedt, 1999: 201. 383. Capedulia calendulae Meyer, 1979: 109; Klip River (Graaff-Reinet—Pearston road), Cape Province, South Africa, ex Osteospermum calendulaceum L. (Asteraceae). Redescription: Ueckermann & Tiedt (1999). Type Depository: SANC. 384. Capedulia maritima Gerson & Meyer, 1980: 9; Dor, Coastal Plain, Israel, ex roots of Limonium meyeri Kuntze (Plumbaginaceae). Redescription: Ueckermann & Tiedt (1999).

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Type Depository: Collection of the Department of Entomology, Faculty of Agriculture, Rehovot, Israel. 385. Capedulia prasadi Ueckermann & Tiedt, 1999: 204; Western Cape, South Africa, ex Aspalathus linearis (N.L. Burm.) (Fabaceae). Type Depository: SANC. Note: Mites were found tightly packed in small galls on the stems and branches on the shrub. All the gall are situated under tube-like plant tissue, running along the stems and branches. 386. Capedulia xeroclei Ueckermann & Tiedt, 1999: 208; Gobabeb, Namibia, ex Trianthema sp. (Aizoaceae). Type Depository: SANC.

Cenopalpus Pritchard & Baker 1958 Cenopalpus Pritchard & Baker, 1958: 190. Type species: Brevipalpus spinosus (Donnadieu, 1875): 114; Akbar & Chaudhri, 1985: 270; Bagdasarian, 1962: 54; Baker & Pritchard, 1960: 561; Chaudhri, 1971a: 71; Chaudhri et al., 1974: 20; Dosse, 1972: 5; Dosse, 1975: 732; Düzgünes, 1965: 124; Düzgünes, 1967: 90; Ehara, 1966: 2; Hatzinikolis, 1987: 54; Hatzinikolis & Emmanouel, 1987: 4; Hatzinikolis et al. 1999: 131; Hatzinikolis et al. 2001: 35; Ma & Li, 1984: 325; Manson, 1963: 222; Mitrofanov, 1973a: 509; Papaioannou-Souliotis, 1986: 133; Pegazzano, 1971: 740; Rimando, 1962: 37; Wainstein, 1960: 236; Welbourn et al. 2003: 108; Zaher & Yousef, 1969: 278. Cenopalpus (Cenopalpoides) Mitrofanov, 1973a: 510; Mitrofanov & Strunkova, 1979: 66. Cenopalpus (Cenopalpus) Mitrofanov, 1973: 510, Castagnoli, 1987: 107. Pritchardipalpus Mitrofanov, 1973a: 509. Type species: Brevipalpus pterinus Pritchard & Baker, 1958; synonymy according to Meyer (1979) who considered Cenopalpus a junior synonym of Brevipalpus. Note: Meyer (1979) and Sepasgosarian (1983) considered Cenopalpus to be a junior synonym of Brevipalpus, whereas Baker & Tuttle (1987) and Welbourn et al. (2003) considered them as different genera. 387. Cenopalpus abaii Khosrowshahi & Arbabi, 1997:7; Fars Province, Iran, ex Quercus sp. (Fagaceae). Type Depository: TUPH, No. T-1022. 388. Cenopalpus aratus Chaudhri, 1971a: 71; Samli, Murree Hills, Punjab, Pakistan, ex Solanum tuberosum L. (Solanaceae). Species designation: Brevipalpus aratus—Meyer (1979). Redescriptions: Akbar & Chaudhri (1985), Chaudhri et al. (1974), Meyer (1979). Type Depository: Department of Entomology, West Pakistan Agricultural University Lyallpur, Pakistan. 389. Cenopalpus arbuti Hatzinikolis & Emmanouel, 1987: 14; Marmari, Evvia, Greece, ex (strawberry tree) Arbutus unedo L. (Ericaceae). Redescriptions: Hatzinikolis (1987), Hatzinikolis et al. (1999). Type Depository: Collection of the Acarology Laboratory of Agricultural Research Centre of Athens, Greece. 390. Cenopalpus bagdasariani (Livschitz & Mitrofanov, 1970): 789; Tajikistan, ex Fraxinus sp. (Oleaceae). Species designations: Brevipalpus bagdasariani Livschitz & Mitrofanov (1970)—original designation, Cenopalpus (Cenopalpus) bagdasariani—Mitrofanov & Strunkova (1979). Redescriptions: Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: Unknown. 391. Cenopalpus bakeri Düzgünes, 1967: 91; Ankara, Turkey, ex Crataegus oxyacantha L. (Rosaceae). Species designation: Brevipalpus bakeri—Meyer (1979). Redescriptions: Ghai & Shenhmar (1984), Hatzinikolis (1987), Hatzinikolis & Emmanouel (1987), Meyer (1979), Khosrowshahi & Arbabi (1997), Mitrofanov & Strunkova (1979). Type Depository: Department of Plant Protection, Faculty of Agriculture, University Ankara, Turkey. Synonym: 391a. Brevipalpus bakeriana Ghai & Shenhmar, 1984: 127; proposed as replacement name for Cenopalpuss bakeri in the process of transferring the species to Brevipalpus. 392. Cenopalpus brachypalpus Hatzinikolis, Panou & Papadoulis, 1999: 277; Levadia, Greece, ex Rubus sp. (Rosaceae). Type Depository: Acari Collection of the Acarology Laboratory of Agricultural University of Athens, Greece. 393. Cenopalpus capacis Chaudhri, 1971a: 72; 1.6 km SW Naran, Kaghan Valley, Pakistan, ex Pinus wallichiana A.B. Jacks (Pinaceae).

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Redescriptions: Akbar & Chaudhri (1985), Chaudhri et al. (1974), Meyer (1979). Type Depository: Department of Entomology, West Pakistan Agricultural University Lyallpur, Pakistan. Note: host originally listed as Pinus willichiana. 394. Cenopalpus capensis (Meyer, 1979): 92—new combination; Stanford, Cape Province, South Africa, ex Elytropappus gnaphaloides L. (Asteraceae). Species designation: Brevipalpus capensis—original designation. Type Depository: SANC. 395. Cenopalpus carpini (Livschitz & Mitrofanov, 1967): 20; Krymskaya, Ukraine, ex Carpinus orientalis Mill. (Betulaceae). Species designations: Brevipalpus carpini—Meyer (1979), Cenopalpus carpini—Hatzinikolis & Emmanouel (1987), Cenopalpus (Cenopalpus) carpini—Mitrofanov & Strunkova (1979). Redescriptions: Hatzinikolis (1987), Hatzinikolis & Emmanouel (1987), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: Unknown. 396. Cenopalpus chitraliensis Akbar & Chaudhri, 1985: 276; Chitral, Pakistan, ex Pyrus malus L. (Rosaceae). Redescription: Chaudhri & Akbar (1985). Type Depository: Acarology Research Laboratory, Department of Agric. Entomology, University of Agriculture, Faisalabad, Pakistan. 397. Cenopalpus crataegi Dosse, 1971: 580; Karadj, Iran, ex Crataegus oxyacantha L. (Rosaceae). Species designation: Brevipalpus crataegi—Meyer (1979). Redescriptions: Khosrowshahi & Arbabi (1997), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: Unknown. 398. Cenopalpus creticus Hatzinikolis, Papadoulis & Panou, 1999: 134; Chani Kokkini, Heraklion, Crete, Greece, ex Salvia officinalis L. (Lamiaceae). Type Depository: Acari Collection of the Laboratory of Agricultural, Zoology and Entomology, of Agricultural University of Athens, Greece. 399. Cenopalpus dignus Akbar & Chaudhri, 1985: 278; Charrapani, Murree Hills, Punjab, Pakistan, ex Musa paradisiaca L. (Musaceae). Redescription: Chaudhri & Akbar (1985). Type Depository: Acarology Research Laboratory, Department of Agric. Entomology, University of Agriculture, Faisalabad, Pakistan. Note: Host originally listed as banana Musa paradisica. 400. Cenopalpus eriobotryi Hatzinikolis, 1969: 57; Raphina, Attica, Greece, ex Eriobotrya japonica Lindl. (Rosaceae). Species designation: Brevipalpus eriobotryi—Meyer (1979). Redescriptions: Hatzinikolis (1987), Hatzinikolis & Emmanouel (1987), Hatzinikolis et al. (2001), Meyer (1979). Type Depository: Collection of the Benaki Phytopathological Institute, Greece. 401. Cenopalpus evini Khosrowshahi, 1991:7; Saveh, Central Provinc, Iran, ex Populus alba L. (Salicaceae). Redescription: Khosrowshahi & Arbabi (1997). Type Depository: TUPH, No. T-108. Note: host originally listed as Populus albae. 402. Cenopalpus favosus Chaudhri, 1971a: 74; Charrapani, Murree Hills, Punjab, Pakistan, ex Pinus sp. (Pinaceae). Species designation: Brevipalpus favosus—Meyer (1979). Redescriptions: Ahmad & Akbar (1984), Akbar & Chaudhri (1985), Chaudhri et al. (1974), Meyer (1979). Type Depository: Department of Entomology, West Pakistan Agricultural University Lyallpur, Pakistan. 403. Cenopalpus halperini Castagnoli, 1987:107; Shaar Hagai, Israel, ex Pinus halepensis Mill. (Pinaceae). Species designation: Cenopalpus (Cenopalpus) halperini—original designation. Type Depository: ISZA.

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404. Cenopalpus haqii Akbar & Chaudhri, 1985: 272; Mastung, Quetta Valley, Punjab, Pakistan, ex Pyrus malus L. (Rosaceae). Redescription: Chaudhri & Akbar (1985). Type Depository: Acarology Research Laboratory, Department of Agric. Entomology, University of Agriculture, Faisalabad, Pakistan. 405. Cenopalpus hederae Papaioannou-Souliotis, 1986: 17; Marathon, Attica, Greece, ex Hedera sp. (Araliaceae) and Cupressus sempervirens L. (Cupressaceae). Type Depository: Acarology Laboratory at the Benaki Phytopathological Institute, Greece. 406. Cenopalpus homalos Akbar & Chaudhri, 1985: 280; Orak Valley, Quetta, Punjab, Pakistan, ex Prunus amygdalus Stokes (Rosaceae). Species designation: Cenopalpus homalus [sic]—Hatzinikolis et al. (1999). Redescriptions: Chaudhri & Akbar (1985), Hatzinikolis et al. (1999). Type Depository: Acarology Research Laboratory, Department of Agric. Entomology, University of Agriculture, Faisalabad, Pakistan. 407. Cenopalpus irani Dosse, 1971: 579; Ainestan, Karadjtal, Iran, ex apple [Malus sp.] (Rosaceae). Species designation: Brevipalpus irani—Meyer (1979). Redescriptions: Khosrowshahi & Arbabi (1997), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: Unknown. 408. Cenopalpus japonicus Hasan, Akbar & Khalid, 2001:1; 1 km E Attock, Punjab, Pakistan, ex Eriobotrya japonica Lindl. (Rosaceae). Type Depository: Acarology Research Laboratory, Department of Entomology, University of Agriculture, Faisalabad, Pakistan. 409. Cenopalpus kritos Hasan, Ashfaq, Li, Wakil & Bashir, 2004: 49; 2 km N Murree, Punjab, Pakistan, ex Pyrus malus L. (Rosaceae). Type Depository: Acarology Research Laboratory, Department of Agric. Entomology, University of Agriculture, Faisalabad, Pakistan. 410. Cenopalpus lanceolatisetae (Attiah, 1956): 436; “Lower and Upper Egypt”, Egypt, ex Prunus domestica L., Prunus armeniaca L., Pyrus malus L. and Pyrus communis L. (Rosaceae). Species designations: Brevipalpus lanceolatisetae—original designation, Cenopalpus lanceolatisetae—Pritchard & Baker (1958), Cenopalpus (Cenopalpus) lanceolatisetae—Mitrofanov & Strunkova (1979). Redescriptions: Al-Gboory (1987), Bagdasarian (1962), Baker & Pritchard (1960), Hatzinikolis (1987), Hatzinikolis & Emmanouel (1987), Khosrowshahi & Arbabi (1997), Meyer (1979), Mitrofanov & Strunkova (1979), Pritchard & Baker (1958), Wafa et al. (1968/1969), Wainstein (1960), Zaher & Yousef (1969), Zaher et al. (1984). Type Depository: Unknown. 411. Cenopalpus limbatus Akbar & Chaudhri, 1985: 280; Garm Chashma, Chitral Valley, Pakistan, ex an unidentified plant. Redescription: Chaudhri & Akbar (1985). Type Depository: Acarology Research Laboratory, Department of Agric. Entomology, University of Agriculture, Faisalabad, Pakistan. 412. Cenopalpus lineola (Canestrini & Fanzago, 1876): 105; Trentino, Trento, Italy, ex pine (Pinaceae). Species designations: Tetranychus lineola—original designation, Brevipalpus lineola—Livschitz & Mitrofanov (1967), Caligonus lineola—Canestrini & Fanzago (1877), Cenopalpus lineola—Pritchard & Baker (1958), Cenopalpus (Cenopalpoides) lineola—Mitrofanov (1973), Tenuipalpus lineola—Berlese (1886). Redescriptions: Bagdasarian (1962), Baker & Pritchard (1952), Berlese (1886, 1887), Canestrini & Fanzago (1877), Donnadieu (1875), Ehara (1966), Gutierrez et al. (1989), Hatzinikolis (1987), Hatzinikolis & Emmanouel (1987), Hatzinikolis et al. (1999), Livschitz & Mitrofanov (1967), Meyer (1979), Mitrofanov (1973), Mitrofanov et al. (1975), Mitrofanov & Strunkova (1979), Pegazzano (1970), Pritchard & Baker (1958), Reck (1951), Rimando (1962), Wainstein (1960), Zaher & Yousef (1969). Type Depository: Unknown.

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Synonym: 412a. Brevipalpus asyntactus Baker & Pritchard, 1952: 612; Holland, intercepted at Hoboken, New Jersey, USA, ex pine forest litter with rhododendrons (Ericaceae).—Pritchard & Baker (1958). Type Depository: USNM, No. 1978. 412b. Brevipalpus kalandadzei Reck, 1951: 295; Georgia, ex Pinus hamata Roezl. (Pinaceae).—Pritchard & Baker (1958). Type Depository: Unknown. 413. Cenopalpus longirostris (Livschitz & Mitrofanov, 1967): 17; Ukraine, ex Quercus sp. (Fagaceae). Species designations: Brevipalpus longirostris Livschitz & Mitrofanov (1967)—original designation, Cenopalpus (Cenopalpus) longirostris—Mitrofanov & Strunkova (1979). Redescriptions: Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: Unknown. 414. Cenopalpus mespili (Livschitz & Mitrofanov, 1967): 21; Ukraine, ex Mespilus sp. (Rosaceae) and apple (Rosaceae). Species designations: Brevipalpus mespili—original designation, Cenopalpus mespili—Hatzinikolis & Emmanouel (1987), Cenopalpus (Cenopalpus) mespili—Mitrofanov & Strunkova (1979). Redescriptions: Hatzinikolis & Emmanouel (1987), Hatzinikolis (1987), Meyer (1979), Mitrofanov & Strunkova (1979). Types Depository: Unknown. 415. Cenopalpus meyerae Khosrowshahi, 1991: 6; Tehran Province, Iran, ex Platanus sp. (Platanaceae). Resdescription: Khosrowshahi & Arbabi (1997). Type Depository: TUPH, No. 612. 416. Cenopalpus mughalii Akbar & Aheer, 1990: 259, 10 km N Pakpattan, Pakistan, ex Momordica charantia L. (Cucurbitaceae). Type Depository: Acarology Research Laboratory, Department of Agric. Entomology, University of Agriculture, Faisalabad, Pakistan. 417. Cenopalpus musai Dosse, 1975: 732; Fanar, Lebanon, ex Rubus sp. (Rosaceae). Species designation: Brevipalpus musai—Meyer (1979). Redescription: Meyer (1979). Type Depository: Unknown. 418. Cenopalpus natalensis (Lawrence, 1943): 45; Karkloof, near Pietermaritzburg, Natal, South Africa, ex Halleria lucida L. (Scrophulariaceae). Species designations: Tenuipalpus natalensis—original designation, Brevipalpus natalensis—Baker (1949), Baker & Pritchard (1952c), Cenopalpus natalensis—Pritchard & Baker (1958), Cenopalpus (Cenopalpus) natalensis—Mitrofanov & Strunkova (1979). Redescriptions: Baker (1949), Baker & Pritchard (1952c, 1960), Meyer (1979), Meyer & Ryke (1959), Mitrofanov & Strunkova (1979), Pritchard & Baker (1958), Wainstein (1969). Type Depository: Unknown. 419. Cenopalpus naupakticus Hatzinikolis, Panou & Papadoulis 1999: 283; Ampelonas, Naupaktia, Aetoloakarnania, Sterea Hellas, Greece, ex Rubus sp. (Rosaceae). Type Depository: Acari Collection of the Acarology Laboratory of Agricultural University of Athens, Greece. 420. Cenopalpus officinalis Papaioannou-Souliotis, 1986: 15; Aghios Gerasimos, Cephalonia, Greece, ex Rosmarinus officinalis L. (Lamiaceae). Redescription: Hatzinikolis et al. (1999). Type Depository: Acarology Laboratory at the Benaki Phytopathological Institute, Greece. 421. Cenopalpus oleunus (Meyer, 1979): 94; Upington, Cape Province, South Africa, ex Olea africana Mill. (Oleaceae). Species designation: Brevipalpus oleunus—original description, Brevipalpus (Cenopalpus) oleunus—Hatzinikolis (1986b). Redescription: Hatzinikolis (1986b). Type Depository: SANC.

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422. Cenopalpus orakiensis Akbar & Chaudhri, 1985: 283; Orak Valley, Quetta, Punjab, Pakistan, ex Pyrus malus L. (Rosaceae). Redescription: Chaudhri & Akbar (1985). Type Depository: Acarology Research Laboratory, Department of Agric. Entomology, University of Agriculture, Faisalabad, Pakistan. 423. Cenopalpus pegazzanoae Castagnoli, 1987: 111; Altovia, Teramo, Italy, ex Abies alba Mill. (Pinaceae). Species designation: Cenopalpus (Cenopalpus) pegazzanoae—original designation. Type Depository: ISZA. 424. Cenopalpus pennatisetis (Wainstein, 1958): 458, Chimkent, S. Kazakhistan, ex Populus sp. (Salicaceae). Species designations: Brevipalpus pennatisetis—original designation, Cenopalpus pennatisetis—Wainstein (1960), Cenopalpus (Cenopalpus) pennatisetis—Mitrofanov & Strunkova (1979). Redescriptions: Hazinikolis & Emmanouel (1987), Hazinikolis (1987), Khosrowshahi & Arbabi (1997), Livschitz & Mitrofanov (1967), Meyer (1979), Mitrofanov & Strunkova (1979), Wainstein (1960). Type Depository: Unknown. 425. Cenopalpus picitilis Chaudhri, 1971a: 75; Kohala, Murree Hills, Punjab, Pakistan, ex Cotoneaster sp. (Rosaceae). Species designation: Brevipalpus picitilis—Meyer (1979), Cenopalpus picilitis [sic] Hatzinikolis et al. (1999). Redescriptions: Akbar & Chaudri (1985), Chaudhri et al. (1974), Hatzinikolis et al. (1999), Meyer (1979). Type Depository: Department of Entomology, West Pakistan Agricultural University Lyallpur, Pakistan. 426. Cenopalpus piger Wainstein, 1960: 240; Kazakhistan, ex an unidentified plant. Species designation: Brevipalpus piger—Meyer (1979). Redescriptions: Bagdasarian (1962), Meyer (1979). Type Depository: Unknown. 427. Cenopalpus pistaciae Hatzinikolis, Papadoulis & Panou, 1999: 133; Sounio, Attiki, Greece, ex Pistacia lentiscus L. (Anacardiaceae). Type Depository: Acari Collection of the Laboratory of Agricultural Zoology and Entomology, Agricultural University of Athens, Greece. 428. Cenopalpus platani (Livschitz & Mitrofanov, 1967): 18; Georgia, ex Platanus sp. (Platanaceae). Species designations: Brevipalpus platani—original designation, Cenopalpus platani—Hatzinikolis & Emmanouel (1987), Cenopalpus (Cenopalpus) platani—Mitrofanov & Strunkova (1979). Redescriptions: Hatzinikolis (1987), Hatzinikolis & Emmanouel (1987), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository : Unknown. 429. Cenopalpus populi (Livschitz & Mitrofanov, 1967): 16; Georgia, ex Populus sp. (Salicaceae). Species designations: Brevipalpus populi—original designation, Cenopalpus populi—Hatzinikolis & Emmanouel (1987), Cenopalpus (Cenopalpus) populi—Mitrofanov & Strunkova (1979). Redescriptions: Al-Gboory (1987), Hatzinikolis (1987), Hatzinikolis & Emmanouel (1987), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: Unknown. 430. Cenopalpus pritchardi Düzgünes, 1967: 90; Bornova, Izmir, Turkey, ex Pyrus communis L. (Rosaceae). Species designation: Brevipalpus pritchardi—Meyer (1979). Redescriptions: Hatzinikolis (1987), Hatzinikolis & Emmanouel (1987), Khosrowshahi & Arbabi (1997), Meyer (1979). Type Depository: Department of Plant Protection, Faculty of Agriculture, University Ankara, Turkey. Note: host originally listed as Pirus communis. 431. Cenopalpus pseudospinosus (Livschitz & Mitrofanov, 1967): 22; Ukranie, ex Rubus sp. (Rosaceae). Species designation: Brevipalpus pseudospinosus—original designation, Cenopalpus pseudospinous—Hatzinikolis & Emmanouel (1987),. Redescriptions: Hatzinikolis (1987), Hatzinikolis & Emmanouel (1987), Hatzinikolis et al. (1999), Meyer (1979). Type Depository: Unknown.

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432. Cenopalpus pterinus Pritchard & Baker, 1958: 195; Palma Mallorca, Spain, ex Adenostoma sp. (Rosaceae). Species designations: Brevipalpus pterinus—Meyer (1979), Pritchardipalpus pterinus—Mitrofanov (1973a). Redescriptions: Hatzinikolis (1987), Hatzinikolis & Emmanouel (1987), Hatzinikolis et al. (1999), Meyer (1979), Mitrofanov (1973a), Mitrofanov & Strunkova (1979). Type Depository: Unknown. 433. Cenopalpus pulcher (Canestrini & Fanzago, 1876): 134; Trevigiano, Italy, ex hedge plant. Species designations: Caligonus pulcher—original designation, Tenuipalpus pulcher—Berlese (1887), Brevipalpus pulcher—Baker (1949), Cenopalpus pulcher—Pritchard & Baker (1958), Cenopalpus (Cenopalpus) pulcher—Mitrofanov & Strunkova (1979), Brevipalpus pulchur [sic]—Gupta & Chatterjee (1997). Redescriptions: Al-Gboory (1987), Berlese (1887), Bagdasarian (1962), Baker (1949), Baker & Pritchard (1960), Duzgunes (1965), Gupta & Chatterjee (1997), Gutierrez et al. (1989), Hatzinikolis (1987), Hatzinikolis & Emmanouel (1987), Khosrowshahi & Arbabi (1997), Livschitz & Mitrofanov (1967), Meyer (1979), Mitrofanov & Strunkova (1979), Pritchard & Baker (1958), Sadana (1997), Wafa et al. (1968/1969), Wainstein (1960), Zaher & Yousef (1969), Zaher et al. (1984). Comment: Bajwa et al. (2001) reported this species from Oregan (USA). Type Depository: Unknown. Synonym: 433a. Brevipalpus ciferrii Lombardini, 1951: 249; Rocca de Giorgi, Pavia, Lombardia, Italy, ex apple (Rosaceae)—Pritchard & Baker (1958). Species designation: Brevipalpus ciferii [sic]—Pritchard & Baker (1958). Type Depository: Unknown. 433b. Brevipalpus pyri Sayed, 1946: 102, Egypt, ex apples, pears, plums and apricots (Rosaceae)—Pritchard & Baker (1958). Redescription: Baker (1949). Type Depository: Unknown. 433c. Tenuipalpus bodenheimeri Berlese in: Bodenheimer, 1930: 240; Palestine, ex apple (Rosaceae)—Pritchard & Baker (1958). Note: a validation of a mnuscript name of Berlese, based on specimens from Palestine, on apple.—Pritchard & Baker (1958), Wainstein (1960). Type Depository: Unknown. 433d. Tenuipalpus oudemansi Geijskes, 1939: 25; Wageningen, Netherlands, ex apple (Rosaceae) and beech (Fagaceae)—Pritchard & Baker (1958). Redescriptions: Baker (1949), Reck (1951). Type Depository: Unknown. 434. Cenopalpus quadricornis (Livschitz & Mitrofanov, 1967): 22, Armenia, ex Rubus sp. (Rosaceae). Species designation: Brevipalpus quadricornis—original designation, Cenopalpus quadricornis—Hatzinikolis et al. (1999), Cenopalpus (Cenopalpus) spinosus quadricornis—Mitrofanov & Strunkova (1979). Redescriptions: Hatzinikolis et al. (1999), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: Unknown. 435. Cenopalpus ramus Manson, 1963: 222; Murree Hills, Punjab, Pakistan, ex fir (Pinaceae). Species designations: Cenopalpus (Cenopalpus) ramus—Mitrofanov & Strunkova (1979), Brevipalpus ramus—Meyer (1979). Redescriptions: Chaudhri & Akbar (1985), Chaudhri et al. (1974), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: USNM, No. 2890. 436. Cenopalpus ruber Wainstein, 1960: 236; Tajikistan, ex an unidentified plant. Species designations: Brevipalpus ruber—Livschitz & Mitrofanov (1967), Cenopalpus ruber Hatzinikolis & Emmanouel (1987), Cenopalpus (Cenopalpus) ruber—Mitrofanov & Strunkova (1979). Redescriptions: Hatzinikolis (1987), Hatzinikolis & Emmanouel (1987), Livschitz & Mitrofanov (1967), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: Unknown.

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437. Cenopalpus saryabiensis Akbar & Chaudhri, 1985: 270; Saryab, Quetta Valley, Punjab, Pakistan, ex Pinus sp. (Pinaceae). Redescription: Chaudhri & Akbar (1985). Type Depository: Acarology Research Laboratory, Department of Agric. Entomology, University of Agriculture, Faisalabad, Pakistan. 438. Cenopalpus scoopsetus Hatzinikolis & Papadoulis, 1999: 289; Argastiri, Chania, Crete, Greece, ex Erica manipuliflora Salisb. (Ericaceae). Type Depository: Acari Collection Laboratory of Agricultural, Zoology and Entomology, Agricultural University of Athens, Greece. 439. Cenopalpus spinosus (Donnadieu, 1875): 114; France, ex Primula vulgaris (Huds.) (Primulaceae). Note: Host type information according to Pritchard & Baker (1958). Species designations: Tenuipalpus spinosus—original designation, Brevipalpus spinosus—Baker (1949), Cenopalpus spinosus—Pritchard & Baker (1958), Cenopalpus (Cenopalpus) spinosus—Mitrofanov & Strunkova (1979). Redescriptions: Bagdasarian (1962), Baker (1949), Baker & Pritchard (1953), Hatzinikolis (1987), Hatzinikolis & Emmanouel (1987), Hatzinikolis et al. (1999); Livschitz & Mitrofanov (1967), Meyer (1979), Mitrofanov & Strunkova (1979), Pritchard & Baker (1958), Wainstein (1960). Type Depository: Faculté des Sciences de Lyon, France. Note: according to Pritchard & Baker (1958), Donnadieu’s illustrations indicate that the nymphs of this species were referred to as Tenuipalpus spinosus, whereas the adult female was described as Tenuipalpus glaber Donnadieu (1875). Synonym: 439a. Tenuipalpus geisenheyneri Rübsaamen, 1910: 127; Germany, ex Cornus sanguinea L. (Cornaceae)—Pritchard & Baker (1958). Species designation: Brevipalpus geisenheyeri [¨sic]—Baker (1949), Tenuipalpus geisenheineri [sic] Wainstein (1960. Redescriptions: Andre (1954), Attiah (1956), Baker (1949), Baker & Pritchard (1952c), Pritchard & Baker (1958). Type Depository: Unknown. 439b. Tenuipalpus glaber Donnadieu, 1875: 114; France, ex ?—Pritchard & Baker (1958)—Wainstein (1960). Type Depository: Unknown or not specified. Synonym: —Flexipalpus donnadieui Oudemans, 1938: LXXIII (73); nov. nom. – Tenuipalpus glaber Donnadieu 1875 said to be preoccupied by Trombidium glabrum Dugès 1834. Species designation: Brevipalpus donnadieui, n. comb. – Baker (1949). Redescription: Baker (1949). Type Depository: Unknown or not specified. 440. Cenopalpus sunniensis Hasan, Ashfaq, Li, Wakil & Bashir, 2004: 48; Sunny Bank, Murree Hills, Punjab, Pakistan, ex an unidentified plant. Type Depository: Acarology Research Laboratory, Department of Agric. Entomology, University of Agriculture, Faisalabad, Pakistan. 441. Cenopalpus tamarixi (Nassar & Kandeel, 1983): 61; Giza and Cairo, Egypt, ex Tamarix articulata Vahl (Tamaricaceae). Species designation: B. tamarixi Nassar & Kandeel, 1983—original designation, Cenopalpus tamarixi—Zaher et al. (1984). Type Depository: Unknown. Note: original description of species not was could not be located. The information provided here was taken from Zaher et al, 1984: 61. Third Egytian Hungary Confr. Plant Protection (under publication). 442. Cenopalpus taygeticus Hatzinikolis, Panou & Papadoulis, 1999: 279; Taygetus Mountain, Messinia, Peleponnessus, Greece, ex Rubus sp. (Rosaceae). Type Depository: Acari Collection of the Acarology Laboratory of Agricultural University of Athens, Greece. 443. Cenopalpus viniferus Hatzinikolis, Papadoulis & Kapaxidi, 2001: 35; Vathi, Avlida; Boeotia, Sterea Hellas, Greece, ex Vitis vinifera L. (Vitaceae). Type Depository: Acari Collection of the Acarology Laboratory of Agricultural University of Athens, Greece.

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444. Cenopalpus virgulatus Akbar & Chaudhri, 1985: 274; Garm Chashma, Chitral Valley, Pakistan, ex an unidentified plant. Redescription: Chaudhri & Akbar (1985). Type Depository: Acarology Research Laboratory, Department of Agric. Entomology, University of Agriculture, Faisalabad, Pakistan. 445. Cenopalpus wainsteini (Livschitz & Mitrofanov, 1967): 16, Ukraine, ex Pinus sp. (Pinaceae). Species designations: Brevipalpus wainsteini—original designation, Cenopalpus wainsteini—Hatzinikolis & Emmanouel (1987), Cenopalpus (Cenopalpoides) wainsteini—Mitrofanov & Strunkova (1979). Redescriptions: Hatzinikolis (1987), Hatzinikolis & Emmanouel (1987), Meyer (1979), Mitrofanov & Strunkova (1979), Pegazzano (1970). Type Depository: Unknown. Synonym: 445a. Cenopalpus fewstrii Zaher & Yousef, 1969: 278; Giza, Egypt, and Gaza Strip, ex Pinus sp. (Pinaceae)—Hatzinikolis (1983). Redescriptions: Hatzinikolis & Emmanouel (1987), Wafa et al. (1968/1969), Zaher et al. (1984). Type Depository: CUE. 446. Cenopalplus xini Ma & Li, 1984: 325; Shihezi, Xinjiang, China, ex Populus nigra L. var. thevestina (Dode) Bean (Salicaceae). Type Depository: Shangai College of Agriculture, Xinjiang, China.

Chaudhripalpus Mesa, Welbourn and Evans, NEW GENUS Type species: Crossipalpus creelae Smiley, Frost & Gerson, 1996: 176. Diagnosis: Palpus 5-segmented with 2 simple setae, terminal podomere short, with one simple seta, an eupathidium and an elongate solenidion. Idiosoma elongate oval. Prodorsum hoodlike, completely covering the gnathosoma with anterior margin rounded and bearing 3 pairs of setae (V2, Sc1, Sc2). Dorsum of opisthosoma with 9 pairs of broadly spatulate setae: c1, c3, d1, d3, e1, e3, f3, h1, h2. Propodosoma and metapodosoma each with a pair of long intercoxal setae (1a, 3a), 4a absent Aggenital, genital and pseudanal setae smooth. Leg setal counts—coxae: 1-2-1-1, trochanters: 1-1-2-1, femora: 4-3-4-2, genua: 1-1-0-0, tibiae: 4-4-3-4, tarsi 6(+1)-6(+1)-4-4. Comparison: The genus Chaudhripalpus is similar to the genus Crossipalpus but species of the latter have 12 pairs of lanceolate dorsal opisthosomal setae. Chaudhripalpus is also similar to the genus Philippipalpus, but the latter has the anterior margin of prodorsum deeply incised and 10 pairs of dorsal opisthosomal setae (f2 present). Etymology: The generic name Chaudhripalpus is given in honor of Dr. W. M. Chaudhri for his many contributions to the taxonomy of mites. 447. Chaudhripalpus creelae (Smiley, Frost & Gerson, 1996): 176;—new combination; 16 km E Mount Mary, South Australia, Australia, ex Allocasuarina luehmannii (R.T. Baker) L.A.S. Johnson (Casuarinaceae). Species designations: Crossipalpus creelae—original designation, Smiley, Frost & Gerson, 1996: 176. Type Depository: WINC.

Coleacarus Meyer 1979 Coleacarus Meyer, 1979: 111. Type species: Coleacarus lithops Meyer 1979: 111. 448. Coleacarus lithops Meyer, 1979: 111; Witputz, Namibia (formerly South West Africa), ex Lithops erniana Tisch. & Jacobsen (Aizoaceae). Type Depository: SANC.

Crossipalpus Smiley, Frost & Gerson 1996 Crossipalpus Smiley, Frost & Gerson, 1996: 174. Type species: Crossipalpus verticillatae Smiley, Frost & Gerson, 1996: 174.

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449. Crossipalpus muellerianae Smiley, Frost & Gerson, 1996: 174; Saint Mary's Peak, Flinders Ranges, South Australia, Australia, ex Allocasuarina muelleriana (Miq.) L.A. S. Johnson subsp. alticola L.A.S. Johnson (Casuarinaceae). Type Depository: WINC. 450. Crossipalpus verticillatae Smiley, Frost & Gerson, 1996: 174; 10 km WNW of Nairne, South Australia, Australia, ex Allocasuarina verticillata (Lam.) L.A.S. Johnson (Casuarinaceae). Type Depository: WINC.

Dolichotetranychus Sayed 1938 Dolichotetranychus Sayed, 1938: 606. Type species: Stigmaeus floridanus Banks 1900: 77; Baker & Pritchard, 1956: 357; Baker & Pritchard, 1960, 570; Baker & Tuttle, 1964: 73; Baker & Tuttle, 1972: 32; Baker & Tuttle, 1987: 110; Ehara, 1966: 5; Evans et al., 1993: 129; Hatzinikolis, 1987: 57; Lo, 1969: 99; Meyer, 1979: 99; Meyer & Gerson, 1980: 76; Mitrofanov, 1973b: 1317; Pritchard & Baker, 1952: 44; Pritchard & Baker, 1958: 250; Rimando, 1962: 44; Sayed, 1950: 1016; Smiley & Gerson, 1995: 36; Wainstein, 1960: 235. Dolichotetranychus (Dolichotetranychus) Mitrofanov, 1973b: 1317; synonymy according to Meyer 1979: 99. Dolichotetranychus (Dolichotetranychoides) Mitrofanov, 1973b: 1317. Type species: Dolichotetranychus summersi Pritchard & Baker, 1952; synonymy according to Meyer 1979: 99; Baker & Tuttle, 1987: Stenotetranychus Mitrofanov, 1973b: 1317. Type species: Siteroptes carnea Banks, 1906; synonymy according to Meyer 1979: 99. 451. Dolichotetranychus alpinus Collyer, 1973b: 747; Mount Barber, 1.1 km Wilmot Pass, west of Lake Manapouri, Southland, New Zealand, ex Chionochloa sp. (Poaceae) with Celmisia laricifolia Hook. (Asteraceae). Species designation: Stenotetranychus alpinus—Mitrofanov & Strunkova (1979). Redescriptions: Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: Collection Entomological Division, Department of Scientific and Industrial Research, Nelson, New Zealand. 452. Dolichotetranychus ancistrus Baker & Pritchard, 1956: 365, Rangitoto Island, New Zealand, ex Hebe salicifolia Pennell (Scrophulariaceae). Species designation: Dolichotetranychus (Dolichotetranychus) ancistrus—Mitrofanov & Strunkova (1979). Redescriptions: Meyer (1979), Mitrofanov & Strunkova (1979), Pritchard & Baker (1958), Zhang & Fan (2004). Type Depository: USNM, No. 2211. 453. Dolichotetranychus apaches Baker & Pritchard, 1956: 379; Coral Gables, Florida, USA, ex sheath of “air plant” (Tillandsia sp. (Bromeliaceae)). Species designation: Dolichotetranychus (Dolichotetranychus) apaches—Mitrofanov & Strunkova (1979). Redescriptions: Meyer (1979), Pritchard & Baker (1958), Mitrofanov & Strunkova (1979). Type Depository: USNM, No. 2214. 454. Dolichotetranychus australianus (Womersley, 1943): 245; Gayndah, South Queensland, Australia, ex Cynodon dactylon (L.) Pers. (Poaceae). Species designations: Trichadenus australianus—original designation, Dolichotetranychus australianus—Baker & Pritchard (1956), Dolichotetranychus (Dolichotetranychus) australianus—Mitrofanov & Strunkova (1979). Redescriptions: Baker & Pritchard (1956, 1960), El-Enany & Solinan (1987), Meyer (1979), Mitrofanov & Strunkova (1979), Pritchard & Baker (1958), Smiley & Gerson (1995), Wafa et al. (1968/1969), Zaher et al. (1984). Type Depository: SAM. 455. Dolichotetranychus babylonicus Al-Gboory, 1987: 76; Abu-Ghraib, Baghdad, Iraq, ex Phragmites australis (Cav.) Trin. & Steud. (Poaceae). Type Depository: Entomology Museum, College of Agriculture, Department of Plant Protection, Abu-Ghraib, Baghdad, Iraq. 456. Dolichotetranychus bakeri El-Enany & Soliman, 1987: 2; between Giza and Fayoum, Egypt, ex an unidentified desert plant. Redescription: Zaher et al. (1984). Type Depository: CUE.

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457. Dolichotetranychus carnea (Banks, 1906): 140; New Mexico and Utah, USA, ex Sporobolus grass (Poaceae). Species designations: Siteroptes carnea—original designation, Dolichotetranychus carnea—Pritchard & Baker (1952), Stenotetranychus carnea—Mitrofanov & Strunkova (1979). Redescriptions: Baker & Pritchard (1956), Meyer (1979), Mitrofanov & Strunkova (1979), Pritchard & Baker (1952, 1958). Type Depository: USNM. Note: host originally listed as Spirobolus grass. Synonym: 457a. Stigmaeodes cinctus Ewing, 1911: 39; Fort Collins, Colorado, USA, ex wild grass—Pritchard & Baker (1952). Type Depository: Unknown. 458. Dolichotetranychus cocos Flechtmann & Fernando, 2000: 145; Bandirippuwa, Lunuwila, Sri Lanka, ex Cocos nucifera L. (Arecaceae). Type Depository: DEES. 459. Dolichotetranychus cracens Pritchard & Baker, 1958: 253; Woodward, Oklahoma, USA, ex Sporobolus cryptandrus (Torr.) Gray (Poaceae). Species designation: Dolichotetranychus (Dolichotetranychus) cracens—Mitrofanov & Strunkova (1979). Redescriptions: Meyer (1979), Mitrofanof & Strunkova (1979). Type Depository: USNM, No. 2212. 460. Dolichotetranychus cymosum Meyer, 1979: 100; Fuller's Bay, East London, Cape Province, South Africa, ex Helichrysum cymosum D. Don (Asteraceae). Type Depository: SANC. 461. Dolichotetranychus elatariae Mohanasundaram, 1983a: 152; Dharwad, Karnataka, India, ex Elettaria cardamomum (L.) Maton (Zingiberaceae). Redescription: Sadana (1997). Type Depository: TNAU. Note: host originally listed as Elataria cardamomi. 462. Dolichotetranychus floridanus (Banks, 1900): 77; Florida, USA, ex pineapple (Bromeliaceae). Species designations: Stigmaeus floridanus—original designation, Pseudoleptus floridanus—Oudemans (1927), Trichadenus floridanus—McGregor (1949), Dolichotetranychus floridanus—Sayed (1938), Dolichotetranychus (Dolichotetranychus) floridanus—Mitrofanov & Strunkova (1979). Redescriptions: Baker & Pritchard (1956), Baker & Tuttle (1987), Chandra & Channabasavanna (1974), El-Enany & Soliman (1987), Ehara (1966), Evans et al. (1993), Hatzinikolis (1987), Lo (1969), McGregor (1949), Meyer (1979), Oudemans (1927), Pritchard & Baker (1952, 1958), Rimando (1962), Sadana (1997), Salas (1983), Sayed (1938), Smiley & Gerson (1995), Wafa et al. (1968/1969), Zaher et al. (1984). Type Depository: Unknown. 463. Dolichotetranychus gramineus Mitrofanov & Strunkova, 1978: 1097; Tajikistan, ex an unidentified cereal. Species designation: Dolichotetranychus (Dolichotetranychus) gramineus—Mitrofanov & Strunkova (1979). Redescription: Mitrofanov & Strunkova (1979). Type Depository: State Nikitsky Botanical Gardens (Yalta) and Institute of Zoology and Parasitology, Academy of Sciences of the Tadjik SSR (Dushanbe) No. 2553/343 464. Dolichotetranychus macer Baker & Pritchard, 1956: 372; Cape Province, South Africa, intercepted at Washington, D.C., USA, ex Aristida namaquensis Trin. & Rupr. (Poaceae). Species designation: Stenotetranychus macer—Mitrofanov & Strunkova (1979). Redescriptions: Baker & Pritchard (1960), Meyer & Ryke (1959), Meyer (1979), Mitrofanov & Strunkova (1979), Pritchard & Baker (1958). Type Depository: USNM, No. 2210. 465. Dolichotetranychus micidus Baker & Pritchard, 1956: 363; Denver, Colorado, USA, ex Muhlenbergia asperifolia (Nees, Meyen & Trin.) Parodi (Poaceae). Species designation: Dolichotetranychus (Dolichotetranychus) micidus—Mitrofanov & Strunkova (1979). Redescriptions: Meyer (1979), Mitrofanov & Strunkova (1979), Pritchard & Baker (1958). Type Depository: USNM, No. 2209.

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Note: host originally listed as “(Sporobolus asperifolius) = Muhlenbergia asperifolia”. 466. Dolichotetranychus muhlenbergia Baker & Tuttle, 1972: 33; McNary, Arizona, USA, ex Muhlenbergia torreyi (Kunth) A.S. Hitchcock & Bush (Poaceae). Species designation: Dolichotetranychus (Dolichotetranychus) muhlenbergia—Mitrofanov & Strunkova (1979). Redescriptions: Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: USNM, No. 3436. 467. Dolichotetranychus repenae Mohanasundaram, 1983a: 151; Pennadam, Tamil Nadu, India, ex Panicum repens L. (Poaceae). Redescription: Sadana (1997). Type Depository: TNAU. 468. Dolichotetranychus salinas Pritchard & Baker, 1952: 46; Modesto, California, USA, ex Distichlis spicata (L.) Greene (Poaceae). Species designation: Dolichotetranychus (Dolichotetranychus) salinas—Mitrofanov & Strunkova (1979). Redescriptions: Baker & Pritchard (1956), Baker & Tuttle (1964), Meyer (1979), Meyer & Ryke (1959), Mitrofanov & Strunkova (1979), Pritchard & Baker (1958). Type Depository: USNM, No. 1933. 469. Dolichotetranychus summersi Pritchard & Baker, 1952: 45; Bond's Corner, Calexico, Imperial Valley, California, USA, ex Bermuda grass (Poaceae). Species designation: Dolichotetranychus (Dolichotetranychoides) summersi—Mitrofanov & Strunkova (1979). Redescriptions: Baker & Pritchard (1956), Baker & Tuttle (1964), Meyer (1979), Mitrofanov & Strunkova (1979), Pritchard & Baker (1958). Type Depository: USNM, No. 1934. Note: according to Pritchard & Baker (1958) the male holotype was found ex Cynodon dactylon (L.) Pers. (Poaceae). 470. Dolichotetranychus tenellae Mohanasundaram, 1983a: 149; Vridhachalam, Tamil Nadu, India, ex Eragrostis tenella (L.) Beauv., Roemer & J.A. Schultes (Poaceae). Redescriptions: Sadana (1997). Type Depository: TNAU. Note: host originally listed as Eragrastis tenella. 471. Dolichotetranychus vandergooti (Oudemans, 1927): 179; Buitenzorg, Java, Indonesia, ex Dendrobium sp. (Orchidaceae). Species designations: Pseudoleptus vandergooti—original designation, Dolichotetranychus vandergooti—Pritchard & Baker (1952), Stenotetranychus vandergooti—Mitrofanov & Strunkova (1979). Redescriptions: Baker & Pritchard (1956), Charanasri et al. (1986), Ehara (1966), Meyer (1979), Mitrofanov & Strunkova (1979), Pritchard & Baker (1952, 1958), Rimando (1962). Type Depository: Unknown. 472. Dolichotetranychus wiesei Meyer, 1979: 101; Edenville, Orange Free State, South Africa, ex grass (Poaceae) and soil. Type Depository: SANC. 473. Dolichotetranychus zoysiae Ehara, 2004: 130; Onnason, Okinawa Island, Japan, ex Zoysia tenuifolia. Willd. & Thiele (Poaceae). Type Depository: NSMT.

Krugeria Meyer 1979 Krugeria Meyer, 1979: 107. Type species: Krugeria ramosa Meyer, 1979: 107. 474. Krugeria ramosa Meyer, 1979: 107; N. 25 of the Grid-map of Kruger National Park, Transvaal, South Africa, ex Grewia bicolor Juss. (Tiliaceae). Type Depository: SANC.

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Larvacarus Baker & Pritchard 1952 Larvacarus Baker & Pritchard, June 1952: 130. Type species: Phytoptipalpus transitans Ewing, 1922: 108; Mitrofanov, 1973: 1315. Phytoptipalpoides Reck, July 1952: 422. 475. Larvacarus transitans (Ewing, 1922): 108; Pusa, New Delhi, India, ex in galls on Ziziphus mauritiana Lam. (Rhamnaceae) Species designations: Phytoptipalpus transitans—original designation, Larvacarus transitans—Baker & Pritchard (1952c), Phytoptipalpoides transitans—Reck (1952). Redescriptions: Al-Gboory (1987), Baker & Pritchard (1952c), Gupta & Chatterjee (1997), Meyer (1979), Mitrofanov & Strunkova (1979), Pritchard & Baker (1958), Reck (1953), Sadana (1997). Type Depository: USNM, No. 24722. Note: host originally listed as Zizyphus jujuba.

Macfarlaniella Baker & Pritchard 1962 Macfarlaniella Baker & Pritchard, 1962: 123. Type species: Raoiella queenslandica Womersley, 1942: 88; Mitrofanov, 1973b: 1316; Mitrofanov & Strunkova, 1979: 122; Meyer, 1979: 114; Smiley & Gerson, 1995: 39. 477. Macfarlaniella queenslandica (Womersley, 1942): 88; Redland Bay, Brisbane, Queensland, Australia, ex Eucalyptus micrantha DC. [=Eucalyptus racemosa] (Myrtaceae). Species designation: Raoiella queenslandica—original designation. Type Depository: SAM. Redescriptions: Baker & Pritchard (1962), Smiley & Gerson (1995).

Magdalenapalpus Mesa, Welbourn and Evans, NEW GENUS Type species: Meyeraepalpus strandtmanni Smiley, Frost & Gerson, 1996: 178, by monotypy. Diagnosis: dorsum—oval shaped; prodorsum hoodlike and completely covering the gnathosoma with anterior margin deeply incised medially and bearing 3 pairs of broadly lanceolate, serrate setae (v2, sc1, sc2); rostral shield absent; opisthosoma with 13 pairs of broadly lanceolate, serrate setae, arranged as follows: c1, c2, c3, d1, d2, d3, e1, e2, e3, f2, f3, h1, h2 venter—palpus 5-segmented, palp tibia with two simple setae, palp tarsus short, with one simple setae, an eupathidium and an elongated solenedion; podosoma and metapodosoma each with a pair of smooth, long intercoxal setae (1a, 3a); 4a absent; pregenital, genital and pseudanal setae smooth. Comparison: The species Magdalenapalpus strandtmanni differs from Australopalpus alphitoniae by having the bases of d1-d1 setae not close together, with the distance between them similar to that of c1-c1 setal bases; and the genital and ventral plate setae smooth; whereas the anterior margin of prodorsum is rounded, slightly pointed at apex; the d1 setae are placed close together, with the distance between them less than half the distance of the c1-c1 setal bases; genital and ventral plate setae barbed in Australopalpus. Discussion: Smiley et al. (1996) described this species and placed it in their new genus Meyeraepalpus, of which they chose Meyeraepalpus delfinadae Smiley, Frost & Gerson as the type speciesfor the genus. We consider Meyeraepalpus delfinadae to be a species of Aegyptobia Sayed 1950. Therefore, the genus name Meyeraepalpus is no longer a valid name since the type species of the genus Meyeraepalpus belongs to a previously described genus. The second species they described in the genus, Meyeraepalpus strandtmanni Smiley, Frost & Gerson, 1996, differs from any previously described genus; however, since the name Meyeraephagus is no longer available due to the synonymy with Aegyptobia, we hereby erect the genus Magdalenapalpus as a replacement name for Meyeraepalpus in honor of Dr. Magdalena Meyer for her excellent work, and many contributions to the taxonomy of the family Tenuipalpidae and other mite families. 478. Magdalenapalpus strandtmanni (Smiley, Frost & Gerson), 1996: 178—new combinatiuon; 16 km east of Mount Mary, South Australia, Australia, ex Allocasuarina luehmannii (R.T. Baker) L.A.S. Johnson (Casuarinaceae). Species designation: Meyeraepalpus strandtmanni Smiley, Frost & Gerson, 1996: 178. Type Depository: WINC.

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Obdulia Pritchard & Baker 1958 Obdulia Pritchard & Baker, 1958: 260. Type species: Obdulia tamaricis Pritchard & Baker, 1958: 260; Hatzinikolis, 1987: 57; Meyer, 1979: 114; Meyer & Gerson, 1980: 76, Mitrofanov, 1973b: 1316; Mitrofanov & Strunkova, 1979: 123; Wainstein, 1960: 235. 479. Obdulia daadi Al-Gboory, 1987: 94; near Bagdad, Iraq, collected from the leaf base of Tamarix pentandra Pall. (Tamaricaceae). Type Depository: Entomology Museum, College of Agriculture, Department of Plant Protection, Abu-Ghraib, Baghdad, Iraq. Note: This species is tentatively retained in the genus Obdulia; however, it differs from the type species, O. tamaricis, by its elongate oval-shaped body with broadly palmate dorsal setae; the c1, d1 and e1 setae are located in the dorsocentral region; and for being collected at the base of the leaf. In O. tamaricis, the body is round, the dorsal setae are long and setiform; the c1, d1 and e1 are located in the sublateral region; and they are found in galls on the twigs of O. tamaricis. 480. Obdulia tamaricis Pritchard & Baker, 1958: 260; Wadi Fukra, Israel, ex twigs galls on Tamarix maris-mortui Gutm. (Tamaricaceae). Redescriptions: Hatzinikolis (1987), Mitrofanov & Strunkova (1979). Type Depository: USNM, No. 2342.

Obuloides Baker & Tuttle 1975 Obuloides Baker & Tuttle, 1975: 453. Type species. Obuloides rajamohani Baker & Tuttle, 1975: 453; Mitrofanov & Strunkova, 1979: 123; Meyer, 1979: 107. 481. Obuloides rajamohani Baker & Tuttle, 1975: 453; Tamil Nadu Agricultural University, Coimbatore, Tamil Nadu, India, ex from galls of Eriophyes hibisci Nalepa on Hibiscus sp. (Malvaceae). Redescriptions: Mitrofanov & Strunkova (1979), Sadana (1997). Type Depository: USNM, No. 3689. 482. Obuloides rimandoi Corpuz-Raros, 1978: 218; Mount Makiling, Puting Lupa, Calamba, Laguna, Philippines, ex Grewia multiflora Juss. (Tiliaceae). Type Depository: USNM.

Pentamerismus McGregor 1949 Pentamerismus McGregor, 1949: 23. Type species: Tenuipalpus erythreus Ewing, 1917: 152; Baker & Pritchard, 1953: 353; Baker & Yin, 1988: 149; Baker & Tuttle, 1972: 26; Baker & Tuttle, 1987: 111; Chaudhri et al. 1974: 29; Duzgunes, 1965: 124; Evans et al. 1993: 129; Hatzinikolis, 1987: 55; Livschitz & Mitrofanov, 1967: 11; Ma & Yuan, 1980: 116; Meyer, 1979: 128; Meyer & Van Dis, 1993: 321; Mitrofanov, 1973a: 508; Mitrofanov et al. 1975: 13; Mitrofanov & Strunkova, 1979: 35; Pritchard & Baker 1952: 8; Pritchard & Baker, 1958: 186; Wainstein, 1960: 234. Brevipalpoides Reck, 1951: 290. Type species: Brevipalpoides juniperi Reck, 1951; synonymy according to Pritchard & Baker 1958: 186; Sepasgosarian (1983). Oligomerismus Mitrofanov, 1973a: 508. Type species: Tenuipalpus taxi Haller, 1877; synonymy according to Meyer 1979: 130; Sepasgosarian (1983), Baker & Tuttle (1987). Livschitzia Mitrofanov, 1973b: 1317. Type species: Pentamerismus tauricus Livschitz & Mitrofanov, 1970; synonymy according to Meyer 1979: 130; Sepasgosarian (1983), Baker & Tuttle (1987). 483. Pentamerismus abnormis Baker & Tuttle, 1987: 112; Chihuahua, Mexico intercepted at El Paso, Texas quarantine station, Texas, USA, ex Juniperus sp. (Cupressaceae). Type Depository: USNM. 484. Pentamerismus arbutusae Baker & Tuttle, 1987: 113; 10 km N San Luis Potosi, Mexico, ex Arbutus sp. (Ericaceae). Type Depository: USNM.

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485. Pentamerismus canadensis McGregor, 1949: 24; British Columbia, Canada, intercepted at Seattle, Washington, DC. USA, ex Thuja sp. (Cupressaceae). Species designation: Oligomerismus canadensis—Mitrofanov et al (1975). Redescriptions: Baker & Pritchard (1953c, 1958), Meyer (1979), Mitrofanov & Strunkova (1979), Mitrofanov et al. (1975). Type Depository: USNM, No. 1530. Note: host originally listed as Thuya sp. 486. Pentamerismus collinus Meyer & Van Dis, 1993: 322; Meiringspoort, Cape Province, South Africa, ex Passerina falcifolia C.H. Wright (Thymelaeaceae). Type Depository: SANC. 487. Pentamerismus coronatus (Canestrini & Fanzago, 1876): 136; Trevignano, Treviso, and Emilia, Modena, Italy, ex Thuja sp. (Cupressaceae). Species designations: Caligonus coronatus—original designation, Tenuipalpus coronatus—Berlese (1886), Brevipalpoides cornatus [sic]—Reck (1951), Pentamerismus coronatus—Baker & Pritchard (1953c), Oligomerismus coronatus—Mitrofanov et al. (1975). Redescriptions: Baker & Pritchard (1953c), Berlese (1886), Hatzinikolis (1987), Meyer (1979), Mitrofanov & Strunkova (1979), Mitrofanov et al. (1975), Pritchard & Baker (1958), Reck (1951). Type Depository: Unknown. 488. Pentamerismus erythreus (Ewing, 1917): 152; Ames, Iowa, USA, ex arbor vitae (Cupressaceae). Species designations: Tenuipalpus erythreus Ewing—original designation, Pentamerismus erythreus—McGregor (1949). Redescriptions: Baker & Pritchard (1953c), Baker & Tuttle (1964, 1987), Evans et al. (1993), McGregor (1949), Meyer (1979), Meyer & Van Dis (1993), Mitrofanov et al. (1975), Mitrofanov & Strunkova (1979), Pritchard & Baker (1952, 1958), Thewke & Enns (1970). Type Depository: Unknown or not specified. 489. Pentamerismus foetidae (Meyer, 1979): 121; Van Rhynsdorp, Cape Province, South Africa, ex Zygophyllum foetidum Schrad. & Wendl. (Zygophyllaceae). Species designation: Aegyptobia foetidae—original designation, Pentamerismus foetidae—Meyer & Van Dis (1993). Redescription: Meyer & Van Dis (1993). Type Depository: SANC. 490. Pentamerismus foliisetis Livschitz & Mitrofanov, 1967: 38; Ukraine, ex (Cupressaceae). Redescriptions: Meyer (1979), Mitrofanov & Strunkova (1979), Mitrofanov et al. (1975). Type Depository: Unknown. 491. Pentamerismus judiciarius De Leon, 1961c: 177; Burnsville, North Carolina, USA, ex Thuja occidentalis L. (Cupressaceae). Species designation: Oligomerismus judiciarius—Mitrofanov et al. (1975). Redescriptions: Meyer (1979), Mitrofanov & Strunkova (1979), Mitrofanov et al. (1975). Type Depository: MCZ. 492. Pentamerismus juniperi (Reck, 1951): 292; Georgia, ex Juniperus sp. (Cupressaceae). Species designation: Brevipalpoides juniperi—original designation, Pentamerismus juniperi—Pritchard & Baker (1958). Redescriptions: Bagdasarian (1962), Hatzinikolis (1987), Livschitz & Mitrofanov (1967), Meyer (1979), Mitrofanov & Strunkova (1979), Mitrofanov et al. (1975), Pritchard & Baker (1958). Type Depository: Unknown. 493. Pentamerismus kunmingensis Ma & Yuan, 1980: 116; Kunming, Yunnan, China, ex Biota orientalis L. (Cupressaceae). Type Depository: Shanghai College of Agriculture, China. 494. Pentamerismus oregonensis McGregor, 1949: 27; Rhododendron, Oregon, USA, ex Libocedrus decurrens Torr. (Cupressaceae). Species designation: Oligomerismus oregonensis—Gutierrez et al. (1989).

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Redescriptions: Bagdasarian (1962), Baker & Pritchard (1953c), Baker & Tuttle (1987), Chandra & Channabasavanna (1974), Chaudhri et al. (1974), Ehara (1962, 1966, 1969), Gutierrez et al. (1989), Hatzinikolis (1987), Livschitz & Mitrofanov (1967), Meyer (1979), Mitrofanov & Strunkova (1979), Mitrofanov et al. (1975), Pritchard & Baker (1952, 1958), Sadana (1997). Type Depository: USNM, No. 1531. 495. Pentamerismus ossianensis Thewke & Enns, 1971: 250; Ossian, Livingston, New York, USA, ex Thuja sp. (Cupressaceae). Species designation: Oligomerismus ossianensis—Mitrofanov & Strunkova (1979). Redescriptions: Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: USNM. 496. Pentamerismus retusus Meyer, 1979: 130; Swartkop, near Upington, Cape Province, South Africa, ex Zygophyllum simplex L. (Zygophyllaceae). Redescription: Meyer & Van Dis (1993). Type Depository: SANC. 497. Pentamerismus shandongensis Wang & Ma, 1993: 3; Shandong, China, ex Celtis sinensis Pers. (Ulmaceae). Type Depository: Unknown or not specified. Note: The original description of this species was not seen. 498. Pentamerismus tauricus Livschitz & Mitrofanov, 1970: 787; Krymskaya, Ukraine, ex Cistus tauricus J. Presl & C. Presl. (Cistaceae). Species designation: Livschitzia tauricus—Mitrofanov (1973b). Redescriptions: Meyer (1979), Mitrofanov (1973). Type Depository: Unknown. 499. Pentamerismus taxi (Haller, 1877): 85; Switzerland, ex Taxus sp. (Taxaceae). Species designations: Tenuipalpus taxi—original designation, Pentamerismus taxi Baker & Pritchard (1953), Oligomerismus taxi—Mitrofanov et al. (1975). Redescriptions: Bagdasarian (1962), Baker & Pritchard (1953), Ehara (1962), Gutierrez et al. (1989), Hatzinikolis (1987), Livschitz & Mitrofanov (1967), Meyer (1979), Mitrofanov (1973a), Mitrofanov & Strunkova (1979), Mitrofanov et al. (1975), Pritchard & Baker (1952, 1958). Type Depository: Unknown. Synonym: 499a. Pentamerismus morishitai Pritchard & Baker, 1952: 12; Berkeley, California, USA, ex Taxus baccata L. (Taxaceae)—Pritchard & Baker (1958). Redescription: Baker & Pritchard (1953c). Type Depository: USNM, No. 1905. Notes: according to Pritchard & Baker (1952), new combination????; previously known as Tenuipalpus taxi Haller (1877). According to Pritchard & Baker (1958), although there is no type and the original figure is inadequate, the occurrence of the same species, restricted to English Yew (T. baccata), in both California and Switzerland, leads to the synonymy of morishitai with taxi. Bowman & Southgate (1980) reported this species on yews in England.

Philippipalpus Corpuz-Raros 1978 Philippipalpus Corpuz-Raros, 1978: 219. Type species: Philippipalpus agohoi Corpuz-Raros, 1978: 220; Smiley, Frost & Gerson, 1996: 172. 500. Philippipalpus agohoi Corpuz-Raros, 1978: 220; Santa Ana, Cagayan, Philippines, ex Casuarina equisetifolia L. (Casuarinaceae). Redescription: Smiley et al. (1996). Type Depository: USNM.

Phyllotetranychus Sayed 1938 Phyllotetranychus Sayed, 1938: 602. Type species: Phyllotetranychus aegyptium Sayed, 1938: 602; Baker & Pritchard, 1960: 571; Meyer, 1979: 115; Meyer & Gerson, 1980: 77; Mitrofanov, 197: 508; Mitrofanov & Strunkova, 1979: 21; Pritchard & Baker, 1958: 254; Sayed, 1950: 1016; Wainstein, 1960: 234.

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501. Phyllotetranychus aegyptium Sayed, 1938: 602; Mataanah Haute, Egypt, ex dates (Arecaceae). Species designation: Phyllotetranychus aegyptiacus [sic]—Sayed (1940), Baker & Pritchard (1960). Redescriptions: Baker & Pritchard (1960), Gerson et al. (1983), Meyer (1979), Mitrofanov & Strunkova (1979), Pritchard & Baker (1958), Sayed (1940). Type Depository: Unknown. 502. Phyllotetranychus romaine Pritchard & Baker, 1958: 255, Mauritius, ex Latania loddigesii Mart. (Arecaceae). Redescriptions: Baker & Pritchard (1960), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: BMNH. Note: host originally listed as Latania loddgesi.

Phytoptipalpus Trägardh 1904 Phytoptipalpus Trägardh, 1904: 9. Type species: Phytoptipalpus paradoxus Trägardh, 1904: 10; Baker & Pritchard, 1960: 561; Baker & Tuttle, 1987: 115; Hatzinikolis, 1987: 53; Meyer, 1979: 111; Meyer & Van Dis, 1993: 324; Mitrofanov, 1973b: 1315; Mitrofanov & Strunkova, 1979: 119; Pritchard & Baker, 1958: 189; Sayed, 1942: 115; Wainstein, 1960: 235; Zaher & Yousef, 1969: 275. Neophytoptipalpus Mitrofanov, 1973b: 1315. Type species: Phytoptipalpus albizziae Pritchard & Baker, 1958; synonymy according to Meyer 1979: 112; Sepasgosarian (1983), Baker & Tuttle (1987). Zaheria Mitrofanov, 1973b: 1316. Type species: Phytoptipalpus aegyptetrapodus Zaher & Yousef, 1969; synonymy according to Meyer 1979: 112; Sepasgosarian (1983), Baker & Tuttle (1987). 503. Phytoptipalpus aegyptetrapodus Zaher & Yousef, 1969: 275; El-Kalubia, El-Sharkia, Giza, Cairo and New Valley, Egypt, ex Acacia nilotica (L.) Willd. & Delile (Mimosaceae). Species designations: Zaheria aegyptetrapodus—Mitrofanov (1973b), Aegyptobia aegyptetrapoda [sic]- Meyer (1979). Redescriptions: Meyer (1979), Meyer & Van Dis (1993), Mitrofanov (1973b), Mitrofanov & Strunkova (1979), Zaher et al. (1984). Type Depository: CUE. 504. Phytoptipalpus albizziae Pritchard & Baker, 1958: 189; Assam, India, ex Albizia procera (Roxb.) Benth. (Mimosaceae). Species designation: Neophytoptipalpus albizziae—Mitrofanov (1973b). Redescriptions: Meyer (1979), Mitrofanov (1973), Mitrofanov & Strunkova (1979), Sadana (1997). Type Depository: USNM, No. 2295. Note: host originally listed as Albizzia procera. 505. Phytoptipalpus alexandriae Meyer & Van Dis, 1993: 328; Naukluft Mountains, Namib-Naukluft Park, Namibia, ex Catophractes alexandri D. Don (Bignoniaceae). Type Depository: SANC. 506. Phytoptipalpus ceibae (De Leon, 1962): 203; about 19.2 km S Guadalajara, Route 15, Jalisco, Mexico, ex Ceiba sp. (Malvaceae). Species designations: Aegyptobia ceibae—original designation, Phytoptipalpus ceibae—Baker & Tuttle (1987), Aegyptobia (Aegyptobia) ceibae—Mitrofanov & Strunkova (1979). Redescriptions: Baker & Tuttle (1987), Baker et al. (1975), Meyer (1979); Mitrofanov & Strunkova (1979). Type Depository: MCZ. 507. Phytoptipalpus cercidium (Baker, Tuttle & Abbatiello, 1975): 2; Mazatlan, Sinaloa, Mexico, ex Cercidium floridum Benth. & Gray (Caesalpinaceae). Species designations: Aegyptobia cercidium—original designation, Phytoptipalpus cercidium—Baker & Tuttle (1987). Redescriptions: Baker & Tuttle (1987), Meyer (1979). Type Depository: USNM, No. 3621. 508. Phytoptipalpus conostegiae Baker & Tuttle, 1987: 117; 9.6 km S Iguala, Guerrero, Mexico, ex Conostegia xalapensis D. Don (Melastomataceae). Type Depository: USNM. Note: host originally listed as Conostegia talapensis (Bump.) D. Don.

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509. Phytoptipalpus harveyi Meyer & Van Dis, 1993: 326; replacement to Aegyptobia albiziae Meyer (1979), name preoccuped by Phytoptipalpus albizziae Pritchard & Baker (1958). (Article 58.7 4th Edition International Code of Zoological Nomenclature). Objective synonym of: Aegyptobia albiziae Meyer—Meyer & Van Dis (1993). 509a. Aegyptobia albiziae Meyer, 1979: 125; Shingwedzi Rest Camp, Kruger National Park, Transvaal, South Africa, ex Albizia harveyi Fourn. (Mimosaceae). Type Depository: SANC. 510. Phytoptipalpus kalahariensis (Meyer, 1979): 121; Kalahari Gemsbok National Park, Cape Province, South Africa, ex Rhigozum trichotomum Burch. (Bignoniaceae); other hosts listed included: Monechma genistifolium C.B. Clarke subsp. australe (P.G. Mey.) J. Munday (Acanthaceae) and Protasparagus pearsonii (Kies) Oberm. (Asparagaceae). Species designation: Aegyptobia kalahariensis—original designation. Redescriptions: Meyer & Ueckermann (1989), Meyer & Van Dis (1993). Type Depository: SANC. 511. Phytoptipalpus lanzhouensis Ma & Yuan, 1981: 194; Lanzhou, China, ex Rosa omeiensis Rolfe (Rosaceae). Type Depository: Shanghai College of Agriculture, China. 512. Phytoptipalpus lepsis Hasan, Ashfaq & Li, 2003: 528; 1 km E Attock, Faisalabad, Pakistan, ex Datura inoxia P. Mill. (Solanaceae). Type Depository: Acarology Research Laboratory, Department of Entomology, University of Agriculture, Faisalabad, Pakistan. Note: host originally listed as Datura metel L. 513. Phytoptipalpus lithos Hasan, Ashfaq & Li, 2003: 527; Kabirwala (Khanewal), Pakistan, ex Mangifera indica L. (Anacardiaceae). Type Depository: Acarology Research Laboratory, Department of Entomology, University of Agriculture, Faisalabad, Pakistan. 514. Phytoptipalpus multistriatum (Meyer, 1979): 123; Pretoria, Transvaal, South Africa, ex Acacia karroo Hayne (Mimosaceae). Species designations: Aegyptobia multistriatum—original designation, Phytoptipalpus multistriatum—Meyer & Van Dis (1993). Redescriptions: Meyer & Ueckermann (1989), Meyer & Van Dis (1993). Type Depository: SANC. 515. Phytoptipalpus muukuorum Meyer, 1979: 112; Pienaarspoort, 16 km from Pretoria, Transvaal, South Africa, ex Acacia karroo Hayne (Mimosaceae). Redescription: Meyer & Van Dis (1993). Type Depository: SANC. 516. Phytoptipalpus nyala (Meyer, 1979): 123; Nyala, Kruger National Park, Transvaal, South Africa, ex Dichrostachys cinerea (L.) subsp. cinerea (L.) Wight & Arn. (Mimosaceae). Species designations: Aegyptobia nyala—original designation, Phytoptipalpus nyalai [sic]—Meyer & Van Dis (1993). Redescription: Meyer & Van Dis (1993). Type Depository: SANC. 517. Phytoptipalpus paradoxus Trägard, 1904: 20; Kaka, White Nile, Sudan, ex Acacia sp. (Mimosaceae). Redescriptions: Meyer (1979), Meyer & Van Dis (1993), Mitrofanov (1979), Pritchard & Baker (1958), Sayed (1942a), Zaher & Yousef (1969), Zaher et al. (1984). Type Depository: Unknown. 518. Phytoptipalpus rosae (Mitrofanof & Strunkova, 1978): 1097; Tajikistan, ex Rosa sp. (Rosaceae). Species designation: Zaheria rosae—original designation, Phytoptipalpus rosae—Meyer (1979). Redescription: Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: State Nikitsky Botanical Gardens (Yalta) and Institute of Zoology and Parasitology, Academy of Sciences of the Tadjik SSR (Dushanbe), No. 2552/366

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519. Phytoptipalpus xianensis Ma & Yuan, 1981: 194; China, ex Buxus microphylla Siebold & Zucc. (Buxaceae). Type Depository: Shanghai College of Agriculture, China.

Priscapalpus De Leon 1961 Priscapalpus De Leon, 1961b: 93. Type species: Priscapalpus macropilis De Leon, 1961: 93; Baker, Tuttle & Abbatiello, 1975: 22; Baker & Tuttle, 1987: 118; De Leon, 1965a: 65; Meyer, 1979: 97; Mitrofanov, 1973a: 510; Mitrofanov & Strunkova, 1979: 77. Deleoniella Mitrofanov, 1973a: 509. Type species: Priscapalpus cherretti De Leon, 1965; synonymy according to Meyer 1979: 97; Sepasgosarian (1983), Baker & Tuttle (1987). 520. Priscapalpus cherretti De Leon, 1965a: 74; near Bartica Nature Reserve, Guyana (= British Guiana), ex Miconia sp. (Melastomataceae). Species designation: Deleoniella cherretti—Mitrofanov (1973a). Redescriptions: Meyer (1979), Mitrofanov (1973), Mitrofanov & Strunkova (1979), Sadana & Sidhu (1989). Type Depository: MCZ. 521. Priscapalpus macropilis De Leon, 1961a: 94; Puerto Vallarta, Jalisco, Mexico, ex "sapodilla" (Sapotaceae). Redescriptions: Baker & Tuttle (1987), Meyer (1979), Mitrofanov (1973), Mitrofanov & Strunkova (1979), Sadana & Sidhu (1989). Type Depository: MCZ. Note: According to Baker et al (1975) the type host probably is Achras zapota L. (Sapotaceae). 522. Priscapalpus thomissus Meyer, 1979: 97; Wolkberg, near Haenertsburg, Transvaal, South Africa, ex Rhoicissus tridentata (L.f.) Wild & Drummond (Vitaceae). Redescription: Sadana & Sidhu (1989). Type Depository: SANC.

Prolixus Beard, Fan & Walter 2005 Prolixus Beard, Fan & Walter, 2005: 164. Type species: Prolixus forsteri Beard, Fan & Walter 2005. 523. Prolixus corruginus Beard, Fan & Walter, 2005: 163; Wondul Stock Route, Wondul Range, near Inglewood, Queensland, Australia, ex. sedge, Gahnia aspera sensu lato (Cyperaceae). Type Depository: QM. 524. Prolixus forsteri Beard, Fan & Walter, 2005: 170; State Forest 1294, Queensland, Australia, ex. sedge, Gahnia aspera sensu lato (Cyperaceae). Type Depository: QM.

Pseudoleptus Bruyant 1911 Pseudoleptus Bruyant, 1911: 340. Type species: Pseudoleptus arechavaletae Bruyant, 1911: 340; Baker & Yin, 1988: 150; Baker & Tuttle, 1972: 26; Baker & Tuttle, 1987: 119; Baker et al. 1975: 1; Duzgunes, 1965: 124; Meyer, 1979: 128; Mitrofanov, 1973b: 1317; Mitrofanov & Strunkova, 1979: 111; Pritchard & Baker, 1952: 6; Pritchard & Baker, 1958: 184; Wainstein, 1960: 234. 525. Pseudoleptus arechavaletae Bruyant, 1911: 340; Montevideo, Uruguay, ex Distichlis scoparia Arechav. (Poaceae). Redescriptions: Akbar & Chaudhri (1983), Baker & Pritchard (1952b), Lahille (1927), Meyer (1979), Mitrofanov & Strunkova (1979), Pritchard & Baker (1958). Type Depository: USNM. Synonym: 525a. Tenuipalpus haumani Lahille, 1927: 671; Argentina, ex Distichlis spicata (L.) Greene (Poaceae)—Pritchard & Baker (1958). Type Depository: Unknown or not specified.

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526. Pseudoleptus bouteloua Baker & Tuttle, 1972: 28; McNary, Arizona, USA, ex Bouteloua gracilis (Kunth) Lagasca ex. Griffiths (Poaceae). Species designation: Pseudoleptus boutelone [sic]—Akbar & Chaudhri (1983). Redescriptions: Akbar & Chaudhri (1983), Baker & Pritchard (1952b), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: USNM, No. 3432. 527. Pseudoleptus graminicola Mitrofanov & Sharonov, 1983; Ukraine, ex an unidentified plant. Type Depository: Unknown. 528. Pseudoleptus graminosus Meyer, 1979: 128; Potchefstroom, Transvaal, South Africa, ex an unidentified grass. Redescription: Akbar & Chaudhri (1983). Type Depository: SANC. 529. Pseudoleptus hilaria Baker & Tuttle, 1972: 27; Dateland, Arizona, USA, ex Hilaria rigida (Thurber) (Poaceae). Redescriptions: Akbar & Chaudhri (1983), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: USNM, No. 3431. 530. Pseudoleptus palustria Pritchard & Baker, 1952: 7; Davis, California, USA, ex Distichlis spicata (L.) Greene (Poaceae). Redescriptions: Akbar & Chaudhri (1983), Baker & Tuttle (1964, 1987), Meyer (1979), Mitrofanov & Strunkova (1979), Pritchard & Baker (1958). Type Depository: USNM, No. 1929. 531. Pseudoleptus panicum Baker & Tuttle, 1972: 26; Portal, Arizona, USA, ex Panicum obtusum H, B & K. (Poaceae). Redescriptions: Akbar & Chaudhri (1983), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: USNM, No. 3429. 532. Pseudoleptus peshawariensis Akbar & Chaudhri, 1983: 49; 4.8 km N Peshawar, Pakistan, ex an unidentified plant. Type Depository: Acarology Research Laboratory, Department of Agric. Entomology, University of Agriculture, Faisalabad, Pakistan. 533. Pseudoleptus tridens Baker & Tuttle, 1972: 26; Picacho, Arizona, USA, ex Tridens pulchellus (H,B &K) Hitchcock (Poaceae). Redescriptions: Akbar & Chaudhri (1983), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: USNM, No. 3430. 534. Pseudoleptus zelihae Pritchard & Baker, 1958: 185; Turkey, intercepted at Washington, District of Columbia, USA, ex Aeluropus sp. (Poaceae). Redescriptions: Akbar & Chaudhri (1983), Düzgünes (1965), Hatzinikolis (1987), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: USNM, No. 2292.

Raoiella Hirst 1924 Raoiella Hirst, 1924: 522. Type species: Raoiella indica Hirst, 1924: 522; Baker & Pritchard, 1960: 570; Chaudhri et al., 1974: 17; Hatzinikolis, 1987: 57; Meyer, 1979: 114; Meyer & Gerson, 1980: 76; Mitrofanov, 1973b: 1316; Mitrofanov & Strunkova, 1979: 120; Pritchard & Baker, 1958: 256; Sayed, 1938: 605; Sayed, 1942: 81; Sayed, 1950: 1016; Smiley & Gerson, 1995: 39; Wainstein, 1960: 234. Neoraoiella Mohanasundaram, 1996: 141. Type species: Neoraoiella eugenia Mohanasundaram, 1996: 141—new synonym. Rarosiella Rimando, 1996: 2. Type species: Rarosiella cocosae Rimando 1996:3, new synonym. 535. Raoiella australica Womersley, 1940: 263; Dee Why, New South Wales, Australia, ex Eucalyptus spp. (Myrtaceae). Redescriptions: Akbar (1990), Akbar & Chaudhri (1987), Chaudhri & Akbar (1985), Meyer (1979), Mitrofanov & Strunkova (1979), Mohanasundaram (1989), Pritchard & Baker (1958), Smiley & Gerson (1995). Type Depository: Unknown.

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536. Raoiella camur Chaudhri & Akbar, 1985:18; 12.8 km E Karachi, Pakistan, ex Phoenix dactylifera L. (Arecaceae). Redescriptions: Akbar (1990), Akbar & Chaudhri (1987). Type Depository: Department Entomology, University of Agriculture, Faisalabad, Pakistan. Note: suspected junior synonym of Raoiella indica Hirst. Synonym: 536a. Raoiella camur Akbar, 1990:78; 12.8 km E Karachi, ex Phoenix dactylifera L. (Arecaceae)—new synonym. Note: The drawings of the original description of R. camur Akbar (1990) is exactly the same as those of the original description of R. camur Chaudhri & Akbar (1985). The site of collection and host are the same. Thus, despite the differences in the indicated collectors, it is assumed that the species are the same. Type Depository: Department of Entomology, University of Agriculture, Faisalabad, Pakistan. 537. Raoiella empedos Chaudhri & Akbar, 1985: 16; 6.4 km E Faisalabad, Pakistan, ex Phoenix dactylifera L. (Arecaceae). Redescriptions: Akbar & Chaudhri (1987). Type Depository: Department of Entomology, University of Agriculture, Faisalabad, Pakistan. Note: suspected junior synonym of Raoiella indica Hirst. Objective synonym: 537a. Raoiella empedos Akbar, 1990:75; 6.4 km E Faisalabad, ex Phoenix dactylifera L. (Arecaceae). Note: The drawings of this species in the original description of R. empedos Akbar (1990) are exactly the same as those of the original description of R. empedos Chaudhri & Akbar (1985). In addition, the collection site and the host of the specimens given in the description in each of these publications are the same. Thus, despite the differences in the indicated collectors, we consider the species to be the same. Type Depository: Department of Entomology, University of Agriculture, Faisalabad, Pakistan. Note: suspected junior synonym of Raoiella indica Hirst. 538. Raoiella eugenia (Mohanasundaram 1996): 141—new combination; Puthur, Mangalore, Karnataka, India, ex Eugenia sp. (Myrtaceae) Species designation: Neoraoiella eugenia Mohanasundaram 1996: 141. Type Depository: TNAU. 539. Raoiella indica Hirst, 1924: 522; Coimbatore, Tamil Nadu, India, ex coconut (Cocos nucifera, Arecaceae). Suspected senior synonym of: Raoiella camur Chaudhri & Akbar, Raoiella empedos Chaudhri & Akbar, Raoiella neotericus Chaudhri & Akbar, Raoiella obelias Hasan & Akbar, Raoiella pandanae Mohanasundaram, Raoiella phoenica Meyer, Raoiella rahii Akbar & Chaudhri. Redescriptions: Akbar (1990), Akbar & Chaudhri (1987), Al-Gboory (1987), Chandra & Channabasavanna (1974), Chaudhri & Akbar (1985), Chaudhri et al. (1974), Gupta (1984, 1992), Meyer (1979), Mitrofanov & Strunkova (1979), Mohanasundaram (1989), Pritchard & Baker (1958), Sadana (1997), Sayed (1942). Type Depository: BMNH. Synonymy: 539a. Rarosiella cocosae Rimando, 1996: 3; Camiguin Island, off Misamis Oriental, in north central Mindanao, Philippines, ex coconut (Arecaceae)—new synonym. Type Depository: University of the Philippines Los Baños, College, Lagua, Department of Entomology and Museum of Natural History, The Philippines. 540. Raoiella macfarlanei Pritchard & Baker, 1958: 257; Rasel Hila, Cyrenaica, Greece, ex olive (Oleaceae). Redescriptions: Akbar (1990), Akbar & Chaudhri (1987), Chandra & Channabasavanna (1974), Chaudhri & Akbar (1985), Hatzinikolis (1987), Meyer (1979), Mitrofanov & Strunkova (1979), Mohanasundaram (1989), Nageshachandra & Channabasavanna (1974), Sadana (1997). Type Depository: BMNH. 541. Raoiella neotericus Chaudhri & Akbar, 1985: 19; University of Agriculture of Faisalabad, Faisalabad, Pakistan, ex Phoenix dactylifera L. (Arecaceae). Redescriptions: Akbar (1990), Akbar & Chaudhri (1987). Type Depository: Department Entomology, University of Agriculture, Faisalabad, Pakistan. Note: suspected junior synonym of Raoiella indica Hirst. Synonym: 541a. Raoiella neotericus Akbar, 1990:78; 6.4 km ; University of Agriculture of Faisalabad, Faisalabad, ex Phoenix dactylifera L. (Arecaceae).—new synonym. Note: The drawings of the original description, the collection locality and host plant of R. neotericus Akbar (1990)

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are exactly the same as those found in the original description of R. neotericus Chaudhri & Akbar (1985). Thus, despite the differences in the names of the collectors listed for each species, we consider that the two species are the same. Type Depository: Department of Entomology, University of Agriculture, Faisalabad, Pakistan. 542. Raoiella obelias Hasan & Akbar, 2000: 11; Jang, Pakistan, ex Phoenix dactylifera L. (Arecaceae). Type Depository: Acarology Research Laboratory, Department of Entomology, University of Agriculture, Faisalabad, Pakistan. Note: suspected junior synonym of Raoiella indica Hirst. 543. Raoiella pandanae Mohanasundaram, 1985: 31; S Arcot, Valadur, Tamil Nadu, India, ex Pandanus sp. (Pandanaceae). Redescription: Sadana (1997). Type Depository: TNAU. Note: suspected junior synonym of Raoiella indica Hirst. 544. Raoiella phoenica Meyer, 1979: 114; Nuri, Sudan, ex Phoenix dactylifera L. (Arecaceae). Redescriptions: Akbar (1990), Akbar & Chaudhri (1987), Mohanasundaram (1989). Type Depository: SANC. Note: suspected junior synonym of Raoiella indica Hirst. 545. Raoiella rahii Akbar & Chaudhri, 1987: 42; University of Agriculture of Faisalabad, Faisalabad, Pakistan, ex Phoenix dactylifera L. (Arecaceae). Redescription: Akbar (1990). Type Depository: Department of Entomology, University of Agriculture, Faisalabad, Pakistan. Note: suspected junior synonym of Raoiella indica Hirst. 546. Raoiella shimapana Meyer, 1979: 115; Ngwanetsi River, Kruger National Park, Transvaal, South Africa, ex Cassine transvaalensis (Burtt Davy) Codd (Celastraceae). Redescriptions: Akbar (1990), Akbar & Chaudhri (1987), Chaudhri & Akbar (1985), Mohanasundaram (1989). Type Depository: SANC.

Raoiellana Baker & Tuttle 1972 Raoiellana Baker & Tuttle, 1972: 32. Type species: Raoiellana allium Baker & Tuttle, 1972: 32; Meyer, 1979: 114, Mitrofanov, 1973b: 1316, Mitrofanov & Strunkova, 1979: 120. 547. Raoiellana allium Baker & Tuttle, 1972: 32; Turkey, intercepted at New York, USA, ex Allium pulchellum G.Don on the bulbs (Alliaceae). Redescription: Mitrofanov & Strunkova (1979). Type Depository: USNM, No. 3437.

Tegopalpus Womersley 1940 Tegopalpus Womersley, 1940: 242. Type species: Tegopalpus conicus Womersley, 1940: 242; Meyer, 1979: 2; Mitrofanov, 1973a: 510; Mitrofanov & Strunkova, 1979: 75; Pritchard & Baker, 1958: 254; Smiley & Gerson, 1995: 39; Sayed, 1950: 1016; Smiley, Frost & Gerson, 1996: 168; Wainstein, 1960: 235. 548. Tegopalpus conicus Womersley, 1940: 242; Avalon Beach, New South Wales, Australia, ex Casuarina sp. (Casuarinaceae). Redescriptions: Meyer (1979), Mitrofanov & Strunkova (1979), Pritchard & Baker (1958), Smiley & Gerson (1995), Smiley et al. (1996). Type Depository: SAM, I. N 1970400. Note: according to Smiley et al. (1996) T. conicus, was briefly described as “probably female” (Womersley, 1940); however, a comparison of Womersley’s drawings with those of Smiley et al. (1996) clearly shows that original description was of a male.

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Tenuilichus Mohanasundaram 1988 Tenuilichus Mohanasundaram, 1988: 235. Type species: Tenuilichus malvae Mohanasundaram 1988: 235.

549. Tenuilichus malvae Mohanasundaram, 1988: 235; Kallar, Tamil Nadu, India, ex malvaceous tree (Malvaceae). Type Depository: TNAU. 550. Tenuilichus striatus Mohanasundaram, 1988: 237; Kallar, Tamil Nadu, India, ex an unidentified tree. Type Depository: TNAU.

Tenuipalpus Donnadieu 1875 Tenuipalpus Donnadieu, 1875: 139; Type species: Tenuipalpus palmatus Donnadieu, 1875: 139; Andre, 1953: 464; Baker, 1945: 34; Baker & Pritchard, 1953: 317; Baker & Pritchard, 1960: 564; Baker & Tuttle, 1964: 73; Baker & Tuttle, 1972: 31; Baker, Tuttle & Abbatiello, 1975: 4; Baker & Tuttle, 1987: 120; Chaudhri et al. 1974: 60; Collyer, 1973c: 915; De Leon, 1965: 65; Dosse, 1971: 582; Duzgunes, 1965: 124; Evans et al. 1993: 129; Gonzalez, 1968: 38; Hatzinikolis, 1987: 56; Livschitz & Mitrofanov, 1967: 9; Lo, 1969: 99; Lo, 1986: 276; Ma & Yuan, 1980: 118; Maninder & Ghai, 1978: 243; McGregor, 1949: 4; Meyer, 1979: 3; Meyer, 1993: 1; Meyer & Gerson, 1980: 68; Mitrofanov, 1973b: 1318; Mitrofanov & Strunkova, 1979: 41; Mitrofanov et al. 1975: 15; Nassar & Ghai, 1981: 381; Pritchard & Baker, 1952: 41; Pritchard & Baker, 1958: 235; Rimando, 1962: 41; Sayed, 1942: 93; Sayed, 1950: 1016; Smiley & Gerson, 1995: 39; Wainstein, 1960: 235; Womersley, 1940: 236. Aegyptopalpus Mitrofanov, 1973b: 1318. Type species: Tenuipalpus granati Sayed, 1946, synonomy according to Meyer 1979:3; Sepasgosarian (1983). Colopalpus Pritchard & Baker, 1958: 258. Type species: Tenuipalpus caudatus Dugés (= T. palmatus Donnadieu), synonomy according to Meyer 1979:3; Sepasgosarian (1983), Baker & Tuttle (1987). Deleonipalpus Mitrofanov, 1973b: 1319. Type species: Tenuipalpus barticanus De Leon, 1965, synonomy according to Meyer 1979:4; Sepasgosarian (1983). Extenuipalpus Reck, 1959. Type species: Tenuipalpus quadrisetosus Lawrence, 1940; synonomy according to Meyer 1979:3; Sepasgosarian (1983). Gnathopalpus Mitrofanov, 1973b: 1318. Type species: Tenuipalpus rosae Kadzhaja, 1955, synonomy according to Meyer 1979:4; Sepasgosarian (1983). Tuttlepalpus Mitrofanov, 1973b: 1318. Type species: Tenuipalpus trisetosus Baker and Tuttle, 1964, synonomy according to Meyer 1979:3; Sepasgosarian (1983). 551. Tenuipalpus aboharensis Sadana & Chhabra, 1980: 151; Abohar, Punjab, India, ex Punica granatum L. (Punicaceae). Redescription: Sadana (1997), Sadana & Gupta (1984). Type Depository: PAUP. 552. Tenuipalpus abutiloni Meyer, 1993: 34; Die Eiland near Tzaneen, Transvaal, South Africa, ex Abutilon austroafricanum Hochr. (Malvaceae). Type Depository: SANC. 553. Tenuipalpus acaciae Ryke & Meyer, 1960: 562; Potchefstroom, Transvaal, South Africa; ex Acacia karroo Hayne (Mimosaceae). Redescription: Meyer (1979, 1993). Type Depository: Collection of the Institute for Zoological Research, Potchefstroom University, South Africa. 554. Tenuipalpus acacii Maninder & Ghai, 1978: 245; Indian Agricultural Research Institute, New Delhi, India, ex Acacia arabica (Lam.) Muhl. & Willd. (Mimosaceae). Redescriptions: Sadana (1997), Sadana & Chhabra (1980), Sadana & Gupta (1984). Type Depository: INPC. 555. Tenuipalpus acritus Meyer, 1993: 8; SE Penge, Sekororo, Natal, Transvaal, South Africa, ex Pouzolzia mixta Solms (Urticaceae). Type Depository: SANC.

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556. Tenuipalpus acuminatae Mohanasundaram, 1983b: 163; S Arcot, Vanniarpalayan, Tamil Nadu, India, ex Tiliacora acuminata Miers (Menispermaceae) Redescription: Sadana (1997). Type Depository: TNAU. Note: host originally listed as Teliacora acuminata. 557. Tenuipalpus aethiopicus Meyer, 1979: 48; Shingwedzi River, Kruger National Park, Transvaal, South Africa, ex Acacia xanthophloea Benth. (Mimosaceae). Redescription: Meyer (1993). Type Depository: SANC. 558. Tenuipalpus africanus Meyer, 1979: 37; Enkeldoornspoort, near Rust de Winter Nature Reserve, Transvaal, South Africa, ex Peltophorum africanum Sond. (Caesalpiniaceae). Redescription: Meyer (1993). Type Depository: SANC. 559. Tenuipalpus albae Meyer, 1979: 66; Chianga, Angola, ex Eclipta prostrata (L.) L. (Asteraceae). Redescription: Meyer (1993). Type Depository: SANC. 560. Tenuipalpus alpinus Collyer, 1973c: 946; Mount Robert Ridge circa 1.1 km, Nelson Lakes National Park, New Zealand, ex mat plants on rocks. Species designation: Tuttlepalpus alpinus—Mitrofanov & Strunkova (1979). Redescriptions: Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: Entomology Division, Department of Scientific and Industrial Research, Auckland, New Zealand. 561. Tenuipalpus amatikulensis Meyer, 1993: 50; KwaZulu, Amatakulu Nature Reserve, Gingindlovu, Natal, South Africa, ex Aspilia natalensis (Sond.) Wild (Asteraceae). Type Depository: SANC. 562. Tenuipalpus amygdalusae Maninder & Ghai, 1978: 245; Maharashtra, Bombay, Chembur, Punjab, India, ex Amygdalus communis L. (Rosaceae). Redescriptions: Sadana (1997), Sadana & Chhabra (1980), Sadana & Gupta (1984). Type Depository: INPC. Note: host originally listed as Amygdalus cammunis. 563. Tenuipalpus anacardii De Leon, 1965a: 67; about 4.8 km S Bartica, Potaro Road, Guyana (=British Guiana), ex Anacardium sp. (probably officinale) (Anacardiaceae). Redescriptions: Collyer (1973b), De Leon (1967), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: MCZ. 564. Tenuipalpus angolensis Meyer, 1979: 56; Ceilunga, Angola, ex Phyllanthus sp. (Euphorbiaceae). Redescription: Meyer (1993). Type Depository: SANC. 565. Tenuipalpus annonae De Leon, 1957: 91 Matias Romero, Oaxaca, Mexico, ex Annona sp. (Annonaceae). Species designation: Tuttlepalpus annonae—Mitrofanov & Strunkova (1979). Redescriptions: Baker & Tuttle (1987), Collyer (1973b), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: MCZ. 566. Tenuipalpus anoplomexus Baker & Tuttle, 1987: 123; north of Tehuantepec, Oaxaca, Mexico, ex Swietenia macrophylla King (Meliaceae). Type Depository: USNM. 567. Tenuipalpus anoplus Baker & Pritchard, 1953b: 320; Coconut Grove, Florida, USA, ex Swietenia mahogani DC. (Meliaceae). Redescriptions: Baker & Tuttle (1987), Collyer (1973b), De Leon (1957, 1965b, 1967), Lo (1969), Meyer (1979), Mitrofanov & Strunkova (1979), Pritchard & Baker (1958). Type Depository: USNM, No. 1999.

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Synonym: 567a. Tenuipalpus swieteniae Livschitz in Livschitz & Salinas, 1968: 122; Santiago de las Vegas, Cuba, ex Swietenia macrophylla King (Meliaceae)—Mitrofanov & Strunkova (1979). Type Depository: Unknown. 568. Tenuipalpus antipodus Collyer, 1964: 436; Little Huia, near Auckland, New Zealand, ex Melicytus ramiflorus J.R. Forst. & G. Forst. (Violaceae). Redescriptions: Collyer (1973b), Lo (1969, 1986), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: BMNH. 569. Tenuipalpus arbuti Mitrofanov & Sharonov, 1983: 948; Ukraine, ex Arbutus unedo L. (Ericaceae). Redescription: Hatzinikolis (1987). Type Depository: Unknown. 570. Tenuipalpus argus Baker & Pritchard, 1953b: 328; Pomona Park, Florida, USA, ex Yucca gloriosa L. (Agavaceae). Redescriptions: Collyer (1973b), Meyer (1979), Mitrofanov & Strunkova (1979), Pritchard & Baker (1958). Type Depository: USNM, No. 2002. 571. Tenuipalpus ariauae Feres & Hernandes, 2006: 64; “Sítio” Sao Luiz, Sao José do Rio Preto, Sao Paulo, Brazil, ex Qualea grandiflora Mart. (Vochysiaceae). Type Depository: Collection of Acari Departamento de Zoologia e Botanica, UNESP, Sao Josè do Rio Preto, State of Sao Paulo, Brazil. 572. Tenuipalpus athrixiae Meyer, 1993: 53; Pretoria, Transvaal, South Africa, ex Athrixia elata Sond. (Asteraceae). Type Depository: SANC. 573. Tenuipalpus attiahi Baker & Pritchard, 1960: 564; Lubero, Kivu, Democratic Republic of Congo (formerly Zaïre), ex Podocarpus usambarensis Pilg. (Podocarpaceae). Redescriptions: Collyer (1973b), Meyer (1979), Meyer (1993), Mitrofanov et al. (1975). Type Depository: USNM, No. 2503. 574. Tenuipalpus aurantiacus Wang, 1983: 61; Yuanming Park, Beijing, China, ex Viola philippica Cav. (Violaceae). Type Depository: Institute of Zoology, Academia Sinica, China. 575. Tenuipalpus auriculatae Meyer, 1979: 17; Alice, Cape Province, South Africa, ex Plumbago auriculata Lam. (Plumbaginaceae). Redescription: Meyer (1993). Type Depository: SANC. 576. Tenuipalpus austrocedri Gonzalez, 1968: 41; Cajon del Rio Claro, Cordillera de Curico, Chile, ex Austrocedrus chilensis (D. Don) Florin & Boutelje (Cupressaceae). Redescriptions: Collyer (1973b), Meyer (1979), Mitrofanov et al. (1975), Mitrofanov & Strunkova (1979). Type Depository: Coleccion de la Estacion Experimental Agronomica, Universidad de Chile (CEA). 577. Tenuipalpus baeri Reck, 1956: 24; Georgia, ex Lagonychium farctum (Banks & Sol.) Bobrov. (Mimosaceae). Redescriptions: Collyer (1973b), Livschitz & Mitrofanov (1967), Meyer (1979), Mitrofanov & Strunkova (1979), Pritchard & Baker (1958). Type Depository: Unknown. 578. Tenuipalpus bagdadensis Al-Gboory, 1987: 128; Abu-Ghraib, Baghdad, Iraq, ex Lagonychium farctum (Banks & Sol.) Bobrov. (Mimosaceae). Type Depository: Entomology Museum, College of Agriculture, Department of Plant Protection, Abu-Ghraib, Baghdad, Iraq. 579. Tenuipalpus bakerdeleonorum Evans, in Evans et al (1993): 148; Rio Telica, near Juticalpa, Olancho, Honduras, ex Chromolaena odorata (L.) King & H.E. Robins. (Asteraceae). Type Depository: USNM. Note: host originally listed as “Chromolaena (= Eupatorium odorata)”.

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580. Tenuipalpus bakeri McGregor, 1949: 7; Cocoa Beach, Florida, USA, ex Magnolia sp. (Magnoliaceae), oak, Sobralia macrantha Lindl. (Orchidaceae) and Yucca gloriosa L. (Agavaceae). Redescriptions: Baker & Pritchard (1953b), Baker & Tuttle (1987), Collyer (1973b), De Leon (1957, 1965b, 1967), Evans et al. (1993), Mairena & Ochoa (1989), Meyer (1979), Mitrofanov & Strunkova (1979), Pritchard & Baker (1958). Type Depository: USNM, No. 1704. 581. Tenuipalpus banahawensis Corpuz-Raros, 1978: 221; Mount Banahaw, Sariaya, Quezon, Philippines, ex (Myrsinaceae). Type Depository: USNM. 582. Tenuipalpus banksiae Gutierrez & Schicha, 1982: 137; New South Wales, Australia, ex Banksia spinulosa Sm. (Proteaceae). Redescription: Smiley & Gerson (1995). Type Depository: Biological and Chemical Research Institute, New South Wales, Department of Agriculture, Rydalmere, New South Wales, Australia. 583. Tenuipalpus barticanus De Leon, 1965a: 67; 8 km S Bartica, Potaro Road, Guyana (=British Guiana), ex shrub. Species designation: Deleonipalpus barticanus—Mitrofanov & Strunkova (1979). Redescriptions: Collyer (1973b), De Leon (1967), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: MCZ. 584. Tenuipalpus bassiae Mohanasundaram, 1988: 238; Coimbatore, Tamil Nadu, India, ex Bassia latifolia Roxb. (Chenopodiaceae). Type Depository: TNAU. 585. Tenuipalpus bellulus Meyer, 1993: 10; Wyllie's Poort, near Louis Trichardt, Transvaal, South Africa, ex Albizia harveyi Fourn. (Mimosaceae). Type Depository: SANC. 586. Tenuipalpus berkheyae Meyer, 1993: 47; Pretoria, Transvaal, South Africa, ex Berkheya radula (Harv.) Burtt Davy (Asteraceae). Type Depository: SANC. 587. Tenuipalpus boninensis Ehara, 1982: 177; Chichi-Jima Island, the Bonin Islands, Japan, ex Morinda boninensis Ohwi (Rubiaceae). Type Depository: BITU. 588 Tenuipalpus boyani De Leon, 1965a: 67; Bartica Nature Reserve, Guyana (= British Guiana), ex Pouteria sp. (Sapotaceae). Redescriptions: Collyer (1973b), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: MCZ. 589. Tenuipalpus bucidae De Leon, 1956a: 57; Fairchild Tropical Garden, Coconut Grove, Florida, USA, ex Bucida buceras L. (Combretaceae). Redescriptions: Collyer (1973b), Meyer (1979), Mitrofanov & Strunkova (1979), Pritchard & Baker (1958). Type Depository: USNM. 590. Tenuipalpus burserae De Leon, 1957: 90; Ocozocoatla, Chiapas, Mexico, ex Bursera sp. (Burseraceae). Redescriptions: Baker & Tuttle (1987), Collyer (1973b), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: MCZ. 591. Tenuipalpus calcarius Meyer, in Meyer & Ueckermann (1988): 9; Mountain Zebra National Park, Cape Province, South Africa, ex Tarchonanthus camphoratus L. (Asteraceae). Redescription: Meyer (1993). Type Depository: SANC. 592. Tenuipalpus caledonicus Meyer, 1979: 63; Caledon, Cape Province, South Africa, ex Aspalathus acuminata Lam. (Fabaceae).

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Redescription: Meyer (1993). Type Depository: SANC. 593. Tenuipalpus capparis Meyer (1979): 68; Shingwidzi River, Kruger National Park, Transvaal, South Africa, ex Capparis sepiaria L. var. citrifolia (Lam.) Toelken (Capparaceae). Redescription: Meyer (1993). Type Depository: SANC. 594. Tenuipalpus capassae Meyer, 1993: 55; Pretoria, Transvaal, South Africa, ex Lonchocarpus capassa Rolfe (Fabaceae). Type Depository: SANC. 595. Tenuipalpus carlosflechtmanni Feres & Hernandes, 2006: 58; “Sítio” Santo Antônio, Sao José do Rio Preto, Sao Paulo, Brazil, ex Qualea grandiflora Mart. (Vochysiaceae) Type Depository: Collection of Acari Departamento de Zoologia e Botanica, UNESP, Sao Josè do Rio Preto, State of Sao Paulo, Brazil. 596. Tenuipalpus carolinensis Baker, 1945: 35; Batesburg, South Carolina, USA, ex “goldenrod” (Asteraceae). Redescriptions: Baker & Pritchard (1953b), Collyer (1973b), McGregor (1949), Meyer (1979), Mitrofanov & Strunkova (1979), Pritchard & Baker (1958), Reck (1951). Type Depository: USNM, No. 1479. 597. Tenuipalpus caudatus (Duges, 1834): 29, 244; France, ex “laurier-tin” Viburnum tinus L. (Caprifoliaceae). Species designation: Trombidium caudatus—original designation, Tenuipalpus caudatus—Pritchard & Baker (1958). Redescriptions: Akbar & Chaudhri (1981), Baker & Pritchard (1953b), Castagnoli & Pegazzano (1979), Collyer (1973b), Ehara & Masaki (2001), Hatzinikolis (1986a, 1987), McGregor (1949), Meyer (1979), Mitrofanov & Strunkova (1979), Pritchard & Baker (1958), Sadana (1997), Sadana & Chhabra (1980), Sadana & Gupta (1984). Type Depository: Unknown. Synonym: 597a. Acarus tini Boisduval, 1867: 91; France, ex “laurestinus” (Caprifoliaceae)—Pritchard & Baker (1958). Redescriptions: Baker (1945), Baker & Pritchard (1953), Berlese (1887), Canestrini (1890), McGregor (1949), Vitzthum (1929). Type Depository: Unknown. 597b. Caligonus calyx Canestrini & Fanzago, 1876: 134; Trevigiano, Italy, ex an unidentified plant—Pritchard & Baker (1958). Type Depository: Unknown. 597c. Tenuipalpus palmatus Donnadieu, 1875: 112; Lyon, France, ex “laurier-tin” Viburnum tinus L. (Caprifoliaceae)—Pritchard & Baker (1958). Redescription: Baker (1945), Baker & Pritchard (1953b), Pritchard & Baker (1958), Reck (1951). Type Depository: USNM. 598. Tenuipalpus cedrelae De Leon, 1957: 90; San Blas, Nayarit, Mexico, ex Cedrela sp. (Meliaceae). Species designations: Tenuipalpus cedrellae [sic] Flechtmann (1979). Redescriptions: Baker & Tuttle (1987), Collyer (1973b), Evans et al. (1993), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: MCZ. 599. Tenuipalpus celtidis Pritchard & Baker, 1958: 242; Davie, Florida, USA, ex Celtis sp. (Ulmaceae). Redescriptions: Collyer (1973b), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: USNM, No. 2297. 600. Tenuipalpus chamaedorea Salas & Ochoa, 1985: 171; La Abigail, Las Nubes de Coronado, San José, Costa Rica, ex Chamaedorea sp. (Arecaceae). Species designations: Tenuipalpus chamaedoreae [sic] Salas & Ochoa (1986), Tenuipalpus uchamaedoreae [sic]—Mairena & Ochoa (1989). Redescriptions: Salas & Ochoa (1986), Mairena & Ochoa (1989).

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Type Depository: Colecciòn de Referencia, Laboratorio Acarologia, Facultad de Agronomia, Universidad de Costa Rica. 601. Tenuipalpus cheladzeae Gomelauri, 1960: 77; Ukraine, ex Taxus baccata L. (Taxaceae). Redescriptions: Livschitz & Mitrofanov (1967), Meyer (1979), Mitrofanov et al (1975), Mitrofanov & Strunkova (1979). Type Depository: Unknown. Synonym: 601a. Tenuipalpus szarvasensis Bozai, 1970: 367; Szarvas, Hungary, ex Abies nordmanniana Spach (Pinaceae)—Mitrofanov & Strunkova (1979). Redescription: Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: Zoological Department of the Hungarian Natural History Museum, Budapest, Hungarian. 602. Tenuipalpus chelinus Meyer, 1993: 56; Blyderivierspoort Nature Reserve, Transvaal, South Africa, ex Rhynchosia monophylla Schltr (Fabaceae). Type Depository: SANC. 603. Tenuipalpus chiclorum De Leon, 1957: 91; Tuxtla Gutierrez, Chiapas, Mexico, ex Achras sapota L. (Sapotaceae). Redescriptions: Baker & Tuttle (1987), Collyer (1973b), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: MCZ. 604. Tenuipalpus chinariensis Akbar & Chaudhri, 1981: 11; Chinari, (Azad Kashmir), Pakistan, ex Punica granatum L. (Punicaceae). Type Depository: Acarology Research Laboratory, Department of Entomology, University of Agriculture, Faisalabad, Pakistan. 605. Tenuipalpus chiococcae De Leon, 1956a: 59; west of Perrine, Florida, USA, ex Chiococca pinetorum Britton & Millsp. (Rubiaceae). Redescriptions: Collyer (1973b), Meyer (1979), Mitrofanov & Strunkova (1979), Pritchard & Baker (1958). Type Depository: USNM. 606. Tenuipalpus cissampelosa Maninder & Ghai, 1978: 249; Buddha Jayanti Park, New Delhi, India, ex Cissampelos pareira L. (Menispermaceae). Redescriptions: Gupta & Chatterjee (1997), Sadana (1997), Sadana & Chhabra (1980), Sadana & Gupta (1984). Type Depository: INPC. Note: host originally listed as Cissampelosa pareira. 607. Tenuipalpus citus Chaudhri, Akbar & Rasool, 1974: 62; 1.6 km N Ochali, Sind, Pakistan, ex Salix babylonica L. (Salicaceae). Redescription: Akbar & Chaudhri (1981). Type Depository: Department of Entomology, University of Agriculture, Lyallpur, Pakistan. 608. Tenuipalpus clematidos Wang, 1983: 61; Beijing, Dongyuan, China, ex Clematis heracleifolia DC (Ranunculaceae). Type Depository: Institute of Zoology, Academia Sinica, China. Note: host originally listed as Clematis neracleifolia. 609. Tenuipalpus coccolobicoloides De Leon, 1965b: 519; Bath Fountain, Saint Thomas Parish, Jamaica, ex Psychotria grandis Sw. (Rubiaceae). Redescriptions: Collyer (1973b), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: MCZ. 610. Tenuipalpus coccolobicolus De Leon, 1956a: 58; Coral Gables, Florida, USA, ex Coccoloba laurifolia Jacq. (Polygonaceae). Redescriptions: Baker & Tuttle (1987), Collyer (1973b), De Leon (1957), Meyer (1979), Mitrofanov & Strunkova (1979), Pritchard & Baker (1958). Type Depository: USNM. Note: host originally listed as Coccolobis laurifolia.

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611. Tenuipalpus coimbatorensis Mohanasundaram, 1981: 401; Coimbatore, Tamil Nadu, India, ex an unidentified plant. Redescription: Sadana (1997), Sadana & Gupta (1984). Type Depository: TNAU. 612. Tenuipalpus comatus Meyer, 1993: 57; Nieuwoudt's Pass, Cederberg Mountains, Cape Province, South Africa, ex Phylica sp. (Rhamnaceae). Type Depository: SANC. 613. Tenuipalpus combreti Meyer, 1979: 27; Enkeldoornspoort, near Rust de Winter Nature Reserve, Transvaal, South Africa, ex Combretum molle R. Br. & G. Don (Combretaceae). Redescription: Meyer (1993). Type Depository: SANC. 614. Tenuipalpus comptus Meyer, 1979: 37; Hankey, Gamtoos Valley, Cape Province, South Africa, ex Maytenus heterophylla (Eckl. & Zeyh.) N. Robson (Celastraceae). Redescription: Meyer (1993). Type Depository: SANC. 615. Tenuipalpus costarricensis Salas & Ochoa, 1986: 203; Centro Agronomico Tropical de Investigacion y Enseñanza, Turrialba, Cartago, Costa Rica, ex Cedrela sp. (Meliaceae). Redescription: Mairena & Ochoa (1989). Type Depository: Colecciòn de referencia, Laboratorio Acarologia, Facultad de Agronomia, Universidad de Costa Rica, Costa Rica. 616. Tenuipalpus couroupita De Leon, 1967: 44; Toco, Trinidad and Tobago, ex Couroupita guianensis Aubl. (Lecythidaceae). Species designation: Tenuipalpus couroupitae [sic]—Meyer (1979). Redescription: Meyer (1979). Type Depository: MCZ. 617. Tenuipalpus coyacus De Leon, 1957: 83; San Blas, Nayarit, Mexico, ex oil palm (Arecaceae). Redescriptions: Baker & Tuttle (1987), Collyer (1973b), Evans et al. (1993), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: MCZ. 618. Tenuipalpus crassulus Baker & Tuttle, 1972: 31; Holland intercepted at Washington, D.C., USA, ex Crassula lycopodioides Lamarck (Crassulaceae). Redescriptions: Collyer (1973b), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: USNM, No. 3443. 619. Tenuipalpus crassus Andre, 1953: 466; Basses-Pyrenees, France, ex Euphorbia echinus Hokk & Coss. (Euphorbiaceae). Species designations: Tenuipalpus orchidarum crassum—original designation, Tenuipalpus crassus—Pritchard & Baker (1958), Colopalpus crassus—Mitrofanov & Strunkova (1979). Redescriptions: Chandra & Channabasavanna (1974), Channabasavanna & Lakkundi (1977), Collyer (1973b), Hatzinikolis (1986a, 1987), Maninder & Ghai (1978), Meyer (1979), Mitrofanov & Strunkova (1979), Pritchard & Baker (1958), Sadana (1997), Sadana & Chhabra (1980), Sadana & Gupta (1984). Type Depository: Unknown. 620. Tenuipalpus crescentiae De Leon, 1957: 88; San Blas, Nayarit, Mexico, ex “cuasticomate” (Bignoniaceae). Redescriptions: Baker & Tuttle (1987), Collyer (1973b), De Leon (1965b, 1967), Meyer (1979), Mitrofanov & Strunkova (1979). Note: According to Baker & Tuttle (1987) the type host is Parmentiera alata Miers (Bignoniaceae). Type Depository: MCZ. Synonym: 620a. Tenuipalpus rillus Manson, 1963: 215; Potrerillos, Honduras, ex an unidentified plant.—De Leon (1965b). Species designation: Tenuipalpus rillos [sic]—Evans et al. (1993). Redescriptions: Collyer (1973b), Evans et al. (1993), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: USNM, No. 2886.

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621. Tenuipalpus crocopontensis Meyer, 1993: 13; Crocodile Bridge, Kruger National Park, Transvaal, South Africa, ex Euclea crispa (Thunb.) Guerke (Ebenaceae). Type Depository: SANC. 622. Tenuipalpus cupressoides Meyer & Gerson, 1980: 68; Kabri, Israel, ex Cupressus sempervirens L. (Cupressaceae). Type Depository: Collection of the Department of Entomology, Faculty of Agriculture, Rehovot, Israel. 623. Tenuipalpus cyatheae Gerson & Collyer, 1984: 142; Mount Bledisloe, near Waitangi, New Zealand, ex Cyathea medullaris (G. Forst.) Sw. (Cyatheaceae). Type Depository: New Zealand Arthropod Collection of the Department of Scientific and Industrial Research, Entomology Division, Mount Albert Research Centre, Auckland, New Zealand. 624. Tenuipalpus daneshvari Khosrowshahi & Arbabi, 1997: 14; Sistan & Baluchestan province, Iran, ex Solanum melongena L. (Solanaceae). Type Depository: TUPH, No. T-1622. 625. Tenuipalpus danxianensis Yin, Cui & Lin, 1995: 61; Danxian, Hainan Province, China ex Manilkara zapota (L.) van Royen (Sapotaceae). Type Depository: Department of Plant Protection, Sheyang Agricultural University, China. Note: host originally listed as Malilkara zapota. 626. Tenuipalpus dasples Baker & Pritchard, 1953b: 324; Oviedo, Florida, USA, ex Sabal megacarpa (Chapm.) Small (Arecaceae). Species designation: Tenuipalpus daples—Mitrofanov & Strunkova (1979). Redescriptions: Baker & Tuttle (1987), Collyer (1973b), De Leon (1957), Meyer (1979), Mitrofanov & Strunkova (1979), Pritchard & Baker (1958). Type Depository: USNM, No. 2001. 627. Tenuipalpus decus Chaudhri, Akbar & Rasool, 1974: 91; 1.6 km E Marghzar, Swat, Pakistan, ex Olea cuspidata Wall. & G. Don (Oleaceae). Redescriptions: Akbar & Chaudhri (1981), Sadana (1997). Type Depository: Department of Entomology, University of Agriculture, Lyallpur, Pakistan. 628. Tenuipalpus dimensus Chaudhri, 1971b: 205; Multan, Punjab, Pakistan, ex Jasminum humile L. (Oleaceae). Species designation: Tenuipalpus odimensus [sic]—Sadana & Chhabra (1980). Redescriptions: Akbar & Chaudhri (1981), Chaudhri (1972c), Chaudhri et al. (1974), Collyer (1973b), Gupta & Chatterjee (1997), Maninder & Ghai (1978), Meyer (1979), Mitrofanov & Strunkova (1979), Nassar & Ghai (1981), Sadana (1997), Sadana & Chhabra (1980), Sadana & Gupta (1984). Type Depository: Department of Entomology, West Pakistan Agricultural University Lyallpur, Pakistan. Synonym: 628a. Tenuipalpus ghanii Kazimi, 1972: 517; Rawalpindi, Pakistan, ex Pyrus communis L. (Rosaceae)—Mitrofanov & Strunkova (1979). Redescriptions: Akbar & Chaudri (1981), Meyer (1979). Type Depository: Unknown. Note: Suspected senior synonym of Tenuipalpus pyrusae Maninder & Ghai and Tenuipalpus raptor Akbar & Chaudhri. 629. Tenuipalpus disparilis Wang & Cui, 1993: 112; Leishan, Guizhou, China, ex bamboo (Poaceae) Type Depository: Institute of Zoology, Acadaemia Sinica, China. (No. 2232). 630. Tenuipalpus dominguensis De Leon, 1965b: 521; San Cristobal, Dominican Republic, ex Catalpa longissima (Jacq.) Dum.-Cours. (Bignoniaceae). Redescriptions: Collyer (1973b), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: MCZ. 631. Tenuipalpus dombeyae Meyer, 1993: 13; Nelspruit, Transvaal, South Africa, ex Dombeya rotundifolia Planch. (Sterculiaceae). Type Depository: SANC.

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632. Tenuipalpus dubinini Reck, 1951: 296; Georgia, ex Ephedra procera Fisch. & Mey. (Ephedraceae). Species designation: Colopalpus dubinini—Mitrofanov & Strunkova (1979). Redescriptions: Bagdasarian (1962), Collyer (1973b), Livschitz & Mitrofanov (1967), Meyer (1979), Meyer & Gerson (1980), Mitrofanov & Strunkova (1979), Pritchard & Baker (1958). Type Depository: Unknown. 633. Tenuipalpus dumus Meyer, 1979: 39; Loskop Dam, near Groblersdal, Transvaal, South Africa, ex Combretum apiculatum Sond. (Combretaceae). Redescription: Meyer (1993). Type Depository: SANC. 634. Tenuipalpus elegans (Collyer, 1973a): 529; Mangarakau, Westhaven Inlet, Nelson Province, New Zealand, ex Senecio hectori Buchanan (Asteraceae). Species designation: Colopalpus elegans—original designation, Tenuipalpus elegans—Meyer (1979). Redescription: Meyer (1979). Type Depository: Entomology Division, Department of Scientific and Industrial Research, Nelson, New Zealand. 635. Tenuipalpus elongatus Meyer, 1979: 44; Bontebok National Park, near Swellendam, Cape Province, South Africa, ex Podocarpus elongata (Aiton) L’Hér. ex Pers. (Podocarpaceae). Redescription: Meyer (1993). Type Depository: SANC. Note: host originally listed as Podocarpus elongatus. 636. Tenuipalpus emeticae Meyer, 1979: 13; Nelspruit, Transvaal, South Africa, ex Trichilia emetica Vahl (Meliaceae). Redescription: Meyer (1993). Type Depository: SANC. Synonym: 636a. Tenuipalpus citri Meyer, 1979: 25; Nelspruit, Transvaal, South Africa, ex Citrus sp. (Rutaceae)—Meyer (1993). Type Depository: SANC. 637. Tenuipalpus engelbrechti Meyer, 1979: 66; Stone Crescent Hotel, Howiesons Poort, Cape Province, South Africa, ex Helichrysum rugulosum Less. (Asteraceae). Redescription: Meyer (1993). Type Depository: SANC. 638. Tenuipalpus ephedrae Livschitz & Mitrofanov, 1970: 787; Ukraine, ex Ephedra distachya L. (Ephedraceae). Redescriptions: Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: Unknown. 639. Tenuipalpus erasus Chaudhri, 1971b: 203; Rahim Yar Khan, Punjab, Pakistan, ex Salvadora oleoides Decne. (Salvadoraceae). Redescriptions: Akbar & Chaudhri (1981), Chaudhri (1972c), Chaudhri et al. (1974), Collyer (1973b), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: Department of Entomology, West Pakistan Agricultural University Lyallpur, Pakistan. 640. Tenuipalpus eremitus Chaudhri, 1972c: 21; Pirowala Forest, Khanewal, Punjab, Pakistan, ex Atriplex crassifolia C. A. Mey. (Chenopodiaceae). Species designation: Tenuipalpus erimitus [sic]- Mitrofanov & Strunkova (1979). Redescriptions: Akbar & Chaudhri (1981), Chaudhri et al. (1974), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: Department of Entomology, West Pakistan Agricultural University Lyallpur, Pakistan. 641. Tenuipalpus eriophyoides Baker, 1948: 59; Margil, Basra, Mesopotamia, Iraq, ex an unidentified plant. Species designation: Colopalpus eriophyoides—Baker & Tuttle (1972). Redescriptions: Al-Gboory (1987), Baker & Pritchard (1953b); Baker & Tuttle (1972), Chaudhri et al. (1974), Khosrowshahi & Arbabi (1997), Meyer (1979), Mitrofanov & Strunkova (1979), Pritchard & Baker (1958). Type Depository: USNM, No. 1800. Synonym: 641a. Tenuipalpus lapidosus Siddiqui & Chaudhri, 1972: 331; Jhang, Punjab, Pakistan, ex Phoenix dactylifera L. (Arecaceae)—Chaudhri et al. (1974).

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Redescriptions: Chaudhri (1972c), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: Department of Entomology, West Pakistan Agricultural University Lyallpur, Pakistan. Note: Baker & Tuttle (1972) state that the species was described from an immature collected in “Mesopotamia”. Baker & Tuttle (1972:35) describe the adult male and female collected from date palm, Bagdad, Iraq. 642. Tenuipalpus eucleae Meyer, 1979: 41; Mkuzi Game Reserve, Natal, South Africa, ex Euclea divinorum Hiern (Ebenaceae). Redescription: Meyer (1993). Type Depository: SANC. 643. Tenuipalpus eugeniae De Leon, 1965b: 521; Green Hills, Portland, Jamaica, ex Eugenia biflora (L.) DC. (Myrtaceae). Redescriptions: Collyer (1973b), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: MCZ. 644. Tenuipalpus euonymi Khosrowshahi, 1991:14, Tehran Province, Iran, ex Euonymus sp. (Celastraceae). Resdescription: Khosrowshahi & Arbabi (1997). Type Depository: TUPH, T-122. 645. Tenuipalpus falcatus Meyer, 1979: 39; Champagne Castle Hostel, Natal, South Africa, ex Podocarpus falcatus (Thunb.) Mirb. (Podocarpaceae). Redescription: Meyer (1993). Type Depository: SANC. 646. Tenuipalpus faresianus Maninder & Ghai, 1978: 251; Mehrauli Road, New Delhi, India, ex Acacia farnesiana (L.) Willd. (Mimosaceae). Species designation: Tenuipalpus farisianus [sic]—Gupta & Chatterjee (1997). Redescriptions: Gupta & Chatterjee (1997), Sadana (1997), Sadana & Chhabra (1980), Sadana & Gupta (1984). Type Depository: INPC. 647. Tenuipalpus faveolus Meyer, 1993: 60; 15 km from George (Oudtshoorn Road), Cape Province, South Africa, ex Helichrysum petiolare Hilliard & Burtt (Asteraceae). Type Depository: SANC. 648. Tenuipalpus feliciae Meyer, 1993: 47; Karoo National Park, near Beaufort West, Cape Province, South Africa, ex Felicia hirsuta DC. (Asteraceae). Type Depository: SANC. 649. Tenuipalpus ferosus Akbar & Chaudhri, 1981: 8; 1.6 km E Muzaffarabad, (Azad Kashmir), Pakistan, ex an unidentified plant. Type Depository: Acarology Research Laboratory, Department of Entomology, University of Agriculture, Faisalabad, Pakistan. 650. Tenuipalpus fici Maninder & Ghai, 1978: 253; Patel Nagar, New Delhi, India, ex Ficus carica L. (Moraceae). Redescriptions: Gupta & Chatterjee (1997), Nassar & Ghai (1981) Sadana (1997), Sadana & Chhabra (1980), Sadana & Gupta (1984). Type Depository: INPC. Note: host originally listed as Ficus carcia. 651. Tenuipalpus filicicola Wang, 1983: 62; Sichuan, Mt. Emei, China, ex Cyclosorus sp. (Thelypteridaceae). Type Depository: Unknown. 652. Tenuipalpus flacourtiae Meyer, 1979: 23; Makoka Research Station, Malawi, ex Flacourtia indica (Burm.) Merr (Flacourtiaceae). Redescription: Meyer (1993). Type Depository: SANC.

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653. Tenuipalpus flechtmanni Mesa, Moraes & Ochoa, 2006: 48; Parque Nacional de Itatiaia, Rio de Janeiro, Brazil, ex Campomanesia sp. (Myrtaceae). Type Depository: DEES. 654. Tenuipalpus frondosus Cromroy, 1958: 84; El Yunque, Puerto Rico, ex a fern tree (Osmundaceae). Redescriptions: Collyer (1973b), De Leon (1965b), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: Collection at Department of Entomology at North Carolina Stage College, USA. 655. Tenuipalpus ghaii Mohanasundaram, 1981: 403; Coimbatore, Tamil Nadu, India, ex an unidentified plant. Redescription: Sadana (1997), Sadana & Gupta (1984). Type Depository: TNAU. 656. Tenuipalpus galpiniae Meyer, 1979: 29; Loskop Dam, near Groblersdal, Transvaal, South Africa, ex Acacia galpinii Burtt Davy (Mimosaceae). Redescription: Meyer (1993). Type Depository: SANC. 657. Tenuipalpus garciniae Meyer & Bolland, 1984: 218; Bambuluwe Lake, near Awing, Cameroon, ex Garcinia sp. (Clusiaceae). Redescription: Meyer (1993). Type Depository: SANC. 658. Tenuipalpus gatoomensis Meyer, 1979: 27; Gatooma, Zimbabwe (formerly Rhodesia), ex Combretum sp. (Combretaceae). Redescription: Meyer (1993). Type Depository: SANC. 659. Tenuipalpus geigeriae Meyer, 1979: 54; S32 on the Kruger National Park grid map, Transvaal, South Africa, ex Geigeria aspera Harv. (Asteraceae). Redescription: Meyer (1993). Type Depository: SANC. 660. Tenuipalpus granati Sayed, 1946: 100; “Lower and Upper Egypt”, Egypt ex pomegranate (Punicaceae). Species designation: Aegyptopalpus granati—Mitrofanov (1973b). Redescriptions: Akbar & Chaudhri (1981), Bagdasarian (1962), Baker & Pritchard (1953b, 1960), Channabasavanna & Lakkundi (1977), Collyer (1973b), Duzgunes (1965), Gupta & Chatterjee (1997), Hatzinikolis (1986a, 1987), Khosrowshahi & Arbabi (1997), Livschitz & Mitrofanov (1967), Maninder & Ghai (1978), Meyer (1979), Mitrofanov & Strunkova (1979), Nassar & Ghai (1981), Pritchard & Baker (1958), Reck (1951), Sadana (1997), Sadana & Chhabra (1980), Sadana & Gupta (1984), Sayed (1942), Wafa et al. (1968/1969), Wainstein (1960), Zaher et al. (1984). Type Depository: Unknown. Note: according to Wainstein (1960) Tenuipalpus orchidarum (Parfitt) Sayed (1942), is synonym of Tenuipalpus granati Sayed. Suspected senior synonym of: Tenuipalpus rodionovi—Pritchard & Baker (1958). 661. Tenuipalpus grevilleae Gutierrez & Schicha, 1982: 138; New South Wales, Australia, ex Grevillea sericea (Sm.) R. Br. (Proteaceae). Type Depository: Biological and Chemical Research Institute, New South Wales, Department of Agriculture, Rydalmere, New South Wales, Australia. 662. Tenuipalpus guamensis Baker, 1945: 36; Sumay, Guam, Tahiti, ex Neottopteris nidus (L.) J. Smith (Aspleniaceae). Redescriptions: Baker & Pritchard (1953b), Collyer (1973c), Gerson & Collyer (1984), Meyer (1979), Mitrofanov & Strunkova (1979), Pritchard & Baker (1958), Reck (1951). Type Depository: USNM, No. 1480. 663. Tenuipalpus guettardae De Leon, 1961c: 176; Key Largo, Florida, USA, ex Guettarda scabra (L.) Vent. (Rubiaceae). Species designation: Tenuipalpus guettardiae [sic]—Collyer (1973b).

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Redescriptions: Collyer (1973b), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: MCZ. 664. Tenuipalpus gumbolimbonis De Leon, 1961c: 176; Coral Glabes, Florida, USA, ex Bursera simaruba (L.) Sarg. (Burseraceae). Redescriptions: Collyer (1973b), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: MCZ. 665. Tenuipalpus guptai Sadana & Gupta, 1984: 142; Solan, India, ex Punica granatum L. (Punicaceae). Redescription: Sadana (1997). Type Depository: PAUP. Note: suspected junior synonym of the Tenuipalpus solanensis Sadana & Gupta. 666. Tenuipalpus haripuriensis Akbar & Chaudhri, 1981: 10; Haripur, Pakistan ex Cannabis sativa L (Cannabaceae). Type Depository: Acarology Research Laboratory, Department of Entomology, University of Agriculture, Faisalabad, Pakistan. 667. Tenuipalpus hastaligni De Leon, 1956a: 57; Coral Gables, Florida, USA, ex Ocotea coriacea Britton (Lauraceae). Species designation: Tenuipalpus hastalagni [sic]—Mitrofanov & Strunkova (1979). Redescriptions: Collyer (1973b), De Leon (1965b), Meyer (1979), Mitrofanov & Strunkova (1979), Pritchard & Baker (1958), Sadana (1997). Type Depository: USNM. 668. Tenuipalpus heteropyxis Meyer, 1993: 36; Blyde River Canyon, Transvaal, South Africa, ex Heteropyxis natalensis Harv. (Heteropyxidaceae). Type Depository: SANC. 669. Tenuipalpus heveae Baker, 1945: 36; Belterra, Para, Brazil, ex a rubber tree (Euphorbiaceae). Redescriptions: Baker & Pritchard (1953b), Collyer (1973b), Meyer (1979), Mitrofanov & Strunkova (1979), Pontier & Flechtmann (1999), Pritchard & Baker (1958), Reck (1951). Type Depository: USNM, No. 1481. 670. Tenuipalpus hondurensis Evans, in Evans et al. (1993): 149; El Lloron, 5 km SE Valle de Los Angeles, Francisco Morazan, Honduras, ex Pinus oocarpa Schiede & Schltdl. (Pinaceae). Type Depository: USNM. Note: host originally listed as Pinus oocarpae. 671. Tenuipalpus hornotinus Chaudhri, 1971b: 207; Abbottabad, Punjab, Pakistan, ex Punica granatum L. (Punicaceae). Redescriptions: Akbar & Chaudhri (1981), Chaudhri (1972c), Chaudhri et al. (1974), Collyer (1973b), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: Department of Entomology, West Pakistan Agricultural University Lyallpur, Pakistan. 672. Tenuipalpus hurae De Leon, 1967: 44; Asa Wright Nature Center (=Simla), Trinidad, Trinidad and Tobago, ex Hura crepitans L. (Euphorbiaceae). Redescriptions: Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: MCZ. 673. Tenuipalpus ilocanus Corpuz-Raros, 1978: 223; Kirsudan, San Isidro Lagangilang, Abra, Philippines, ex Litsea glutinosa (Lour.) C.B. Rob. (Lauraceae). Type Depository: USNM. 674. Tenuipalpus imias Cao, 1982: 14; Imias, Guantanamo, Cuba, ex Hippomane mancinella L. (Euphorbiaceae). Type Depository: Colleciòn de la Estación Biologica Docente (EBD) de la Facultad de Biologia, Universidad de la Habana. 675. Tenuipalpus indicus Maninder & Ghai, 1978: 253; Haryana, Faridabad, Punjab, India, ex Acacia farnesiana (L.) Willd. (Mimosaceae). Redescriptions: Sadana (1997), Sadana & Chhabra (1980), Sadana & Gupta (1984). Type Depository: INPC.

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676. Tenuipalpus inophylli Gutierrez & Bolland, 1981: 26; NE Koumac, New Caledonia, ex Codiaeum inophyllum Mull. Arg. (Euphorbiaceae). Type Depository: Museum National d’Historie Naturelle de Paris, France. 677. Tenuipalpus insularis Meyer, 1993: 17; Die Eiland, near Tzaneen, Transvaal, South Africa, ex an unidentified wild tree. Type Depository: SANC. 678. Tenuipalpus isabelae Mesa, Moraes & Ochoa, 2006: 45; Reserva Ecologica de Ibicatu, Piracicaba—SP., Brazil, ex Actinostemon sp. (Euphorbiaceae). Type Depository: DEES. 679. Tenuipalpus ixorae Maninder & Ghai, 1978: 254; Haldwani, Uttar Pradesh, India; ex Ixora parviflora Vahl (Rubiaceae). Redescriptions: Sadana (1997), Sadana & Chhabra (1980), Sadana & Gupta (1984). Type Depository: INPC. 680. Tenuipalpus jagatkhanaens Sadana & Gupta, 1984: 144, Jaget Khana, Nalagarh, India, ex Achras sapota L. (Sapotaceae). Species designation: Tenuipalpus jagatkhanaensis—Sadana (1997). Redescription: Sadana (1997), Sadana & Gupta (1984). Type Depository: PAUP. Note: Species description based on male, female unknown 681. Tenuipalpus jamaicensis De Leon, 1965b: 521; between Lluidas Vale and Top Hill, Saint Catherine, Jamaica, ex Tetrazygia pallens Cogn. (Melastomataceae). Redescriptions: Collyer (1973b), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: MCZ. 682. Tenuipalpus japonicus Nishio, 1956: 80; Okayama, Japan, ex Diospyros kaki L. var domestica Makino (Ebenaceae). Redescriptions: Collyer (1973b), Ehara (1956), Meyer (1979, 1993). Type Depository: Hokkaido National Agric. Expt. Stat. Japan. Note: according to Mitrofanov & Strunkova, 1979, Tenuipalpus japonicus Nishio, 1956 is junior synonym to Tenuipalpus zhizhilashviliae Reck, 1953 683. Tenuipalpus jasmini Khan, 1970: 292; Lawrence Road, Lahore, Pakistan, ex Jasminum sp. (Oleaceae). Type Depository: Unknown. 684. Tenuipalpus jawadii Hasan, Wakil & Bashir, 2004: 98; Sadiqabad, Pakistan, ex Salvadora sp. (Salvadoraceae). Type Depository: Acarology Research Laboratory, Department of Agric. Entomology, University of Agriculture, Faisalabad, Pakistan. 685. Tenuipalpus jianfengensis Ma & Yuan, 1980: 119; Jianfeng, Hainan Island, Guangdong, China, ex an unidentified host. Type Depository: CMNS. 686. Tenuipalpus jonkeri Meyer, 1993: 61; Jonkershoek, near Stellenbosch, Cape Province, South Africa, ex Salvia africana-caerulea L. (Lamiaceae). Type Depository: SANC. 687. Tenuipalpus jordaani Meyer, 1979: 63; Pafuri, Kruger National Park, Transvaal, South Africa, ex unidentified wild plant. Redescription: Meyer (1993). Type Depository: SANC. 688. Tenuipalpus jussiaeae De Leon, 1961c: 175; Everglades National Park, Florida, USA, ex Jussiaea sp. (Onagraceae). Species designation: Tenuipalpus jussiae [sic]—Collyer (1973b). Redescriptions: Collyer (1973b), Meyer (1979), Mitrofanov & Strunkova (1979).

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Type Depository: MCZ. Note: host originally listed as Jussiaeae sp. 689. Tenuipalpus kamalii Khosrowshahi & Arbabi, 1997: 11; Tehran province, Iran, ex Fraxinus excelsior L. (Oleaceae). Type Depository: TUPH, No. T-1422. 690. Tenuipalpus kapoki De Leon, 1957: 89; Tuxtla Gutierrez, Chiapas, Mexico, ex Ceiba pentandra (L.) Gaertn. (Bombacaceae). Redescriptions: Baker & Tuttle (1987), Collyer (1973b), De Leon (1965b), Evans et al. (1993), Gerson & Collyer (1984), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: MCZ. 691. Tenuipalpus karrooi Meyer, 1979: 51; Pretoria, Transvaal, South Africa, ex Acacia karroo Hayne (Mimosaceae). Redescription: Meyer (1993). Type Depository: SANC. 692. Tenuipalpus keiensis Meyer, 1979: 63; Great Kei River Pass, Cape Province, South Africa, ex Hypoestes sp. (Acanthaceae). Redescription: Meyer (1993). Type Depository: SANC. 693. Tenuipalpus kenos Hasan, Wakil & Bashir, 2004: 41; Daddar, Chakwal, Pakistan, ex Prunus persica (L.) Batsch. (Rosaceae). Type Depository: Acarology Research Laboratory, Department of Agric. Entomology, University of Agriculture, Faisalabad, Pakistan. Note: suspected junior synonym of Tenuipalpus mandraensis Hasan, Wakil & Bashir. 694. Tenuipalpus kesari Sadana, Gupta & Goyal, 1985: 162; Malodh, Ludhiana, India, ex Brassica oleracea L. Alef. var. botrytis L. (Brassicaceae). Redescriptions: Sadana (1985, 1997). Type Depository: PAUP. 695. Tenuipalpus knorri Baker & Pritchard, 1953b: 320; Concordia, Argentina, ex Salix alba L. (Salicaceae). Redescriptions: Collyer (1973b), Meyer (1979), Mitrofanov & Strunkova (1979), Pritchard & Baker (1958). Type Depository: USNM, No. 2000. 696. Tenuipalpus kobachidzei Reck, 1951: 295; Georgia, ex Clematis vitalba L. (Ranunculaceae). Redescriptions: Collyer (1973b), Livschitz & Mitrofanov (1967), Meyer (1979), Mitrofanov & Strunkova (1979), Pritchard & Baker (1958). Type Depository: Unknown. 697. Tenuipalpus kraussianae Meyer, 1993: 39; Hans Merensky Nature Reserve, Die Eiland, near Tzaneen, Transvaal, South Africa, ex Acacia nilotica (L.) Willd. & Delile subsp. kraussiana (Benth.) Brenan (Mimosaceae). Type Depository: SANC. 698. Tenuipalpus lalbaghensis Channabasavanna & Lakkundi, 1977: 19; Bangalore, Lalbagh, Karnataka, India, ex Artocarpus integrifolia L. (Moraceae). Redescriptions: Maninder & Ghai (1978), Meyer (1979), Nassar & Ghai (1981), Sadana (1997), Sadana & Chhabra (1980), Sadana & Gupta (1984). Type Depository: Zoological Survey of India and Department of Entomology, University of Agricultural Sciences, Bangalore, India. 699. Tenuipalpus laminasetae Mohanasundaram, 1981: 401; Coimbatore, Tamil Nadu, India, ex an unidentified plant. Redescription: Sadana (1997), Sadana & Gupta (1984). Type Depository: TNAU. 700. Tenuipalpus lanceae Meyer, 1979: 23; Roodeplaat, near Pretoria, Transvaal, South Africa, ex Rhus lancea L. (Anacardiaceae).

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Redescription: Meyer (1993). Type Depository: SANC. 701. Tenuipalpus latiseta Aranda, in: Flechtmann, 1976: 61; Corumbatai, São Paulo, Brazil, ex (Celastraceae). Type Depository: DEES. 702. Tenuipalpus lawrencei Baker & Pritchard, 1960: 566; Leopoldville (formerly Kinshasa), Democratic Republic of Congo (formerly Zaïre), ex an unidentified large tree. Redescriptions: Collyer (1973b), Meyer (1979, 1993), Mitrofanov & Strunkova (1979). Type Depository: USNM, No. 2504. 703. Tenuipalpus legatus Chaudhri, 1972c: 23; 14 km N of Karachi, Sind, Pakistan, ex Alhagi maurorum Medik. (Fabaceae). Redescriptions: Akbar & Chaudhri (1981), Chaudhri et al. (1974), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: Department of Entomology, West Pakistan Agricultural University Lyallpur, Pakistan. Note: host originally listed as Alhagi camelorum 704. Tenuipalpus leipoldti Meyer, 1993: 62; Pakhuis Pass, Christian Louis Leipoldt´s grave, Cape Province, South Africa, ex Rhus undulata Jacq. (Anacardiaceae). Type Depository: SANC. 705. Tenuipalpus leonorae Meyer, 1979: 15; Adelaide, Cape Province, South Africa, ex Acacia karroo Hayne (Mimosaceae). Redescription: Meyer (1993). Type Depository: SANC. 706. Tenuipalpus leucospermi Meyer, 1993: 64; 30 km from Willowmore (Baviaanskloof Road), Cape Province, South Africa, ex Leucospermum conocarpodendron Buek (Proteaceae). Type Depository: SANC. 707. Tenuipalpus lineosetosus Wang, 1983: 61; Qiemo, Xinjiang, China, ex poplar (Populus sp.—Salicaceae). Type Depository: Institute of Zoology, Academia Sinica, China. 708. Tenuipalpus lucumae De Leon, 1957: 84; Tuxtla Gutierrez, Chiapas, Mexico, ex Lucuma salicifolia H.B. & K. (Sapotaceae). Redescriptions: Baker & Tuttle (1987), Collyer (1973b), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: MCZ. 709. Tenuipalpus ludhianaensis Sadana & Chhabra, 1980: 152; Ludhiana, Punjab, India, ex Pyrus communis L. (Rosaceae). Redescriptions: Sadana (1997), Sadana & Gupta (1984). Type Depository: PAUP. 710. Tenuipalpus lulinicus Ma & Yuan, 1980: 118; Haikou, Hainan Island, Guangdong, China, ex Ruellia drymophila Hand.-Mazz. (Acanthaceae). Type Depository: CMNS. 711. Tenuipalpus lunatus Meyer, 1979: 25; Komatipoort, Transvaal, South Africa, ex Acacia nigrescens Oliver (Mimosaceae). Redescription: Meyer (1993). Type Depository: SANC. 712 Tenuipalpus lustrabilis Chaudhri, 1971b: 211; Muzaffargarh, Punjab, Pakistan, ex Punica granatum L. (Punicaceae). Redescriptions: Chaudhri (1972c), Chaudhri et al. (1974), Collyer (1973b). Type Depository: Department of Entomology, West Pakistan Agricultural University Lyallpur, Pakistan. Note: according to Collyer (1973b), from the description of Tenuipalpus lustrabilis Chaudhri, 1971 it is not possible to separate this species from Tenuipalpus punicae Pritchard & Baker, 1958; Meyer (1979) suspected Tenuipalpus lustrabilis Chaudhri, to be a junior synonym of Tenuipalpus punicae Pritchard & Baker.

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713. Tenuipalpus lycioides Meyer, 1979: 15; Gatooma Research Station, Zimbabwe (formerly Rhodesia), ex Diospyros lycioides Desf. (Ebenaceae). Redescription: Meyer (1993). Type Depository: SANC. 714. Tenuipalpus lygodii De Leon, 1965b: 519; Ipswich, Saint Elizabeth, Jamaica, ex Lygodium volubile Sw. (Lygodiaceae). Redescriptions: Al-Gboory (1987), Collyer (1973b), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: MCZ. 715. Tenuipalpus magalismontani Meyer, 1993: 21; Venda, South Africa, ex Bequaertiodendron magalismontanum (Sond.) Heine & J. H. Hemsl. (Sapotaceae). Type Depository: SANC. 716. Tenuipalpus mahoensis Collyer, 1964: 438; Waitakeres, near Auckland, North Island, New Zealand, ex Melicytus ramiflorus J. R. Forst. & G. Forst. (Violaceae). Redescriptions: Collyer (1973b), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: BMNH. 717. Tenuipalpus malligai Mohanasundaram, 1981: 399; Coimbatore, Tamil Nadu, India, ex an unidentified plant. Redescription: Sadana (1997), Sadana & Gupta (1984). Type Depository: TNAU. 718. Tenuipalpus mallotae Mohanasundaram, 1983b: 165; Vridhachalam, Tamil Nadu, India, ex Mallotus sp. (Euphorbiaceae). Redescription: Sadana (1997). Type Depository: TNAU. 719. Tenuipalpus mandraensis Hasan, Wakil & Bashir, 2004: 41; 2 km SW Mandra, Pakistan, ex an unidentified plant. Type Depository: Acarology Research Laboratory, Department of Agric. Entomology, University of Agriculture, Faisalabad, Pakistan. Note: suspected junior synonym of Tenuipalpus kenos, Hasan, Wakil & Bashir. 720. Tenuipalpus mansoni De Leon, 1965b: 521; Palisadoes Park, Kingston, Jamaica, ex Tabebuia sp. (Bignoniaceae). Redescriptions: Collyer (1973b), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: MCZ. 721. Tenuipalpus mansoniculus Ghai & Shenhmar, 1985: 117; replacement to Colopalpus mansoni Collyer (1973b), name pre-occuped by Tenuipalpus mansoni De Leon (1965b). (Article 58.7 4th Edition International Code of Zoological Nomenclature). Redescriptions: Ghai & Shenhmar (1984), Meyer (1979), Smiley & Gerson (1995). Type Depository: Collection of Entomology, Division, Department of Sciences and Industrial Research, Division, Nelson, New Zealand. Synonym: 721a. Colopalpus mansoni Collyer, 1973a: 531; intercepted in quarantine from Australia ex Pomaderris sp. (Rhamnaceae)—Ghai & Shenhmar (1984). Species designation: Tenuipalpus mansoni—Meyer (1979). Redescriptions: Ghai & Shenhmar (1985), Meyer (1979), Smiley & Gerson (1995). Type Depository: Collection of Entomology, Division, Department of Sciences and Industrial Research, Division, Nelson, New Zealand. 722. Tenuipalpus matthyssei (Pritchard & Baker, 1958): 259; Laguna, Philippines, ex Melanolepis multiglandulosa (Reinw. ex Blume) Rchb.f. & Zoll. (Euphorbiaceae). Species designations: Colopalpus matthyssei—original designation; Tenuipalpus mattyssei—Meyer (1979). Redescriptions: Meyer (1979), Mitrofanov & Strunkova (1979), Rimando (1962). Type Depository: USNM, No. 2343. Note: host originally listed as Melanolipes multiglandulosa.

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723. Tenuipalpus melhaniae Meyer, 1979: 11; Tshokwane, Kruger National Park, Transvaal, South Africa, ex Melhania forbesii Planch. & Mast. (Sterculiaceae) Redescription: Meyer (1993). Type Depository: SANC. Note: host originally listed as Melhania forhesii. 724 Tenuipalpus menglunensis Yin & Cui, 1996: 73; Menglun, Yuman Province, China, ex a kind of tree. Type Depository: Department of Plant Protection, Shenyang Agricultural University, China. 725. Tenuipalpus metis Hasan, Akbar & Bashir, 2003: 205, 3 km E Muree, Pakistan, ex Prunus amygdalus Stokes (Rosaceae). Type Depository: Acarology Research Laboratory, Department of Entomology, University of Agriculture, Faisalabad, Pakistan. Note: Suspected junior synonym of Tenuipalpus haripuriensis Akbar & Chaudhri. 726. Tenuipalpus metopii De Leon, 1956a: 56; Coral Gables, Florida, USA, ex Metopium toxiferum (L.) Krug & Urban (Anacardiaceae). Redescriptions: Collyer (1973b), Meyer (1979), Mitrofanov & Strunkova (1979), Pritchard & Baker (1958). Type Depository: USNM. 727. Tenuipalpus micheli Lawrence, 1940: 111; Durban, Umhloti Beach, Natal, South Africa, ex Chaetachme aristata (Ulmaceae). Redescriptions: Baker (1945), Baker & Pritchard (1953b, 1960), Chandra & Channabasavanna (1974), Channabasavanna & Lakkundi (1977), Collyer (1973b), Lawrence (1943), Maninder & Ghai (1978), Meyer (1979, 1993), Meyer & Ryke (1959), Mitrofanov & Strunkova (1979), Pritchard & Baker (1958), Reck (1951), Sadana (1997), Sadana & Chhabra (1980), Sadana & Gupta (1984). Type Depository: SANC. Note: host originally listed as Chaetaeme. 728. Tenuipalpus microphylli Meyer, 1979: 48; Limpopo River, Citrus Research Station, near Messina, Transvaal, South Africa, ex Combretum paniculatum Vent. subsp. microphyllum (Klotzsch) Wickens (Combretaceae). Redescription: Meyer (1993). Type Depository: SANC. 729. Tenuipalpus mkuziensis Meyer, 1979: 51; Mkuzi Game Reserve, Natal, South Africa, ex Pteleopsis myrtifolia Engl. & Diels. (Combretaceae). Redescription: Meyer (1993). Type Depository: SANC. 730. Tenuipalpus molinai Evans, in Evans et al. (1993): 149; Lago Yojoa, Cortes, Honduras, ex Asteraceae. Type Depository: USNM. 731. Tenuipalpus montanus Collyer, 1973c: 949; Mount Grey, 1.1 km Turret Range, Lake Manapouri area, New Zealand, ex Pimelea sericeovillosa Hook. (Thymelaeaceae), with litter and soil from beneath this shrub. Species designation: Tuttlepalpus montanus—Mitrofanov & Strunkova (1979). Redescriptions: Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: Entomology Division, Department of Scientific and Industrial Research, Auckland, New Zealand. 732. Tenuipalpus mopaneae Meyer, 1979: 21; Beit Bridge, near Messina, Transvaal, South Africa, ex Colophospermum mopane J. Kirk & Benth. (Fabaceae). Redescription: Meyer (1993). Type Depository: SANC. 733. Tenuipalpus moraesi Feres & Hernandez, 2006:60, Estaçao Ecológica de Paulo de Faria, Paulo de Faria, Sao Paulo, Brazil, ex Sebastiana brasiliensis Spreng (Euphorbiaceae). Type Depository: Collection of Acari Departamento de Zoologia e Botanica, UNESP, Sao Jose do Rio Preto, State of Sao Paulo, Brazil.

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734. Tenuipalpus morianus Meyer, 1993: 23; Lebowa, Transvaal, South Africa, ex Combretum molle R. Br. & G. Don (Combretaceae). Type Depository: SANC. 735. Tenuipalpus mourerae De Leon, 1965a: 71; Bartica Nature Reserve, Guyana (= British Guiana), ex Mourera sideroxylon. (Podostemaceae) Redescriptions: Collyer (1973b), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: MCZ. Note: the host listed in original description could not be verified, possibly ex Mouriri sideroxylon Sagot & Triana (Melastomataceae). 736. Tenuipalpus muguanicus Ma & Yuan, 1980: 120; Qufu, Shandong, China, ex Carica papaya L. (Caricaceae). Type Depository: CMNS. 737. Tenuipalpus mustus Chaudhri, 1972c: 25; Pirowala Forest area, Pakistan, ex Bauhinia purpurea L. (Caesalpiniaceae). Redescriptions: Akbar & Chaudhri (1981), Chaudhri et al. (1974), Meyer (1979), Mitrofanov & Strunkova (1979), Sadana (1997), Sadana & Chhabra (1980), Sadana & Gupta (1984). Type Depository: Department of Entomology, West Pakistan Agricultural University Lyallpur, Pakistan. 738. Tenuipalpus myrtus Al-Gboory, 1987: 135; Abu-Ghraib, Baghdad, Iraq, ex Myrtus communis L. (Myrtaceae). Type Depository: Entomology Museum, College of Agriculture, Department of Plant Protection, Abu-Ghraib, Baghdad, Iraq. 739. Tenuipalpus namaensis Meyer, 1993: 25; De Hoek Farm, Vanrhynsdorp, Cape Province, South Africa, ex Rhus lancea L. (Anacardiaceae). Type Depository: SANC. 740. Tenuipalpus nenaxi Meyer, 1993: 76; Karoo National Park, near Beaufort West, Cape Province, South Africa, ex Nenax microphylla (Sond.) Salter (Rubiaceae). Type Depository: SANC. 741. Tenuipalpus niekerkae Meyer, 1979: 70; Charter's Creek, Saint Lucia, Natal, South Africa, ex Apodytes dimidiata E. Mey. & Arn. (Icacinaceae). Redescriptions: Meyer (1993), Ueckermann & Meyer (1988). Type Depository: SANC. 742. Tenuipalpus nigerianus Meyer, 1979: 56; Ibadan, Nigeria, ex Aspilia africana (Pers.) C.D. Adams (Asteraceae). Redescription: Meyer (1993). Type Depository: SANC. 743. Tenuipalpus obvelatus Wang, 1983: 61; Mount Badaling, Beijing, China, ex Cyperaceae. Type Depository: Institute of Zoology, Academia Sinica, China. 744. Tenuipalpus oliveirai Flechtmann, 1994: 184; Barretos, São Paulo, Brazil, ex Peschiera fuchsiaefolia Miers (Apocynaceae). Type Depository: Collection of the Department of Zoology, ESALQ, University of Sao Paulo, Brazil. Note: host originally listed as Peschiera fuchsiaefoliae. 745. Tenuipalpus ombrensis Meyer, 1993: 41; Farm Belle Ombre (road between Tzaneen and Hans Merensky Nature Reserve), Transvaal, South Africa, ex Acacia gerrardii Benth. (Mimosaceae). Type Depository: SANC. 746. Tenuipalpus orchidofilo Moraes & Freire, 2001: 2; Piracicaba, São Paulo, Brazil, ex Arundina graminifolia Hochr. (Orchidaceae). Type Depository: DEES. 747. Tenuipalpus oribiensis Meyer, 1979: 61; Oribi Gorge, Natal, South Africa, ex Loxostylis alata Spreng. & Rchb. (Anacardiaceae).

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Redescription: Meyer (1993). Type Depository: SANC. 748. Tenuipalpus orilloi Rimando, 1962: 42; Lipa, Batangas, Philippines, ex citrus (Rutaceae). Redescriptions: Collyer (1973b), De Leon (1965a, 1967), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: Department of Entomology, University of the Philippines, Philippines. Synonym: 748a. Tenuipalpus spathiphyllus Manson, 1963: 217; Indonesia, ex Spathiphyllum sp. (Araceae).—De Leon (1967). Type Depository: USNM, No. 2887. 749. Tenuipalpus ortus Chaudhri, 1972c: 22; Lyallpur, Multan, Pakistan, ex Morus alba L. (Moraceae). Redescriptions: Akbar & Chaudhri (1981), Chaudhri et al. (1974), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: Department of Entomology, West Pakistan Agricultural University Lyallpur, Pakistan. 750. Tenuipalpus ovalis Meyer & Ryke, 1959: 322; Kaap Muiden, Transvaal, South Africa, ex an unidentified wild shrub. Redescriptions: Meyer (1979, 1993), Mitrofanov & Strunkova (1979). Type Depository: Collection of the Institute for Zoological Research of the Potchefstroom University for C.H.E., Potchefstroom, South Africa. 751. Tenuipalpus oxalis (Flechtmann), in: Flechtmann 1976: 63; Piracicaba, São Paulo, Brazil, ex Oxalis sp. (Oxalidaceae). Species designation: Colopalpus oxalis—original designation, Tenuipalpus oxalis—Meyer (1979). Redescription: Meyer (1979). Type Depository: DEES. 752. Tenuipalpus pacificus Baker, 1945: 38; Panama Canal Zone, Panama intercepted at Hoboken, New Jersey, USA, ex Phalaenopsis stuartiana Rchb. (Orchidaceae). Redescriptions: Baker & Pritchard (1953b), Charanasri et al. (1986), Collyer (1973b), Ghai & Shenhmar (1984), Hatzinikolis (1986a, 1987), Meyer (1979, 1993), Mitrofanov & Strunkova (1979), Pritchard & Baker (1952, 1958), Rimando (1962), Smiley & Gerson (1995). Type Depository: USNM, No. 1482. 753. Tenuipalpus pagesae Rimando, 1962: 43; Guinobatan, Albay, Philippines, ex Lagerstroemia speciosa (L.) Pers. (Lythraceae). Redescriptions: Collyer (1973b), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: Department of Entomology, University of the Philippines, Philippines. 754. Tenuipalpus pagina Chaudhri, Akbar & Rasool, 1974: 75; 16 km E Karachi, Sind, Pakistan, ex Fagonia cretica L. (Zygophyllaceae). Redescription: Akbar & Chaudhri (1981). Type Depository: Department of Entomology, University of Agriculture, Lyallpur, Pakistan. 755. Tenuipalpus palosapis Corpuz-Raros, 1978: 227; Makiling Botanic Gardens, Mount Makiling, Laguna, Philippines, ex Anisoptera thurifera Blume (Dipterocarpaceae). Type Depository: USNM. 756. Tenuipalpus panici De Leon, 1965c: 196; south of Baranquetas, Puerto Rico, ex Panicum maximum Jacq. (Poaceae). Redescriptions: Collyer (1973b), De Leon (1967), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: MCZ. 757. Tenuipalpus papiothalensis Meyer, 1993: 67; Baviaanskloof, Cape Province, South Africa, ex Rhus macowanii Schonl. (Anacardiaceae). Type Depository: SANC. 758. Tenuipalpus pareriophyiodes Meyer & Gerson, 1980: 71; En Yahav, Israel, ex Phoenix dactylifera L. (Arecaceae). Redescription: Gerson et al. (1983). Type Depository: Collection of the Department of Entomology, Faculty of Agriculture, Rehovot, Israel.

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759. Tenuipalpus parsii Khosrowshahi & Arbabi, 1997: 13; Tehran province, Iran, ex Plantago lanceolata L. (Plantaginaceae). Type Depository: TUPH, No. T-1522. 760. Tenuipalpus pedrus (Manson, 1963): 222; Manacai, San Pedro, Honduras, ex an unidentified tree. Species designation: Colopalpus pedrus—original designation, Tenuipalpus pedrus—Meyer (1979). Redescriptions: Evans et al. (1993), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: USNM, No. 2891. 761. Tenuipalpus pernicis Chaudhri, Akbar & Rasool, 1974: 85; 3.2 km E Campbellpur, Rawalpindi, Punjab, Pakistan, ex Chrysanthemum sp. (Asteraceae). Redescriptions: Akbar & Chaudhri (1981), Gupta & Chatterjee (1997), Maninder & Ghai (1978), Sadana (1997), Sadana & Chhabra (1980). Type Depository: Department of Entomology, University of Agriculture, Lyallpur, Pakistan. 762. Tenuipalpus persicae Sadana, Chhabra & Gupta, 1982: 399, Punjab Agriculture University Ludhiana, Punjab, India, ex Prunus persica (L.) Batsch (Rosaceae). Redescriptions: Sadana (1985, 1997). Type Depository: PAUP. 763. Tenuipalpus philippinensis (Corpuz-Raros, 1978): 216; Nasipit Lumber Company, Tungao Butuan, Agusan, Philippines, ex Pterocarpus indicus Willd. (Fabaceae). Species designation: Colopalpus philippinensis—original designation, Tenuipalpus phillippinensis—Meyer (1979). Redescription: Meyer (1979). Type Depository: Unknown. 764. Tenuipalpus pieteri Meyer, 1979: 32; 15 km S Potgietersrus, Transvaal, South Africa, ex Terminalia sericea Burch. & DC. (Combretaceae). Redescription: Meyer (1993). Type Depository: SANC. 765. Tenuipalpus pigrus Pritchard & Baker, 1952: 43; Mount Diablo, California, USA, ex Umbellularia californica (Hook. & Arn.) Nutt. (Lauraceae). Redescriptions: Baker & Pritchard (1953b), Collyer (1973b), Meyer (1979), Mitrofanov & Strunkova (1979), Pritchard & Baker (1958). Type Depository: USNM, No. 1911. 766. Tenuipalpus pisinnus Chaudhri, Akbar & Rasool, 1974: 68; 4.8 km W Campbellpur, Punjab, Pakistan, ex Heliotropium strigosum Willd. (Boraginaceae). Redescription: Akbar & Chaudhri (1981). Type Depository: Department of Entomology, University of Agriculture, Lyallpur, Pakistan. 767. Tenuipalpus placitus Chaudhri, 1971b: 209; Rahim Yar Khan, Punjab, Pakistan, ex Punica granatum L. (Punicaceae). Species designation: Tenuipalpus placitis [sic]—Chaudhri (1972c). Redescriptions: Akbar & Chaudhri (1981), Chaudhri (1972c), Chaudhri et al. (1974), Collyer (1973b), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: Department of Entomology, West Pakistan Agricultural University Lyallpur, Pakistan. 768. Tenuipalpus platycaryae Wang, 1983: 62; Zhejiang, Hangzhou, China, ex Platycarpa strobilacea Sieb. & Zucc. (Juglandaceae). Type Depository: Institute of Zoology, Academia Sinica, China. 769. Tenuipalpus podocarpi Lawrence, 1943: 40; Cathkin Peak, Drakensberg Mountains, South Africa, ex Podocarpus falcatus (Thunb.) R.Br. ex Mirb. (Podocarpaceae). Redescriptions: Baker & Pritchard (1953b, 1960), Collyer (1973b), Meyer & Ryke (1959), Meyer (1979, 1993), Mitrofanov et al (1975), Mitrofanov & Strunkova (1979), Pritchard & Baker (1958). Type Depository: SANC.

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770. Tenuipalpus populi Al-Gboory, 1987; Baghdad, Iraq, ex Populus nigra L. (Salicaceae). Type Depository: Entomology Museum, College of Agriculture, Department of Plant Protection, Abu-Ghraib, Baghdad, Iraq. Note: Meyer (1979: 11) states the types slides have the following collection data: Champagne Castle Hostel, Natal, from Podocarpus falcatus, May 1941. 771. Tenuipalpus portulacae Parsi, Khosrowshahi & Farid, 1990: 39, Kerman province (Jiroft), Iran, ex Portulaca oleracea L. (Portulacaceae). Redesciption: Khosrowshahi & Arbabi (1997). Type Depository: TUPH, No. 703. 772. Tenuipalpus proctori De Leon, 1965b: 521; Ipswich, Saint Elizabeth, Jamaica, ex Hohenbergia proctori L.B. Sm. (Bromeliaceae). Redescriptions: Collyer (1973b), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: MCZ. 773. Tenuipalpus proteae Meyer, 1979: 59; Kirstenbosch Botanical Garden, Cape Province, South Africa, ex Protea nana Thunb. (Proteaceae). Redescription: Meyer (1993). Type Depository: SANC. 774. Tenuipalpus protectus Meyer, 1979: 32; Mkuze Game Reserve, Natal, South Africa, ex an unidentified Caesalpiniaceae. Redescription: Meyer (1993). Type Depository: SANC. 775. Tenuipalpus protumidus Meyer, 1993: 69; Boulder Kloff, W Buffelspoort Dam, Transvaal, South Africa, ex Berkheya carlinopsis Welw. & O Hoffm. (Asteraceae). Type Depository: SANC. 776. Tenuipalpus pruni Maninder & Ghai, 1978: 255; Himachal Pradesh, India; ex Prunus persica (L.) Batsch (Rosaceae). Redescriptions: Nassar & Ghai (1981), Sadana (1997), Sadana & Chhabra (1980), Sadana & Gupta (1984). Type Depository: INPC. 777. Tenuipalpus prunioides Meyer, 1979: 34; N151 on Kruger National Park, grid map, Transvaal, South Africa, ex Terminalia prunioides M. Laws. (Combretaceae). Redescription: Meyer (1993). Type Depository: SANC. 778. Tenuipalpus pseudocedrelae Livschitz in Mitrofanov & Strunkova, 1979: 56; replacement to Tenuipalpus cedrelae Livschitz in Livschitz & Salinas (1968), name pre-occuped by Tenuipalpus cedrelae De Leon (1957). (Article 58.7 4th Edition International Code of Zoological Nomenclature). Type Depository: Unknown. Synonym: 778a. Tenuipalpus cedrelae Livschitz in Livschitz & Salinas, 1968: 120; Santiago de las Vegas, Cuba, ex Cedrela mexicana M.Roem. (Meliaceae)—Mitrofanov & Strunkova (1979). Type Depository: Unknown. 779. Tenuipalpus punicae Pritchard & Baker, 1958: 240; Bet Maskita, Palestine, ex Punica granatum L. (Punicaceae). Suspected senior synonym of: Tenuipalpus lustrabilis Chaudhri, 1971b—Meyer (1979) Species designation: Tenuipalpus punciae [sic]—Sadana & Chhabra (1980). Redescriptions: Akbar & Chaudhri (1981), Al-Gboory (1987), Bagdasarian (1962), Channabasavanna & Lakkundi (1977), Chaudhri (1972c), Chaudhri et al. (1974), Collyer (1973b), Gupta (1995), Gupta & Chatterjee (1997), Hatzinikolis (1986a, 1987), Khosrowshahi & Arbabi (1997), Livschitz & Mitrofanov (1967), Maninder & Ghai (1978), Meyer (1979), Mitrofanov & Strunkova (1979), Nassar & Ghai (1981), Sadana (1997), Sadana & Chhabra (1980), Sadana & Gupta (1984), Strunkova (1978), Wafa et al. (1968/1969), Wainstein (1960), Zaher et al. (1984). Type Depository: USNM, No. 2206.

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780. Tenuipalpus punjabensis (Maninder & Ghai, 1978): 111; Phagwara, Punjab, India, ex Nerium indicum Mill. (Apocynaceae). Species designation: Colopalpus punjabensis—original designation. Redescriptions: Gupta (1995), Meyer (1979), Sadana (1997), Sadana & Gupta (1984). Type Depository: INPC. 781. Tenuipalpus pyroides Meyer, 1979: 54; Loskop Dam, near Groblersdal, Transvaal, South Africa, ex Rhus pyroides Burch. (Anacardiaceae). Redescription: Meyer (1993). Type Depository: SANC. 782. Tenuipalpus pyrusae Maninder & Ghai, 1978: 257; Indian Agricultural Research Institute, New Delhi, India, ex Pyrus malus L. (Rosaceae). Redescriptions: Gupta & Chatterjee (1997), Nassar & Ghai (1981), Sadana (1997), Sadana & Chhabra (1980), Sadana & Gupta (1984). Type Depository: INPC. Note: suspected junior synonym of Tenuipalpus dimensus Chaudhri. 783. Tenuipalpus qingchengensis Wang, 1983: 62; Mount Qingcheng, Sichuan, China, ex Viburnum brachybotryum Hemsl., Forb. & Hemsl. (Caprifoliaceae). Type Depository: Institute of Zoology, Academia Sinica, China. 784. Tenuipalpus rangiorae Collyer, 1964: 436; Piha, near Auckland, North Island, New Zealand, ex Brachyglottis repanda J.R. Forster & G. Forster (Asteraceae). Redescriptions: Collyer (1973b), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: BMNH. 785. Tenuipalpus raphiae Meyer & Bolland, 1984: 219; Bamenda, Cameroon, ex Raphia sp. (Arecaceae). Redescription: Meyer (1993). Type Depository: SANC. 786. Tenuipalpus raptor Akbar & Chaudhri, 1981: 6; 4.8 km E Faisalabad, Pakistan, ex an unidentified plant. Type Depository: Acarology Research Laboratory, Department of Entomology, University of Agriculture, Faisalabad, Pakistan. Note: suspected junior synonym of Tenuipalpus dimensus Chaudhri. 787. Tenuipalpus rarus Chaudhri, 1972c: 20; Gatwala Forest Park, Punjab, Pakistan, ex Peganum harmala L. (Zygophyllaceae). Redescriptions: Akbar & Chaudhri (1981), Chaudhri et al. (1974), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: Department of Entomology, West Pakistan Agricultural University Lyallpur, Pakistan. Note: host originally listed as Peganum hermala. 788. Tenuipalpus reticulus Siddiqui & Chaudhri, 1972c: 329; West Pakistan Agricultural University, Lyallpur Pakistan, ex Vitis vinifera L. (Vitaceae). Redescriptions: Akbar & Chaudhri (1981), Chaudhri (1972c), Chaudhri et al. (1974), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: Department of Entomology, West Pakistan Agricultural University Lyallpur, Pakistan. 789. Tenuipalpus rhagicus Pritchard & Baker, 1952: 42; Crescent Lake, California, USA, ex Vaccinium ovatum Pursh (Ericaceae). Redescriptions: Baker & Pritchard (1953b), Collyer (1973b), Meyer (1979), Mitrofanov & Strunkova (1979), Pritchard & Baker (1958). Type Depository: USNM, No. 1910. 790. Tenuipalpus rhizophorae De Leon, 1967: 41; Coroni Swamp, near Experiment Station Farm, Trinidad, Trinidad and Tobago, ex Rhizophora mangle L. (Rhizophoraceae). Redescription: Meyer (1979). Type Depository: MCZ.

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791. Tenuipalpus rhusi Meyer, 1979: 59; Bontebok National Park, near Swellendam, Cape Province, South Africa, ex Rhus undulata Jacq. (Anacardiaceae). Redescription: Meyer (1993). Type Depository: SANC. 792. Tenuipalpus rhysus Baker & Pritchard, 1953b: 330; Glen Saint Mary, Florida, USA, ex Cyrilla racemiflora L. (Cyrillaceae). Species designation: Tenuipalpus rhyssus [sic]—De Leon (1957). Redescriptions: Baker & Tuttle (1987), Collyer (1973b), De Leon (1957), Mairena & Ochoa (1989), Meyer (1979), Mitrofanov & Strunkova (1979), Pritchard & Baker (1958), Salas & Ochoa (1986). Type Depository: USNM, No. 2003. Note: Evans et al. (1993) synomyzed T. rhysus with T. bakeri McGregor (1949), but examination of new material led us to conclude that they are not synonyms. 793. Tenuipalpus robustae Meyer, in Ueckermann & Meyer (1988): 36; Addo Elephant National Park, Cape Province, South Africa, ex Grewia robusta Burch. (Tiliaceae). Redescription: Meyer (1993). Type Depository: Unknown. 794. Tenuipalpus rodionovi Chalilova, 1953: 550; Azerbaidjan, ex Cydonia sp. (Rosaceae). Suspected synonym of: Tenuipalpus granati Sayed—Pritchard & Baker (1958), Wainstein (1960). Type Depository: Unknown or not specified. 795. Tenuipalpus rosae Kadzhaja, 1955: 65; Ukraine, ex Rosa sp. (Rosaceae). Species designations: Gnathopalpus rosae—Mitrofanov (1973b), Colopalpus rosae—Mitrofanov & Strunkova (1979). Redescriptions: Collyer (1973b), Hatzinikolis (1986a, 1987), Livschitz & Mitrofanov (1967), Meyer (1979), Mitrofanov (1973), Mitrofanov & Strunkova (1979), Pritchard & Baker (1958). Type Depository: Unknown. 796. Tenuipalpus rusapensis Meyer, 1979: 15; Lesapi River, 3 km from Rusape, Zimbabwe (formerly Rhodesia), ex Brachystegia manga De Wild. (Caesalpiniaceae). Redescription: Meyer (1993). Type Depository: SANC. 797. Tenuipalpus sagittus Meyer, 1993: 71; Swellendam, Cape Province, South Africa, ex unidentified species of Restionaceae. Type Depository: SANC. 798. Tenuipalpus salicis Al-Gboory, 1987: 150; Kerbala, Iraq, ex Salix alba L. (Salicaceae). Type Depository: Entomology Museum, College of Agriculture, Department of Plant Protection, Abu-Ghraib, Baghdad, Iraq. 799. Tenuipalpus sanblasensis De Leon, 1957: 89; San Blas, Nayarit, Mexico, ex “naranjilla” (Solanaceae). Redescriptions: Baker & Tuttle (1987), Collyer (1973b), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: MCZ. 800. Tenuipalpus sandyi De Leon, 1965a: 69; near Bartica Nature Reserve, Guyana (= British Guiana), ex Humiria balsamifera Mart. var. floribunda (Mart.) Cuatrec. (Humiriaceae). Redescriptions: Collyer (1973b), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: MCZ. 801. Tenuipalpus santae Manson, 1963: 213; Santa Rosa, Costa Rica, ex fence tree. Redescriptions: Collyer (1973b), Mairena & Ochoa (1989), Meyer (1979), Mitrofanov & Strunkova (1979), Salas & Ochoa (1986). Type Depository: USNM, No. 2885. 802. Tenuipalpus sanyaensis Yin, Cui & Lin, 1995: 62, Sanya City, Hainan Province, China, ex Lannea grandis Engl. (Anacardiaceae). Type Depository: Department of Plant Protection, Sheyang Agricultural University, China.

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803. Tenuipalpus scitulus Meyer, 1993: 45; Skukuza, Kruger National Park, Transvaal, South Africa, ex Cassia sp. (Caesalpiniaceae). Type Depository: SANC. 804. Tenuipalpus sclerocaryae Meyer, 1979: 17; Pongola, Transvaal, South Africa, ex Sclerocarya caffra Sond. (Anacardiaceae). Redescription: Meyer (1993). Type Depository: SANC. 805. Tenuipalpus senecionis Collyer, 1973c: 945; Waitakere Ranges, Auckland, New Zealand, ex Senecio kirkii Hook. & Kirk (Asteraceae). Redescriptions: Meyer (1979), Mitrofanov & Strunkova (1979), Mitrofanov & Sharonov (1983). Type Depository: Entomology Division, Department of Scientific and Industrial Research, Auckland, New Zealand. 806. Tenuipalpus sharmai Sadana & Gupta, 1984: 143; Agricultural University Ludhiana, Punjab, India, ex Helianthus annuus L. (Asteraceae). Redescriptions: Sadana (1997), Sadana & Gupta (1984). Type Depository: PAUP. Note: host originally listed as Helianthus annus. 807. Tenuipalpus simarubae De Leon, 1965c: 196; N Coamo, Puerto Rico, ex Bursera simaruba (L.) Sarg. (Burseraceae). Redescriptions: Collyer (1973b), De Leon (1965b), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: MCZ. 808. Tenuipalpus simplex Vitzthum, 1920: 8; Weimar, Germany, ex apple (Rosaceae). Species designations: Tenuipalpus palmatus var. simplex—original designation, Tenuipalpus simplex—Baker & Pritchard (1953b). Redescriptions: Baker & Pritchard (1953b), Pritchard & Baker (1958), Vitzthum (1929). Type Depository: Unknown. 809. Tenuipalpus simplychus Cromroy, 1958: 85; Maricao Insular Forest, Puerto Rico, ex Cordia sulcata DC. (Boraginaceae). Redescriptions: Collyer (1973b), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: Collection at Department of Entomology at North Carolina Stage College, USA. 810. Tenuipalpus smithi Meyer, 1979: 21; Pongola, Transvaal, South Africa, ex Acacia tortilis (Forsskal) Hayne (Mimosaceae). Redescription: Meyer (1993). Type Depository: SANC. 811. Tenuipalpus solanensis Sadana & Gupta, 1984: 141; Solan, India, ex Punica granatum L. (Punicaceae). Redescriptions: Sadana (1997), Sadana & Gupta (1984). Type Depository: PAUP. Note: suspected senior synonym of Tenuipalpus guptai Sadana & Gupta. 812. Tenuipalpus sophiae Meyer, 1979: 13; Great Fish River, Cape Province, South Africa, ex Acacia karroo Hayne (Mimosaceae). Redescription: Meyer (1993). Type Depository: SANC. 813. Tenuipalpus sparsus Chaudhri, Akbar & Rasool, 1974: 62; 3.2 km E Hyderabad, Sind, Pakistan, ex Ipomoea biloba Forssk. (Convolvulaceae). Resdescription: Akbar & Chaudhri (1981). Type Depository: Department of Entomology, University of Agriculture, Lyallpur, Pakistan. 814. Tenuipalpus spatulatus Wang, 1983: 61; Haian Island, Guangdong, China, ex an unidentified Asteraceae. Type Depository: Institute of Zoology, Academia Sinica, China.

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815. Tenuipalpus stativus Chaudhri, Akbar & Rasool, 1974: 65; 3.2 km N Leiah, Punjab, Pakistan, ex Aerva javanica Juss. (Amaranthaceae). Redescription: Akbar & Chaudhri (1981). Type Depository: Department of Entomology, University of Agriculture, Lyallpur, Pakistan. Note: host originally listed as Aerua javanica. 816. Tenuipalpus stefani Meyer, 1993: 44; Sekororo, southeast of Penge, Transvaal, South Africa, ex Euphorbia sp. (Euphorbiaceae). Type Depository: SANC. 817 Tenuipalpus striolatus Meyer, 1979: 34; 16 km S Potgietersrus, Transvaal, South Africa, ex Terminalia sericea Burch. & DC. (Combretaceae). Redescription: Meyer (1993). Type Depository: SANC. 818. Tenuipalpus tabebuiae De Leon, 1957: 85; San Blas, Nayarit, Mexico, ex Tabebuia pentaphylla DC. (Bignoniaceae). Redescriptions: Baker & Tuttle (1987), Collyer (1973b), De Leon (1965b), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: MCZ. 819. Tenuipalpus taonicus Ma & Yuan, 1980: 121; Ruijin, Jiangxi, China, ex Prunus persica (L.) Batsch (Rosaceae). Type Depository: CMNS. 820. Tenuipalpus tapirirae De Leon, 1965a: 69; near Bartica Nature Reserve, Guyana (= British Guiana), ex Tapirira guianensis Aubl. (Anacardiaceae). Redescriptions: Collyer (1973b), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: MCZ. 821. Tenuipalpus tauricus (Mitrofanov & Strunkova, 1978): 1098; Ukraine, ex Thymus sp. (Lamiaceae). Species designations: Gnathopalpus tauricus—original designation; Colopalpus tauricus—Mitrofanov & Strunkova (1979); Tenuipalpus tauricus—Meyer (1979). Redescriptions: Ghai & Shenhmar (1984), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: State Nikitsky Botanical Gardens (Yalta) and Institute of Zoology and Parasitology, Academy of Sciences of the Tadjil SSR (Dushanbe). 822. Tenuipalpus tectonae Mohanasundaram, 1981: 397; Panaji, Tamil Nadu, India, ex Tectona grandis L. (Verbenaceae). Redescription: Sadana (1997), Sadana & Gupta (1984). Type Depository: TNAU. 823. Tenuipalpus tepicanus De Leon, 1957: 85; San Blas, Nayarit, Mexico, ex “capulincillo” (cherry) (Ericaceae). Species designation: Tenuipalpus tepicensis [sic]—Baker & Tuttle (1987). Redescriptions: Baker & Tuttle (1987), Collyer (1973b), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: MCZ. 824. Tenuipalpus terminaliae De Leon, 1965a: 71; near Bartica Nature Reserve, Guyana (= British Guiana), ex Terminalia amazonica (J.F Gmel.) Exell (Combretaceae). Redescriptions: Collyer (1973b), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: MCZ. 825. Tenuipalpus tortulus Meyer, 1993: 72; Poung Kloof, Penge, Transvaal, South Africa, ex Xymalos monospora (Harv.) Baill. & Warb. (Monimiaceae). Type Depository: SANC. 826. Tenuipalpus toowongi Smiley & Gerson, 1995: 41; Toowong, Brisbane, Queensland, Australia, ex Eucalyptus sp. (Myrtaceae). Type Depository: QDPC.

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827. Tenuipalpus tetrazygiae De Leon, 1956a: 55; W Perrine, Florida, USA, ex Tetrazygia bicolor (P. Mill.) Cogn. (Melastomataceae). Redescriptions: Collyer (1973b), Meyer (1979), Mitrofanov & Strunkova (1979), Pritchard & Baker (1958), Sadana (1997). Type Depository: according to Pritchard & Baker (1958) in USNM. 828. Tenuipalpus transvaalensis Meyer, 1979: 68; Elephants River Gorge, Kruger National Park, Transvaal, South Africa, ex Boscia albitrunca Gilg & Benedict (Capparaceae). Redescription: Meyer (1993). Type Depository: SANC. 829. Tenuipalpus trichiliae De Leon, 1965b: 522; 22.4 km E Kingston, Morant Bay Road, Jamaica, ex Trichilia hirta L. (Meliaceae). Redescriptions: Collyer (1973b), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: MCZ. 830. Tenuipalpus trifoliatae Mohanasundaram, 1983a: 148; Vriddhachalam, Tamil Nadu, India, ex an unidentified hedge plant. Redescription: Sadana (1997). Type Depository: TNAU. 831. Tenuipalpus trisegmentus Siddiqui & Chaudhri, 1972: 328; Lyallpur, Faisalabab, Punjab, Pakistan, ex Morus alba L. (Moraceae). Species designation: Tenuipalpus trisegmentum [sic]—Mitrofanov & Strunkova (1979). Redescriptions: Akbar & Chaudhri (1981), Chaudhri (1972c), Chaudhri et al. (1974), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: Department of Entomology, West Pakistan Agricultural University Lyallpur, Pakistan. 832. Tenuipalpus trisetosus Baker & Tuttle, 1964: 73; Sells, Arizona, USA, ex Yucca elata (Engelm.) Engelm. (Agavaceae). Species designation: Tuttlepalpus trisetosus—Mitrofanov (1973). Redescriptions: Collyer (1973b), Meyer (1979), Mitrofanov (1973), Mitrofanov & Strunkova (1979). Type Depository: USNM, No. 3035. 833. Tenuipalpus tuttlei Ochoa, 1988: 225; replacement to Tenuipalpus chamaedoreae Baker & Tuttle (1987), name preoccuped by Tenuipalpus chamaedorea Salas & Ochoa (1985). (Article 58.7 4th Edition International Code of Zoological Nomenclature). Type Depository: USNM. Synonym: 833a. Tenuipalpus chamaedoreae Baker & Tuttle, 1987: 125; Mexico, intercepted at New York City, USA, ex Chamaedorea sp. (Arecaceae)—Ochoa (1988). Type Depository: USNM. 834. Tenuipalpus ueckermanni Meyer, 1979: 29; Mkuze Game Reserve, Natal, South Africa, ex Peltophorum africanum Sond. (Caesalpiniaceae). Redescription: Meyer (1993). Type Depository: SANC. 835. Tenuipalpus umarii Hasan, Wakil & Bashir, 2006: 384; Holotype, female, 2 km N Taxila, Pakistan, 26/10/2002, H. Hasan coll, ex Punica gramatum (Punicaceae). Redescription: Meyer (1993). Type Depository: SANC. 836. Tenuipalpus unimerus De Leon, 1957: 84; San Blas, Nayarit, Mexico, ex “guamacate” (Lauraceae). Redescriptions: Baker & Tuttle (1987), Collyer (1973b), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: MCZ. Note: According to Baker & Tuttle (1987) the type host in the original description is Persea americana P. Mill. (Lauraceae).

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837. Tenuipalpus unonopsonis De Leon, 1965a: 66; Bartica Nature Reserve, Guyana (= British Guiana), ex Unonopsis guatterioides (DC.) R.E. Fr. (Annonaceae) Redescriptions: Collyer (1973b), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: MCZ. Note: host originally listed as Unonopsis guatteroides. 838. Tenuipalpus uvae De Leon, 1962: 205; San Blas, Nayarit, Mexico, ex “uva” (a large tree). Redescriptions: Baker & Tuttle (1987), De Leon (1967), Mairena & Ochoa (1989), Meyer (1979), Mitrofanov & Strunkova (1979), Salas & Ochoa (1986). Type Depository: MCZ. 839. Tenuipalpus velitor Hasan, Akbar & Bashir, 2003: 208, 1 km E Hasanabdal, Pakistan, ex an unidentified plant. Type Depository: Acarology Research Laboratory, Department of Entomology, University of Agriculture, Faisalabad, Pakistan. 840. Tenuipalpus venustus Collyer, 1973c: 951; Mount Evans Track, Canaan Area, Takaka Hill, Nelson, New Zealand, ex Libocedrus bidwillii Hook. (Cupressaceae). Redescriptions: Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: Entomology Division, Department of Scientific and Industrial Research, Auckland, New Zealand. Note: host originally listed as Libocedrus bidwilli. 841. Tenuipalpus vernoniae Meyer, 1993: 76; Rustenburg Nature Reserve, Transvaal, South Africa, ex Vernonia oligocephala Sch. Bip. & Walp. (Asteraceae). Type Depository: SANC. 842. Tenuipalpus vexus De Leon, 1965b: 520; Santo Domingo, Dominican Republic, ex Tabebuia sp. (Bignoniaceae). Redescriptions: Collyer (1973b), De Leon (1967), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: MCZ. 843. Tenuipalpus victoriae De Leon, 1967: 43; Cleaver Reserve, Arima, Trinidad and Tobago, ex an unidentified shrub or young tree. Redescription: Meyer (1979). Type Depository: MCZ. 844. Tenuipalpus vitexi Meyer, 1993: 31; Rustenburg, Transvaal, South Africa, ex Vitex harveyana H.H.W. Pearson (Verbenaceae). Type Depository: SANC. 845. Tenuipalpus viticola Al-Gboory, 1987: 149; Abu-Ghraid, Baghdad, Iraq, ex Vitis vinifera L. (Vitaceae). Type Depository: Entomology Museum, College of Agriculture, Department of Plant Protection, Abu-Ghraib, Baghdad, Iraq. 846. Tenuipalpus vriddagiriensis Mohanasundaram, 1983a: 147; Vriddhachalam, Tamil Nadu, India, ex an unidentified hedge plant. Redescription: Sadana (1997). Type Depository: TNAU. 847. Tenuipalpus waqasii Hasan, Wakil & Bashir, 2004: 97; 2 km E Alipur, Muzzafarghar, Punjab, Pakistan, ex Aerva sp. (Amaranthaceae). Type Depository: Acarology Research Laboratory, Department of Agric. Entomology, University of Agriculture, Faisalabad, Pakistan. Note: host originally listed as Aerua sp. 848. Tenuipalpus xerocolus Meyer, 1993: 79; Karro National Park, near Beaufort West, Cape Province, South Africa, ex an unidentified wild herb. Type Depository: SANC. 849. Tenuipalpus xylosmae De Leon, 1965b: 519; Bath Fountain, Saint Thomas, Jamaica, ex Xylosma G. Forst. aff. nitida (Salicaceae).

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Redescriptions: Collyer (1973b), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: MCZ. 850. Tenuipalpus yarensis Hasan, Bashir & Wakil, 2003: 359; Faisalabad, Punjab, Pakistan, ex Phoenix dactylifera L. (Arecaceae). Type Depository: Acarology Research Laboratory, Department of Agric. Entomology, University of Agriculture, Faisalabad, Pakistan. 851. Tenuipalpus yousefi Nassar & Ghai, 1981: 384; Indian Agricultural Research Institute, New Delhi, India, ex Psidium guajava L. (Myrtaceae). Species designation: Tenuipalpus yoursefi [sic]—Sadana (1997). Redescriptions: Gupta & Chatterjee (1997), Sadana (1997), Sadana & Gupta (1984). Type Depository: INPC. 852. Tenuipalpus zanthus De Leon, 1965b: 521; 22.4 km E Kingston, Jamaica, ex Zanthoxylum elephantiasis Macfad. (Rutaceae). Species designation: Tenuipalpus santhus—Mitrofanov & Strunkova (1979). Redescriptions: Collyer (1973b), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: MCZ. 853. Tenuipalpus zeyheri Meyer, 1993: 31; Vygeboom Dam, near Badplaas, Transvaal, South Africa, ex Scolopia zeyheri (Nees) Harv. (Flacourtiaceae). Type Depository: SANC. 854. Tenuipalpus zhengzhouensis Xu & Yin, 1994: 308; Zhengzhou, Henan Province, China, ex Magnolia denudata Desr (Magnoliaceae). Type Depository: Department of Plant Protection, Shenyang Agricultural University, China. 855. Tenuipalpus zhizhilashviliae Reck, 1953: 179; Georgia, ex Diospyros kaki L. (Ebenaceae). Redescriptions: Collyer (1973b), Ehara (1964), Hatzinikolis (1986a, 1987), Livschitz & Mitrofanov (1967), Ma & Yuan (1977), Meyer (1979), Livschitz & Mitrofanov (1967), Mitrofanov & Strunkova (1979), Pritchard & Baker (1958), Tseng (1974). Type Depository: Unknown. Suspected senior synonym of: Tenuipalpus japonicus Nishio—Mitrofanov & Strunkova (1979). 856. Tenuipalpus zuluensis Meyer, 1979: 46; Mkuzi Game Reserve, Natal, South Africa, ex Euclea divinorum Hiern (Ebenaceae). Redescription: Meyer (1993). Type Depository: SANC.

Terminalichus Anwarullah & Khan 1973 Terminalichus Anwarullah & Khan, 1973: 5. Type species: Terminalichus karachiensis Anwarullah & Khan, 1973: 154; Gupta & Chatterjee (1997), Nassar & Ghai, 1981: 379; Sadana 1997: 100. Comment: Sadana (1997) provided a key to the Indian species of Terminalichus. 857. Terminalichus delhiensis Maninder & Ghai, 1978: 219; Indian Agrigultural Research Institute, New Delhi, India, ex Terminalia arjuna Wight & Arn. (Combretaceae). Redescriptions: Gupta & Chatterjee (1997), Nassar & Ghai (1981), Sadana (1997), Sadana & Sidhu (1989). Type Depository: Indian Agricultural Research Institute Campus, New Delhi, India. 858. Terminalichus dehlonensis Sadana & Sidhu, 1989: 107; Dehlon, Ludhiana, India, ex Psidium guajava L. (Myrtaceae). Redescription: Sadana (1997). Type Depository: Unknown. 859. Terminalichus karachiensis Anwarullah & Khan, 1973: 6; Karachi, Sind, West Pakistan, ex Terminalia catappa L. (Combretaceae).

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Redescriptions: Gupta & Chatterjee (1997), Maninder & Ghai (1978), Nassar & Ghai (1981), Sadana (1997), Sadana & Sidhu (1989). Type Depository: Collections of the Acarology Section, P.C.S.I.R. Laboratories, Karachi, Pakistan. 860. Terminalichus panajiensis Maninder & Ghai, 1978: 217; Panaji, Goa, India, ex Terminalia sp. (Combretaceae). Species designation: Terminalicus penajiensis [sic]—Sadana & Sidhu (1989). Redescription: Sadana (1997). Type Depository: Indian Agricultural Research Institute Campus, New Delhi, India. 861. Terminalichus psidi Mohanasundaram, 1983c: 221; Villupuram, Tamil Nadu, India, ex Psidium guajava L. (Myrtaceae). Redescription: Sadana (1997), Sadana & Sidhu (1989). Type Depository: Unknown or not specified. Note: host originally listed as Psidium gujava. 862. Terminalichus serratus Nassar & Ghai, 1981: 380; Indian Agricultural Research Institute, New Delhi, India, ex Psidium guajava L. (Myrtaceae). Resdescriptions: Gupta & Chatterjee (1997), Sadana (1997). Type Depository: INPC.

Ultratenuipalpus Mitrofanov 1973 Ultratenuipalpus Mitrofanov, 1973b: 1318. Type species: Tenuipalpus meekeri De Leon, 1957: 82; Baker & Yin, 1988: 151; Baker & Tuttle, 1987: 136; Meyer, 1979: 3; Meyer, 1993: 39; Mitrofanov & Strunkova, 1979: 38; Smiley & Gerson, 1995: 41; Wang & Cui, 1993: 111. Extenuipalpus Reck, 1959. Type species: Tenuipalpus quadrisetosus Lawrence, 1940; new synonomy. 863. Ultratenuipalpus aberrans (Collyer, 1973c)—new combination: 926; Buller River, New Zealand, ex Pyrus malus L. (Rosaceae) in Nothofagus (Fagaceae) forest. Species designations: Tenuipalpus aberrans—original designation, Amblypalpus aberrans—Mitrofanov & Strunkova (1979). Redescriptions: Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: Entomology Division, Department of Scientific and Industrial Research, Auckland, New Zealand. Notes: it appears that Nothofagus spp. are the main host; when the mite was collected from other hosts, the plants were located inside Nothofagus forest. 864. Ultratenuipalpus acharis (Gonzalez, 1968): 45—new combination; 30 km S Los Angeles, Malleco, Chile, ex Gevuina avellana Molina (Proteaceae). Species designation: Tenuipalpus acharis—original designation, Extenuipalpus acharis—Mitrofanov & Strunkova (1979). Redescriptions: Collyer (1973b), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: Coleccion de la Estacion Experimental Agronomica, Universidad de Chile (CEA); Museo de Historia Natural Santiago (MHNS). Chile. 865. Ultratenuipalpus arboreus (Collyer, 1973c): 941—new combination; Canaan Trig, Takaka Hill, Nelson Province, New Zealand, ex Phyllocladus alpinus Hooker (Phyllocladaceae). Species designation: Tenuipalpus arboreus—original designation, Extenuipalpus arboreus—Mitrofanov & Strunkova (1979). Redescriptions: Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: Entomology Division, Department of Scientific and Industrial Research, Auckland, New Zealand. 866. UItratenuipalpus asteliae (Collyer, 1973c): 937—new combination; Chateau, Tongariro National Park, New Zealand, ex Astelia sp. (Asteliaceae). Species designation: Tenuipalpus asteliae—original designation, Extenuipalpus asteliae—Mitrofanov & Strunkova (1979). Redescriptions: Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: Entomology Division, Department of Scientific and Industrial Research, Auckland, New Zealand.

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867. Ultratenuipalpus asteliicola (Collyer, 1973c): 935—new combination; Waitakere Ranges, Auckland, New Zealand, ex Astelia sp. (Asteliaceae). Species designation: Tenuipalpus asteliicola—original designation, Extenuipalpus asteliicola—Mitrofanov & Strunkova (1979). Redescriptions: Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: Entomology Division, Department of Scientific and Industrial Research, Auckland, New Zealand. 868. Ultratenuipalpus baltazarae Rimando, 2001: 108; Luzon Island: Magat Reforestation Project, 22, Philippines, ex fern. Type Depository: University of the Philippines Los Baños, Museum of Natural History, College, Laguna, Philippines. Note: according to Rimando (2001), the species identified by Corpuz-Raros (1978, 1986) was misidentified. 869. Ultratenuipalpus canelae (Gonzalez, 1968): 45—new combination; Parque Nacional de Fray Jorge, Coquimbo, Chile, ex Drymis winteri Forst. (Winteraceae). Species designation: Tenuipalpus canelae—original designation, Extenuipalpus canelae—Mitrofanov & Strunkova (1979). Redescriptions: Collyer (1973b), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: Coleccion de la Estacion Experimental Agronomica, Universidad de Chile (CEA); Museo de Historia Natural Santiago (MHNS). 870. Ultratenuipalpus carpodeti (Collyer, 1973c): 930—new combination; Buller Gorge, near Inangahua, Nelson Province, New Zealand, ex Carpodetus serratus JR Forst. et G. Forst. (Carpodetaceae). Species designation: Tenuipalpus carpodeti—original designation, Extenuipalpus carpodeti—Mitrofanov & Strunkova (1979). Redescriptions: Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: Entomology Division, Department of Scientific and Industrial Research, Auckland, New Zealand. 871. Ultratenuipalpus charlini (Gonzalez, 1968): 43—new combination; Parque Nacional de Fray Jorge, Coquimbo, Chile, ex Aextoxicon punctatum Ruiz & Pav. (Aextoxicaceae). Species designation: Tenuipalpus charlini—original designation, Extenuipalpus charlini—Mitrofanov & Strunkova (1979). Redescriptions: Collyer (1973b), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: Colecciòn de la Estacion Experimental Agronomica, Universidad de Chile (CEA); Museo de Historia Natural Santiago (MHNS), Chile. 872. Ultratenuipalpus coprosmae (Collyer, 1964): 434—new combination; Hunua, near Auckland, New Zealand, ex Coprosma sp. (Rubiaceae). Species designation: Tenuipalpus coprosmae—original designation, Extenuipalpus coprosmae—Mitrofanov & Strunkova (1979). Redescriptions: Collyer (1973b), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: BMNH. 873. Ultratenuipalpus coprosmicus (Collyer, 1973c): 931—new combination; Cobb Reservoir, Upper Takaka, Nelson, North West Forest Park, New Zealand, ex Coprosma sp. (Rubiaceae). Species designation: Tenuipalpus coprosmicus—original designation, Extenuipalpus coprosmicus—Mitrofanov & Strunkova (1979). Redescriptions: Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: Entomology Division, Department of Scientific and Industrial Research, Auckland, New Zealand. 874. Ultratenuipalpus cyathodis (Collyer, 1973c): 942—new combination; Palmers Bush, Eves Valley, Nelson, New Zealand, ex Cyathodes fasciculata (G. Forst.) Allan (Epacridaceae). Species designation: Tenuipalpus cyathodis—original designation, Extenuipalpus cyathodis—Mitrofanov & Strunkova (1979). Redescriptions: Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: Entomology Division, Department of Scientific and Industrial Research, Auckland, New Zealand. 875. Ultratenuipalpus filicicola (Wang, 1983): 62; Mount Emei, Sichuan, China, ex Cyclosorus sp. (Thelypteridaceae). Species designations: Tenuipalpus filicicola Wang (1983)—original designation; Ultratenuipalpus filicicola—Wang & Cui (1993).

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Resdescrition: Wang & Cui (1993). Type Depository: Institute of Zoology, Academia Sinica, China. 876. Ultratenuipalpus hainanensis (Wang, 1980): 387; Hainan Island, Guandong, China, ex Lygodium japonicum (Thunb.) Sw. (Lygodiaceae). Species designations: Tenuipalpus hainanensis—original designation; Ultratenuipalpus hainanensi—Wang & Cui (1993). Redescription: Wang & Cui (1993). Type Depository: Institute of Zoology, Academia Sinica, China. 877. Ultratenuipalpus lacorpuzrarosae Rimando, 2001: 110; Panaon Island off Leyte: Bgy. Sta. Paz, San Francisco, Southern Leyte, Philippines, ex fern. Type Depository: University of the Philippines Los Baños, Museum of Natural History, College, Laguna, Philippines. 878. Ultratenuipalpus meekeri (De Leon, 1957): 82; San Blas, Nayarit, Mexico, ex fern in mangrove swamp. Species designations: Tenuipalpus meekeri—original designation, Ultratenuipalpus meekeri—Mitrofanov (1973b). Redescriptions: Baker & Tuttle (1987), Baker et al. (1975), Collyer (1973b), Corpuz-Raros (1989), De Leon (1967), Meyer (1979, 1993), Mitrofanov (1973b), Mitrofanov & Strunkova (1979), Rimando (2001), Sadana (1997), Sadana & Gupta (1983), Sadana & Sidhu (1990), Wang & Cui (1993). Type Depository: MCZ. 879. Ultratenuipalpus nothofagi (Collyer, 1973c): 934—new combination; Abel Tasman National Park, New Zealand, ex Nothofagus fusca (Hook) Oerst. (Fagaceae). Species designation: Tenuipalpus nothofagi—original designation, Extenuipalpus nothofagi—Mitrofanov & Strunkova (1979). Redescriptions: Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: Entomology Division, Department of Scientific and Industrial Research, Auckland, New Zealand. 880. Ultratenuipalpus pteriphilus Meyer, 1993: 42; Magaliesberg Mountains, near Maanhaarrand, Transvaal, South Africa, ex Pteridium aquilinum (L.) Kuhn (Dennstaedtiaceae). Redescription: Rimando (2001). Type Depository: SANC. 881. Ultratenuipalpus quadrisetosus (Lawrence, 1940): 114—new combination; Stella Bush, Durban, South Africa, ex Cryptocarya woodii Engl. (Lauraceae). Species designation: Tenuipalpus quadrisetosus—original designation, Extenuipalpus quadrisetosus—Mitrofanov & Strunkova (1979). Redescriptions: Baker & Pritchard (1953b, 1960), Collyer (1973b), Lawrence (1943), Meyer (1979, 1993), Meyer & Ryke (1959), Mitrofanov & Strunkova (1979), Pritchard & Baker (1958), Sadana (1997). Type Depository: Unknown. Synonym: 881a. Tenuipalpus gibbus Lawrence, 1943: 42; Stella Wood, Durban, South Africa, ex Cryptocarya woodii Engl. (Lauraceae)—Baker & Tuttle (1953). Type Depository: Unknown. 882. Ultratenuipalpus rubi (Collyer, 1964): 432—new combination; Little Huia, near Auckland, New Zealand, ex Rubus sp. (Rosaceae). Species designation: Tenuipalpus rubi—original description, Extenuipalpus rubi—Mitrofanov & Strunkova (1979). Redescriptions: Collyer (1973b), Meyer (1979), Mitrofanov & Strunkova (1979). Type Depository: BMNH. 883. Ultratenuipalpus umtataensis Meyer, 1979: 3; 46.7 km from Umtata, Transkei, South Africa, ex an unidentified plant. Species designation: Ultratenuipalpus umatataensis [sic]—Sadana & Sidhu (1990). Redescriptions: Meyer (1993), Rimando (2001), Sadana & Sidhu (1990), Wang & Cui (1993). Type Depository: SANC. 884. Ultratenuipalpus womersleyi (Pritchard & Baker, 1958): 238—new combination; Redland Bay, Queensland, Australia, ex Leptospermum sp. (Myrtaceae).

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Species designations: Tenuipalpus womersleyi—original designation, Extenuipalpus womersleyi—Mitrofanov & Strunkova (1979). Redescriptions: Collyer (1964, 1973b), Meyer (1979), Mitrofanov & Strunkova (1979), Rimando (2001), Smiley & Gerson (1995). Type Depository: BMNH. 885. Ultratenuipalpus younguisti Baker & Tuttle, 1987: 137; Mexico, intercepted at San Pedro, California quarantine station, California, USA, ex unidentified plant. Redescription: Meyer (1993), Rimando (2001), Wang & Cui (1993). Type Depository: USNM.

Urigersonus Mesa, Ochoa & Evans, NEW GENUS Type species: Ultratenuipalpus bunyai Smiley & Gerson, 1995: 41, by monotypy. Generic Diagnosis: Palpus 4-segmented, with solenidion on palptarsus inserted at the base of the segment. Rostral shield deeply notched medially. Shape of idiosoma similar to that of Ultratenuipalpus species. Opisthosoma slightly narrower than proprodosoma and somewhat truncate caudally. Dorsum of opisthosoma with 9 pairs of setae: c1, c3, d1, d3, e1, e3, f3, h1, h2; setae c1, d1 and e1 setae minute and slender; d3 slightly longer and thicker; remaining setae (c3, e3, h2 and h1) large and broadly spatulate.setae f3 and sc2 very long and whiplike, more than 10 times longer than other dorsal opisthosomal setae. Pregenital and genital areas with smooth, fine striae. Comparison; In Ultratenuipalpus setae f3 and sc2 are not long and whiplike, but rather similar in size and shape to other opisthosomal dorsal setae; the palpus is usually 4-segmented (rarely 3) but the solenidion on palptarsus is inserted at the apex of the segment. Etymology: This genus is named in honor of Dr. Uri Gerson for his many contributions to the study of mites. 886. Urigersonus bunyai (Smiley & Gerson) 1995: 41—new combination; Toowong, Australia, ex Araucaria bidwilli Hook. (Araucariaceae). Species designations: Ultratenuipalpus bunyai Smiley & Gerson, 1995: 41—original designation. Redescriptions: Rimando (2001). Type Depository: QDPC.

species Incertae sedis 887. Priscapalpus gurdaspurensis Kaur & Sadana, 1999: 126; Gurdaspur, Punjab, India, ex Vitis vinifera L. (Vitaceae). Type Depository: INPC. Note: according to the characters mentioned it is most likely an immature. 888. Priscapalpus piarai Sadana & Sidhu, 1989: 75; Gurdaspur, Punjab, India, ex Psidium guajava L. (Myrtaceae). Redescription: Sadana (1997). Type Depository: Unknown. Note: according to the characters mentioned it is most likely an immature. 889. Ultratenuipalpus gonianaensis Sadana & Sidhu, 1990: 358; Goniana, Faridkot, Punjab, India, ex Citrus limon L. Burm (Rutaceae). Redescriptions: Meyer (1993), Rimando (2001), Sadana (1997). Type Depository: PAUP. Note: according to the characters mentioned it is most likely an immature.

Addendum Acaricis Beard & Gerson 2009: 31 Acaricis Beard & Gerson 2009: 32; Type species: Acaricis plana Beard & Gerson, 2009.

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890. Acaricis danutae Beard & Gerson, 2009: 32. Cooyal, New South Wales, Australia, ex Carex sp. (Cyperaceae). Type Depository: ASCU. 891. Acaricis plana Beard & Gerson, 2009: 32. 10 km NNW Murgon, Queensland, Australia, ex Scleria sphacelata F. Muell. (Cyperaceae). Type Depository: QM.

FINAL CONSIDERATIONS A total of 34 genera and 891 valid species of tenuipalpids are reported herein. Each of a few genera are comprised of relatively large numbers of species, whereas others contain one or a few species. The number of species in the five largest genera are: Tenuipalpus 305, Brevipalpus 281, Cenopalpus 59, Aegyptobia 93, and Dolichotetranychus 22. The following genera have less than 10 species: Acaricis, Prolixus, Pseudoleptus, Terminalichus, Afronychus, Capedulia, Crossipalpus, Priscapalpus, Obdulia, Obuloides, Phyllotetranychus and Tenuilichus. The following genera are monospecific: Australopalpus, Chaudhripalpus, Coleacarus, Krugeria, Larvacarus, Lisaepalpus, Macfarlaniella, Magdalenapalpus, Philippipalpus, Raoiellana, Tegopalpus and Urigersonus (Table 1). The largest number of species have been described from the United States of America (17.3%), followed by Mexico (16.3%), Pakistan (16%) and South Africa (13.8%). The tenuipalpid fauna of China, Southeast Asia, the Pacific Islands, Central and South America, Russia, Australia and several other countries is not well known. The number and distribution of species do not show a clear ecological pattern at the family level. A few genera, especially Brevipalpus, Dolichotetranychus and Tenuipalpus, are represented by species from multiple continents and on a wide range of host plants. The genus Brevipalpus is most speciose in the Nearctic, Tenuipalpus in the Afrotropical, Raoiella in the Oriental, and Ultratenuipalpus in the Pacific Island regions (Table 1). Other genera are geographically restricted. For example Afronychus, Capedulia, Coleacarus and Krugeria, are restricted to Africa; Australopalpus, Chaudhripalpus, Crossipalpus, Lisaepalpus, Macfarniella, Magdalenapalpus and Tegopalpus, to Australia; Larvacarus, Neoraoiella, Obuloides, Philippipalpus, Rarosiella, Tenuilichus and Terminalichus, to Southeast Asia; Obdulia, to the Mideast; Phyllotetranychus, to Egypt and Mauritius Island; and Raoiellana, to Turkey (Table 1). This publication is based mostly on an evaluation of the current literature. An adequate revision, with examination of type specimens, may indicate a considerable number of synonyms, resulting from the occurrence of intraspecific morphological variation. Welbourn et al. (2003) conducted a detailed study of the morphological variation present within Brevipalpus phoenicis and compared this variation with B. californicus and B. obovatus. Similar studies are needed for other species of Tenuipalpidae. We believe that there are likely to be several to many tenuipalpid species that will prove to be synonymous with previously described species once we have a better understanding of the intraspecific variation that occurs within tenuipalpid species.

RECOMMENDATIONS After reading hundreds of species descriptions, comparing hundreds of type specimens present at our institutions, and studying several genera under Low Temperature Scanning Electron Microscope (LT-SEM), we noticed several important systematic problems. Based on these findings, we recommend that future research on the Tenuipalpidae observe the following protocol: a. Descriptions should include line drawings illustrating the complete dorsal and ventral features of the mite. The ornamentation of the dorsum of the propodosoma and opisthosoma as well as that of the ventral plates provide important clues for determining the species’ identity. The shape of the dorsal opisthosomal setae and the number of palpal segments and chaetotaxcy are also important. When

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b.

c.

d.

e.

phase contrast, differential phase contrast, or SEM micrographs are used, it is essential that they be accompanied by line drawings showing details of the key characters. Description of new taxa based on a single specimen is generally discouraged, especially for new species that are considered to be very similar to known species. Single specimens may represent aberrant variations of already described species or may appear to have distinct characteristics due to age of the mite, what it has fed upon, the type of clearing agent, mounting medium or technique used to mount the specimen. Contrary to the common name of the Tenuipalpidae as a whole, the dorsum of “flat mites” is not actually flat, but rather covered with ridges, bumps and protuberances, especially on the central and submedial areas. The integument of the mite becomes soft when Nesbitt’s solution, lactophenol, Vitzthum’s fluid, or KOH are used as clearing agents, so that when the cover slip is put in place, these sculptural characters can be distorted or made smoother, making the specimen apparently different from the species’ normal appearance. Furthermore, the color and the strength of the integument sculpture depend upon the age and food content in the mite. Older females often exhibit more prominent and detailed ornamentation than younger females. The food content directly affects the coloration of the mite (Morishita 1954, Haramoto 1969). Also, specimens mounted in Hoyer’s can be overcleared if dried for more than 3-5 days and/or at a temperature higher than 35–55 o C. Determining the family, genus and species of the host plant may facilitate identification because some genera and species show significant host preference and this can provide a clue for the determination of genus or species. For example, almost all species in the genus Raoiella are associated with dicotyledons, whereas most species of Pentamerismus are associated with conifers in the family Cupressaceae. Most of the tenuipalpids important to agriculture, however,are generalists and feed on a wide range of host plants. Wherever possible, all stages of the species, egg, larva, protonymph and deutonymph, in addition to the adult male and female should be collected. Quite often, closely related tenuipalpid species can be distinguished by characters found only in the immature form, especially the deutonymph. A description of the deutonymph, preferably with a drawing of the entire dorsum of the mite, should be included in the description. For example, the adults of Brevipalpus trinidadensis and B. californicus are very similar; however, their deutonymphs are readily distinguished. We consider important the record of the habits and location of the mite on the plant, and unusual host features determined by the mite, such as galls, chlorosis, necrosis, and cracking of the epidermis of the fruit or leaves which may result from the mite feeding, and any fungus or fungal spores associated with the mite. These may be symptoms of diseases especially considering the close association of certain species of the genus Brevipalpus with plant viruses and other diseases (Kitajima et al. 2003, Ochoa et al. 1994).

The widespread epidemic of the leprosis virus in the Americas (Childers et al. 2003) and the recent introduction of invasive mites such as Raoiella indica to the Americas (Flechtmann and Etienne 2004) demonstrate the importance of the flat mites to the agricultural economy of the nations of the world. It is the hope of the authors that this catalogue will provide a tool and guide to future faunistic studies conducted in Asia, South America, Australia and other areas where the species composition, diversity and biology of flat mites are not well known.

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MORPHOLOGY The body of all mites is divided in two parts—(i) gnathosoma, the anterior portion of the body which includes the mouthparts, and (ii) idiosoma, the posterior portion of the body. The idiosoma is composed of three regions: (i) propodosoma (bearing legs I and II), (ii) metapodosoma (bearing legs III and IV) and (iii) opisthosoma (posterior to legs IV). The plane separating the first two regions is called sejugal plane, while the plane separating the last two regions is called disjugal plane. Transversal furrows are usually found at those planes; they are called sejugal and disjugal furrows, respectively. In the Acariformes, the superorder to which the Tenuipalpidae belong, the dorsum of the metapodosoma is considered to have disappeared, so that the posterior edge of the dorsum of the propodosoma becomes aposed to the anterior edge of the dorsum of the opisthosoma, or those edges are bridged by the merged sejugal and disjugal furrows.

FIGURE 1. Aegyptobia tragardhi female dorsum and venter (after Sayed 1950).

The Gnathosoma The mouthparts of tenuipalpids consists of a pair of recurved styletiform chelicerae, the stylophore and the rostrum; the latter houses the pharynx and bears a pair of simple palpi having 1-5 segments. The number of palpal segments is often used in characterizing the different genera (Figure 2). Tenuipalpids differ from the other families in the superfamily Tetranychoidea in lacking a claw on the penultimate segment of the palpus. The Idiosoma A. Dorsum 1. Propodosoma The propodosoma of the tenuipalpids has two pairs of simple eyes, with the exception of Raoiellana allium Baker and Tuttle which lacks eyes. This region bears 3 pairs of setae, with the exception of Tenuipalpus elegans, with only 2 pairs. The anteriormost pair (v2), lacking in T. elegans, are followed by the sc1 setae towards the lateral margin and just anterior to the eyes, and the sc2 setae, immediately posterior to the eyes.

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The anterior margin of the propodosoma of certain genera is modified into a sclerotized platelike structure called the rostral shield or prodorsal sclerite, which overhangs the gnathosoma.

FIGURE 2. Types of palpi found in the family Tenuipalpidae (after Pritchard & Baker 1958).

2. Opisthosoma The opisthosoma of tenuipalpids bears between 5 and 13 pairs of setae, that are located in 5 transverse rows. Setae of the most anterior row, along the anterior margin of the opisthosoma, are designated by the letter c with the dorsocentral pair (DC) of setae designated as (c1), the dorsosublateral pair (DSL) as c2, and the dorsolateral pair (DL) as c3. These setae are followed successively by setae in the d, e, f, and h rows; the letter g is not used to designate rows or setae on the dorsum of Acariformes, to preclude confusion with the genital setae on the venter, which is designated as g. The maximum number of dorsal opisthosomal setae consists of 3 pairs in each of the c, d and e rows, labelled as the c1, c2, c3, d1, d2, d3 and e1, e2, e3, respectively; and a maximum of 2 pairs in each of the f and h rows, and labelled as f2, f3 and h1, h2, respectively. 3. Determination of the dorsal setae To determine the dorsal setae, first locate the stable setae, present in nearly all tenuipalpid species. Based on our examination of the descriptions, illustrations and specimens of the known tenuipalpid species, we consider sc1, sc2, c3, d3, e3, h1 and h2 to be stable setae; they are present in all known species of tenuipalpids, with the possible exception of Krugeria ramose, depending upon whether one designates the pair of setae located on a same plate as e3, f3 or h2. With some reservations, we designate this pair of setae as h2, to be consistent with the presumed stability of these setae. Setae f3 are present in almost all species, with the exception of those species which have only 5 pairs of dorsolateral setae (Krugeria, Obuloides, Terminalichus species and Priscapalpus thomissus); these setae should be accounted for first when examining a tenuipalpid specimen and determining its setal complement. They are usually located along the lateral margin with the h1 setae being at the posterior apex. However, the position of one or more pairs of these setae may shift to the sublateral position, as in Dolichotetranychus. Therefore, where only one pair of setae is present laterad to the dorsocentral region, they are designated as c3, d3, e3 and h2. If there are two pairs of setae laterad to the dorsocentral region, the setae in between the dorsocentral pair and the dorsolateral pair (usually located in the submedial region) are designated as c2, d2, e2 or f2 depending upon their relative position on the dorsum, and after the stable setae have been accounted for.

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Next, account for the dorsocentral setae (c1, d1, e1). The majority of tenuipalpid species have three pairs of dorsocentral setae. Setae h1 are not counted as dorsocentral setae and setae f1 do not occur, or are not designated as such, in the Tenuipalpidae. At least one of the dorsocentral pairs of setae is absent in all known species of Terminalichus, Tenuilichus, Obuloides, Dolichotetranychus and in certain species of Tenuipalpus, Priscapalpus and Afronychus. Lastly, account for the sublateral setae (c2, d2, e2, f2). Because of the occasional shifting of the dorsocentral and/or dorsolateral setae to the sublateral region, we recommend that the dorsosublateral setae be designated as such when they are found located between the dorsocentral and the dorsolateral pair of setae in their respective row, with the exceptions of the f row which lacks setae f1, and the h row which lacks setae h3. For example, the 3 pairs of setae along the anterior margin of the opisthosoma (row c) of Aegyptobia species are designated c1, c2 and c3, with the c2 setae located between the c1 (dorsocentral) and the c3 (dorsolateral) pair of setae. The lateral margin of many species appears to have 6 or 7 pairs of setae (8 in Pentamerimus erythreus), due to the shifting of the dorsosublateral setae, c2, d2, e2 and/or f2, to the lateral margin. B. Venter The ventral setae have seldom been used in the classification of tenuipalpid genera and species, except for the designation of certain species groups in some of the genera. Setae 1b, 1c, 2b, 2c, 3b, 4b are associated with the coxae and setae 1a, 3a, 4a are intercoxal, inserted centrally on the venter. 1. Propodosoma—Normally, there are 5 pairs of setae on the venter of the propodosoma (1a, 1b, 1c, 2b, 2c). Setae 1a are located medially between the bases of coxae I; setae 1b are located anteromesad to 1a, and 1c is located posterolaterad to 1b (1b and 1c are associated with coxa I). Setae 2a are not known to occur in tenuipalpid species. Setae 2b are located usually at the base of coxae II and setae 2c are located posterolaterad to 2b. 2. Metapodosoma—There are two pairs of setae, often elongate setae, in the median area of the metapodosoma. Setae 3a are usually located between the bases of coxae III and setae 4a are usually located between the bases of coxae IV. Setae 3b and the 4b are usually short and located on coxae III and IV, respectively. 3. Opisthosoma—Normally, there are 5 pairs of ventral opisthosomal setae (ag, g1, g2, ps1, ps2) pairs of ventral opisthosomal setae and a maximum of 7 pairs when a third pair of genital setae (g3) and a third pair of anal plate setae (ps3) are present. When present, setae ag are located on the ventral plate or between the 4a metapodosomal setae and the genital setae (g1 and g2), the latter located on the genital plate. The anal plates bear 1–3 pairs of setae, ps1–ps3 (ps3 rarely present). TABLE 2. Opisthosomal setae, body shape, rostral shield and number of palpal segments. Genus/species Acaricis plana Aegyptobia tragardhi Afronychus cliffortiae Afronychus diastellae Afronychus mulibrinus Amblypalpus masakii Amblypalpus narsusikulovi Australopalpus alphitoniae Brevipalpus obovatus Brevipalpus californicus Capedulia calendulae Cenopalpus spinosus

Total 12 13 9 11 11 7 8 12 9 10 10 12

C1 C2 C3 D1 D2 D3 E1 E2 E3 F2 F3 H1 H2 BS RS Ps 0 0 1 1 0 1 1 0 1 1 1 1 1 1 1 4 1 1 1 1 1 1 1 1 1 1 1 1 1 4 0 5 1 0 1 1 0 1 1 0 1 0 1 1 1 1 0 3 1 0 1 1 0 1 1 1 1 1 1 1 1 1 0 3 1 1 1 1 0 1 1 1 1 0 1 1 1 1 0 3 0 0 1 1 0 1 1 0 1 0 0 1 1 3 1 3 1 0 1 1 0 1 1 0 1 0 0 1 1 3 1 3 1 1 1 1 1 1 1 0 1 1 1 1 1 6 0 5 1 0 1 1 0 1 1 0 1 0 1 1 1 3 1 4 1 0 1 1 0 1 1 0 1 1 1 1 1 3 1 4 1 1 1 1 0 1 1 0 1 0 1 1 1 4 0 2 1 1 1 1 0 1 1 0 1 1 1 1 1 3 1 4 .....continued on the next page

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TABLE 2 (continued) Genus/species Chaudhripalpus creelae Coleacarus lithops Crossipalpus verticillatae Dolichotetranychus floridanus Krugeria ramosa Larvacarus transitans Lisaepalpus smileyae Macfarlaniella queenslandica Magdalenapalpus strandtmanni Obdulia daadi Obdulia tamaricis Obuloides rajamohani Pentamerismus oregonensis Pentamerismus erythreus Philippipalpus agohoi Phyllotetranychus aegyptium Phytoptipalpus albizziae Phytoptipalpus paradoxus Priscapalpus cherretti Priscapalpus thomissus Priscapalpus macropilis Prolixus corruginus Prolixus forsteri Pseudoleptus arechavaletae Pseudoleptus graminosus Raoiella macfarlanei Raoiella indica Raoiellana allium Tegopalpus conicus Tenuilichus malvae Tenuipalpus capassae Tenuipalpus salicis Tenuipalpus albae Tenuipalpus elegans Tenuipalpus keiensis Tenuipalpus vernoniae Tenuipalpus viticola Tenuipalpus granati Tenuipalpus proteae Tenuipalpus caudatus Terminalichus karachiensis Terminalichus panajiensis Ultratenuipalpus quadrisetosus Ultratenuipalpus meekeri Urigersonus bunyai

Total 9 10 12 9 7 10 9 11 13 11 11 7 12 13 10 13 13 12 8 8 9 8 9 12 13 12 13 12 9 6 7 7 8 8 8 8 8 8 9 10 5 6 9 10 9

C1 C2 C3 D1 D2 D3 E1 E2 E3 F2 F3 H1 H2 BS 1 0 1 1 0 1 1 0 1 0 1 1 1 6 1 1 1 1 0 1 1 0 1 0 1 1 1 1 1 1 1 1 1 1 1 1 1 0 1 1 1 6 1 1 1 1 0 1 0 0 1 0 1 1 1 1 1 0 1 1 0 1 0 0 1 0 0 1 1 5 1 1 1 1 0 1 1 0 1 0 1 1 1 4 1 0 1 1 0 1 1 0 1 0 1 1 1 2 1 1 1 1 0 1 1 1 1 0 1 1 1 5 1 1 1 1 1 1 1 1 1 1 1 1 1 6 1 1 1 1 1 1 1 0 1 0 1 1 1 4 1 1 1 1 1 1 1 0 1 0 1 1 1 5 1 0 1 1 0 1 0 0 1 0 0 1 1 1 1 1 1 1 1 1 1 0 1 1 1 1 1 4 1 1 1 1 1 1 1 1 1 1 1 1 1 4 1 0 1 1 1 1 1 0 1 0 1 1 1 6 1 1 1 1 1 1 1 1 1 1 1 1 1 4 1 1 1 1 1 1 1 1 1 1 1 1 1 4 1 1 1 1 1 1 1 0 1 1 1 1 1 4 1 0 1 1 0 1 0 0 1 0 1 1 1 3 1 0 1 1 0 1 1 0 1 0 0 1 1 3 1 0 1 1 0 1 1 0 1 0 1 1 1 3 1 0 1 0 0 1 1 0 1 0 1 1 1 1 1 0 1 0 0 1 1 1 1 0 1 1 1 1 1 1 1 1 1 1 1 1 1 0 1 1 1 4 1 1 1 1 1 1 1 1 1 1 1 1 1 4 1 1 1 1 1 1 1 1 1 0 1 1 1 5 1 1 1 1 1 1 1 1 1 1 1 1 1 5 1 1 1 1 1 1 1 1 1 0 1 1 1 4 1 0 1 1 0 1 1 0 1 0 1 1 1 6 0 0 1 0 0 1 0 0 1 0 1 1 1 4 1 0 1 0 0 1 0 0 1 0 1 1 1 2 1 0 1 0 0 1 0 0 1 0 1 1 1 2 0 0 1 1 0 1 1 0 1 0 1 1 1 2 1 0 1 0 0 1 0 0 1 1 1 1 1 2 1 0 1 1 0 1 1 0 1 0 1 1 1 2 0 0 1 1 0 1 1 0 1 0 1 1 1 2 1 0 1 0 0 1 0 0 1 1 1 1 1 2 1 0 1 0 0 1 0 0 1 0 1 1 1 2 1 0 1 1 0 1 1 0 1 0 1 1 1 2 1 0 1 1 0 1 1 0 1 1 1 1 1 2 0 0 1 0 0 1 0 0 1 0 0 1 1 3 1 0 1 0 0 1 0 0 1 0 0 1 1 3 1 0 1 1 0 1 1 0 1 0 1 1 1 3 1 0 1 1 0 1 1 0 1 1 1 1 1 2 1 0 1 1 0 1 1 0 1 0 1 1 1 3

RS 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 1 1 1 0 0 1 1 0 0 0 0 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1

Ps 5 2 5 3 1 2 1 4 5 1 1 2 5 5 5 2 5 5 2 3 2 3 3 5 5 2 2 2 2 3 3 2 1 1 2 2 2 3 2 2 3 3 4 4 4

Total = total number of opisthosomal setae; BS = body shape (see descriptions further in this paper); Rs = rostral shield (0=absent; 1=present); PS = number of palpal segments.

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Body shapes 1. Dolichotetranychine—body long and slender, at least 1.6 x longer than wide, parallel-sided with posterior margin of propodosoma as wide as anterior margin of hysterosoma; several rows of transverse striations present above the C-series of setae; d3 and sometimes e3 setae located in submedial region; e1 setae absent in most species; rostral shield absent; palpus 2–3 segmented. Genera: Acaricis, Afronychus, Coleacarus, Dolichotetranychus, Obuloides, Prolixus, Tegopalpus. 2. Tenuipalpine—body with propodosoma distinctly wider than hysterosoma, anterior margin of hysterosoma abruptly narrowing; rostral shield present; palpus 1–3 and h2 setae long and flagellate except in Ultratenuipalpus and Urigersonus which have 4-segmented palpi and h2 not long andf lagellate). Genera: Lisaepalpus, Tenuipalpus (most species), Ultratenuipalpus, Urigersonus. 3. Brevipalpine—body with posterior margin of propodosoma as wide as anterior margin of hysterosoma, the latter tapering slightly posteriorly; exteneded membraneous rostral shield present; ventral plate usually present; palpus 3-4 segmented; dorsosublateral setae absent (except c2 present in Cenopalpus). Genera: Amblypalpus; Brevipalpus, Cenopalpus, Priscapalpus, Tenuipalpus (few species), Terminalichus. 4. Aegyptobine—similar to brevipalpine but with hysterosoma more globular, not tapering gradually towards the posterior apex; membraneous rostral shield absent; palpus 1–5 segmented; dorsosublateral setae present (except in Tenuilichus). Genera: Aegyptobia, Capedulia,; Obdulia, Pentamerismus, Phyllotetranychus, Phytoptipalpus, Pseudoleptus, Raoiellana, Tenuilichus. 5. Macfarlanielline—body round or nearly so, hysterosoma as wide or wider than propodosoma; palpus 2-3 segmented; rostral shield absent. Genera: Krugeria, Macfarlaniella, Raoiella. 6. Tegopalpine—propodosoma hoodlike, covering the gnathosoma. a) anterior margin of propodosoma deeply incised. Genera: Australopalpus, Magdalenapalpus, Philippipalpus, Tegopalpus. b) anterior margin of propodosoma rounded. Genera: Crossipalpus, Chaudhripalpus.

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KEY TO THE WORLD GENERA OF TENUIPALPIDAE (BASED ON ADULT FEMALES) 1a. 1b.

Penultimate dorsal setae (h2) long and whip-like (Fig. 3A), at least 4x longer than other dorsal opisthosomal setae (Fig. 1), dorsum of opisthosoma with 6–10 pairs of setae, setae c2 absent ................................................ 2 Penultimate dorsal setae (h2) not long and attenuate, but similar in length to other opisthosomal setae (Fig. 3B); c2 present or absent; dorsum of opisthosoma with 5–13 pairs of setae ....................................................... 6

Figure 3. A). Tenuipalpus tetrazygiae female dorsum (after Pritchard & Baker 1958). B) Brevipalpus colpodes female (after Pritchard & Baker 1958). 2a (1a).

Dorsal opisthosoma with 6 pairs of setae, c1, d1 and e1 absent; idiosoma oval-shaped (Fig. 4); 2 species, Oriental (India) ............................................................................................. Tenuilichus Mohanasundaram 1988

FIGURE 4. Tenuilichus malvae female dorsum, palp and venter (after Mohanasundaram 1988).

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2b.

Dorsal opisthosoma with 7–10 pairs of setae, at least one pair of either c1, d1 or e1 present; most species with idiosoma very broad, abruptly narrowing at anterior margin of opisthosoma, the latter, usually slender ......... 3

3a (2b).

Idiosoma elongate and slender, parallel-sided, at least 4x longer than wide; prodorsal sclerite absent; opisthosoma with 8–9 pairs of dorsal setae; genital setae (g1-2) removed from posterior margin and widely separated, with g1 located far anterior to g2; palp 3-segmented; 2 species, Australia on sedge (Fig. 5) .............. ...................................................................................................................... Prolixus Beard, Fan & Walter 2006 Idiosoma less than 2.75x as long as wide, most species with prodorsum very broad, abruptly narrowing at anterior margin of opisthosoma, the latter, usually slender; few species with idiosoma oval-shaped; prodorsal sclerite present; opisthosoma with 7–10 pairs of dorsal setae; g1 and g2 located at the posterior margin of the genital plate and proximal ................................................................................................................................... 4

3b.

FIGURE 5. Prolixus forsteri female dorsum and venter (after Beard et al. 2006).

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4a (3b). 4b.

Palp 1-segmented; femora of legs I and II each with 2–3 setae; sc2 setae minute, much smaller than the v2 setae; 1 species, Australia (Fig. 6)................................................................Lisaepalpus Smiley & Gerson 1995 Palp usually 2 to 4- segmented (rarely 1-segmented); femora of leg I with 4 setae, femora of leg II with 4 (rarely 3) setae; sc2 setae not minute and longer than the v2 setae ....................................................................... 5

FIGURE 6. Lisaepalpus smileyae female dorsum and venter (after Smiley & Gerson 1995). 5a (4b).

Palp 2 or 3- segmented (Fig. 2C, G, K, L), rarely 1 segmented; c1 setae present in most species; 300 species, worldwide (Fig. 7) .................................................................................................. Tenuipalpus Donnadieu 1875

FIGURE 7. Tenuipalpus bucidus female dorsum, palp and venter (after Pritchard & Baker 1958).

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5b.

Palp 4-segmented; c1 setae absent; 2 species, Australia (Fig. 8) ..........................Acaricis Beard & Gerson 2009

FIGURE 8. Acaricis plana female dorsum and venter (after Beard & Gerson 2009).

6a (1b) 6b.

Setae c2 absent (Fig. 9–20) .................................................................................................................................. 7 Setae c2 present (Fig. 21–38)............................................................................................................................. 19

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7a (6a)

Opisthosoma with 7 pairs of long and ramose (tree-like) dorsal setae inserted on tubercles; h2 inserted on a plate together with setae e3; palp 1-segmented; 1 pair of anal setae; body round; 1 species, South Africa (Fig. 9) ........................................................................................................................................ Krugeria Meyer 1979

FIGURE 9. Krugeria ramosa female dorsum, tarsus, venter and leg I (after Meyer 1979).

7b.

Dorsal opisthosoma variable but not long and ramose (tree-like) and inserted on tubercles; h2 not inserted on a plate together with setae e3; palp segments and number of anal setae variable ................................................. 8

8a (7b)

Propodosoma with a rostral shield of varying development (Fig. 14) (idiosoma of most species not elongate, elliptical or slender) ............................................................................................................................................ 9 Propodosoma without a rostral shield (idiosoma of most species somewhat elongate, elliptical or slender) . 15

8b. 9a (8a)

9b.

Idiosoma widest at posterior margin of propodosoma, narrowing abruptly at base of opisthosoma, then expanding posteriorly; some of the dorsal opisthosomal setae broadly palmate; palp 4-segmented; venter with or without plates in genito-anal area. ................................................................................................................ 10 Idiosoma usually oval-shaped, with opisthosoma tapering slightly posteriorly; dorsal opisthosomal setae usually not broadly palmate; palp 1 to 4-segmented; venter with plates in genito-anal .................................. 12

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10a (9a) Setae sc2 and f3 very long and whiplike, more than 10 times longer than other dorsal opisthosomal setae; palpus 4-segmented, with solenidion of palptarsus inserted at base of the segment; 1 species, Australia (Fig. 10).............................................................................................. Urigersonus Mesa, Ochoa & Evans, new genus

FIGURE 10. Urigersonus bunyai female dorsum and palp (after Smiley & Gerson 1995). 10b.

Setae sc2 and f3 not long and whiplike, similar in length and shape to other dorsal opisthosomal setae; palpus usually 4-segmented (rarely 3); solenidion on palptarsus inserted at the apex of the segment ............................ .............................................................................................Ultratenuipalpus sensu lato Mitrofanov 1973 .... 11

11a (10b) Venter without plates in genito-anal area, so that oopore has wavy walls, similar to that in Tetranychidae; 20+ species, Nearctic, Neotropical, Afrotropical, Australian, Pacific Islands (Fig. 11) ............................................ ..............................................................................................................................Ultratenuipalpus sensu stricto

FIGURE 11. Ultratenuipalpus meekeri female dorsum and venter (after Baker & Tuttle 1987).

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11b.

Venter with plates in genito-anal area, so that walls of oopore are not wavy (Fig. 12) ........................................ .................................................................................................................. Ultratenuipalpus quadrisetosus group

FIGURE 12. Ultratenuipalpus quadrisetosus female dorsum, palpus and venter (after Meyer 1979). 12a (9b) Opisthosoma with 5–6 pairs of dorsal setae; dorsocentral setae absent (except c1 present in T. panajiensis); sc1 extremely long, much longer than sc2 in known species; palp 3-segmented; 6 species, India and Pakistan (Fig. 13).........................................................................................................Terminalichus Anwarullah & Khan 1973

FIGURE 13. Terminalichus karachiensis dorsum, venter and palp (after Anwarullah & Khan 1973). 12b.

Opisthosoma with 7–10 (rarely 11) pairs of dorsal setae; at least one pair of dorsocentral setae present; sc1 not extremely long and more or less subequal to sc2 .............................................................................................. 13

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13a (12b) Opisthosoma with 7–8 pairs of short and setiform dorsal setae (c1 present or absent); ventral plate without defined limits; palp 3-segmented; 2 species, Western Palearctic (Fig. 14) ........................................................... .................... ....................................................................................Amblypalpus Mitrofanov & Strunkova 1978 13b. Opisthosoma with 8–10 pairs of dorsal setae, variable in length and shape (c1 present); ventral plate with defined margins or fused with genital plate to form a single distinct plate (Fig. 14, 15); palp 2 to 4-segmented ............................................................................................................................................................................14

FIGURE 14. A. narsikulovi female dorsum, palp and venter (after Mitrofanov & Strunkova 1978). 14a (13b) Genital and ventral plates separated, the former elliptical, wider than long, and the latter rectangular, usually with a striate cuticle forming a frame around the plate; palpus 4-segmented (Fig. 2J) (rarely 3 segmented) (Fig. 2C; lateral margin of opisthoma with 6 or 7 pairs of setae (i.e. f2 present or absent); sc1 , c1 and e1 not extremely long and lanceolate; 275+ species, worldwide (Fig. 15) ...................... Brevipalpus Donnadieu 1875

FIGURE 15. Brevipalpus phoenicis female dorsum, venter and deutonymph (after Baker & Tuttle 1987).

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14b.

Genital and ventral plates fused to form a single, enlarged plate; palpus 2 or 3-segmented; lateral margin of opisthosoma with 5–6 pairs of setae; v2, sc1 , c1 and e1 extremely long and lanceolate in known species; 3 species; Neotropical, Afrotropical (Fig. 16)............................................................. Priscapalpus DeLeon 1961

FIGURE 16. Priscapalpus macropilis female dorsum and venter (after De Leon 1961). 15a (8b) Prodorsum hoodlike (Fig. 17–19), completely covering the gnathosoma with anterior margin of prodorsum deeply incised or rounded; opisthosoma with 9–10 pairs of dorsal short, spatulate or palmate dorsal setae ............................................................................................................................................................................16 15b. Prodorsum not hoodlike or completely covering the gnathosoma with anterior margin of prodorsum deeply incised (Fig. 20–21); opisthosoma with 7–11 pairs of dorsal setae of variable length and shape .................... 18 16a (15a). Palp 2-segmented (Fig. 2H); opisthosoma with 9 pairs of dorsal setae; 1 species, Australia (Fig. 17) ............... .................................................................................................................................Tegopalpus Womersley 1940

FIGURE 17. Tegopalpus conicus female dorsum and venter (after Smiley et al. 1996).

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16b.

Palp 5-segmented; opisthosoma with 9–10 pairs of dorsal setae ...................................................................... 16

17a (16b) Anterior margin of prodorsum deeply incised; setae d2 present (10 pairs of dorsal opisthosomal setae); 1 species, Philippines (Fig. 18) ........................................................................ Philippipalpus Corpuz-Raros 1978

FIGURE 18. Philippipalpus agohoi female dorsum, venter and palp (after Smiley et al. 1996). 17b.

Anterior margin of prodorsum rounded; setae d2 absent (9 pairs of dorsal opisthosomal setae); 1 species, Australia (Fig. 19) .........................................................Chaudhripalpus Mesa, Welbourn & Evans, new genus

FIGURE 19. Chaudhripalpus creelae female dorsum (after Smiley et al. 1996).

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18a (15b). Opisthosoma with 7 pairs of dorsal setae; e1 absent; all dorsal setae very long, several longer than half the width the body, and serrate; palp 1-segmented; c2 absent; 2 species, India and Philippines (Fig. 20) ................ .............................................................................................................................Obuloides Baker & Tuttle 1975

FIGURE 20. Obuloides rajamohani female dorsum and venter (after Baker & Tuttle 1975).

18b.

Opisthosoma with 9–11 pairs of dorsal setae; e1 present; dorsal setae not as long, none longer than half the width of the body; palp 3-segmented (Fig. 19B); setae c2 absent (except in A. mulibrinus); 4 species, South Africa) (Fig. 21) ................................................................................................Afronychus Meyer 1979 (in part)

FIGURE 21. A–C) Afronychus amnicus female A) dorsum, B) palps and C) venter (after Meyer1979).

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19a (6b) Opisthosoma with 11 or fewer pairs of dorsal setae; d2 and e2 absent [e2 present in Macfarlaniella (Australia) in sublateral or lateral position]............................................................................................................................. 20 19b. Opisthosoma with 12–13 pairs of dorsal setae; d2 and/or e2 present..................... ............................................27 20a (19a) Genital and ventral plates distinct and separated, the former elliptical, wider than long, and the latter rectangular, usually with a striate cuticle forming a frame around the plate; palpus 4-segmented (rarely 3segmented) (Fig. 2F); idiosoma broadly oval or slightly tapering posteriorly; 50+ species, worldwide (Fig. 22) . ....................................................................................................................Cenopalpus Pritchard & Baker 1958

FIGURE 22. Cenopalpus capensis female dorsum and venter (after Meyer 1979).

20b.

Genital and ventral plates lacking, incomplete or fused, ventral plate not rectangular and surrounded by a frame (Fig. 22, 26, 28); palpus 1–5 segmented (rarely 4-segmented in Ultratenuipalpus and Macfarlaniella); idiosoma shape variable; with 0–3 pairs of dorsocentralsetae (i.e. c1 , d1 ,and/or e1) ........................................ 21

21a (20b) Idiosoma elongate, more than 1.5 x as long as wide, slender and parallel-sided, tapering posteriorly; d3 and e3 usually inserted in the submedial region; rostral shield absent; opisthosomal setae usually setiform or serrate, not ramose or lanceolate nor longer than half the width of the idiosoma ........................................................ 22 21b. Idiosoma elliptical, oval or round, less than 1.5 x as long as wide, not parallel sided; d3 and e3 inserted along the lateral margin; rostral shield present or absent; dorsal opisthosomal setae ramose or lanceolate, very long, at least half the width of the idiosoma............................................................................................................... 23

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22a (21a) Posterior end of idiosoma with 4 pairs of dorsal setae (e2 present, f2 absent); genu IV without setae; 20 species, worldwide (Figs. 2N, 23) ...................................................................................Dolichotetranychus Sayed 1938

FIGURE 23. Dolichotetranychus floridanus female dorsum, palp and venter (after Baker 1956). 22b.

Posterior end of idiosoma with 5 pairs of dorsal setae (e2 present, f2 absent); genu IV with 1 seta; 1 species (A. mulibrinus) with c2 present; South Africa (Fig. 24) ..........................................Afronychus Meyer 1979 (in part)

FIGURE 24. Afronychus mulibrinus female dorsum (after Meyer 1979).

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23a (21b) With 3 pairs of legs, found in galls on Ziziphus; 1 species, Oriental (India) (Figs. 2P, 25).................................. ..................................................................................................................... Larvacarus Baker & Pritchard 1952

FIGURE 25. Larvacarus transitans female dorsum and venter (after Baker & Pritchard 1952).

23b.

With 4 pairs of legs; usually not known to occur in galls (except Capedulia prasadi on Aspalathus and Obdulia tamaricis on Tamaricis)....................................................................................................................... 24

24a (23b) Idiosoma roundish, with 11 pairs of dorsal setae, of which 8 pairs laterad of the dorsocentral setae; e2 present and d2 present or absent ..................................................................................................................................... 25 24b. Idiosoma elongate elliptical, with 10 pairs of dorsal setae, of which 7 pairs laterad of the dorsocentral setae; d2 and e2 absent ...................................................................................................................................................... 26 25a (24a) Palp 4-segmented; e2 absent and d2 present; 1 species, Australia on Eucalyptus (Fig. 26) .................................. ................................................................................................................Macfarlaniella Baker & Pritchard 1962

FIGURE 26. Macfarlaniella queenlandica female dorsum and venter (after Baker & Pritchard 1962).

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25b.

Palp 1-segmented; d2 present and e2 absent; 2 species, Oriental, Western Palearctic in galls or on leaves of Tamarix (Fig. 27) ...............................................................................................Obdulia Pritchard & Baker 1958

FIGURE 27. Obdulia tamaricis female dorsum (original), venter and palps (after Pritchard & Baker 1958). 26a (24b) Paired claws and empodium short and padlike; each claw with 2 short inner tenent hairs and a long outer hair (see tarsus I in Fig. 28); empodium with 2 rows of short tenent hairs; 4 species, Afrotropical (South Africa) (Fig. 28) ............................................................................................................................ Capedulia Meyer 1979

FIGURE 28. Capedulia calendulae female dorsum, venter and tarsus (after Meyer 1979). 26b.

Paired claws and empodium relatively long; each claw with 2 similar rows of hairs; 1 species, Africa (South Africa) (Fig. 29) ............................................................................................................. Coleacarus Meyer 1979

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FIGURE 29. Coleacarus lithops female dorsum, gnathosoma, venter and tarsus I (after Meyer 1979). 27a (19b) Eyes absent; with 3 pairs of legs; opisthosoma with 12 pairs of dorsal setae (f2 absent); palp 2-segmented; 1 species; Turkey (Fig. 30) ................................................................................... Raoiellana Baker & Tuttle 1972

FIGURE 30. Raoiellana allium female dorsum and venter (after Baker & Tuttle 1972). 27b.

Eyes present; with 4 pairs of legs (some Phytoptipalpus with 3 pairs); opisthosoma with 12–13 pairs of dorsal setae; palp 1 to 5-segmented ............................................................................................................................. 28

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28a (27b) Dorsal opisthosomal setae long and rod-like, with clavate tips; opisthosoma with 13 pairs of dorsal setae; idiosoma round; palpus 2-segmented; 12 species, Neotropical (introduced), western Palearctic, Afrotropical, Oriental, Australasian (Figs. 2D, 31) ................................................................................... Raoiella Hirst 1924

FIGURE 31. Raoiella indica female and male (after Pritchard & Baker 1958). 28b.

Dorsal opisthosomal setae setiform, lanceolate or palmate; opisthosoma with 12–13 pairs of dorsal setae; shape of idiosoma variable; palpus 2 or 5-segmented....................................................................................... 29

29a (28b) Rostral shield present, horn-like, with acute apices; ventral plate absent; idiosoma somewhat elongate and slender, all dorsal opisthosomal setae relatively short and setiform; palpus 4 or 5-segmented (Fig. 2B, 2M); 10 species, worldwide except Australian and Pacific Island regions (Fig. 32).............. Pseudoleptus Bruyant 1911

FIGURE 32. Pseudoleptus arechavaletae female dorsum and venter (after Pritchard & Baker 1958).

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MESA ET AL.

29b.

Rostral shield absent or, if present, not horn-like; ventral plate present or absent, when present, more or less quadrate; idiosoma oval; dorsal opisthosomal setae relatively long and lanceolate or palmate; palpus 2 or 5segmented.......................................................................................................................................................... 30

30a (29b) Prodorsum hoodlike, completely covering the gnathosoma, with anterior margin rounded or deeply incised medially; prodorsal shield absent; only known from Australia ........................................................................ 31 30b. Prodorsum not hoodlike, not completely covering the gnathosoma, with anterior margin rounded or incised; prodorsal shield present or absent; not only known from Australia.................................................................. 33 31a (30a) Opisthosomal dorsal setae short and broadly lanceolate or palmate; f2 absent; anterior margin of prodorsum rounded; 2 species, Australia (Fig. 33)............................................. Crossipalpus Smiley, Frost & Gerson 1996

FIGURE 33. Crossipalpus verticillatae female dorsum, venter of gnathosoma and palpi (after Smiley et al. 1996).

31b.

Opisthosomal dorsal setae long and lanceolate dorsal setae; f2 present or absent; anterior margin of prodorsum rounded or deeply incised medially................................................................................................................... 32

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32a (31b) Anterior margin of prodorsum rounded, slightly pointed at apex; bases of setae d1 close together, with the distance between them corresponding to less than half the distance between bases of setae c1; genital and ventral plate setae barbed; 1 species, Australia (Fig. 34)........................Australopalpus Smiley & Gerson 1995

FIGURE 34. Australopalpus alphitoniae female dorsum and venter (after Smiley & Gerson 1995). 32b.

Anterior margin of prodorsum deeply incised medially; bases of setae d1 not close together, with the distance between about the same as the distance between setae c1; genital and ventral plate setae smooth; 1 species, Australia (Fig. 35) ....................................................... Magdalenapalpus Mesa, Welbourn & Evans, new genus

FIGURE 35. Magdalenapalpus strandtmanni female dorsum (after Smiley et al. 1996).

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MESA ET AL.

33a (30b) Palp 2-segmented (Fig. 2O); opisthosoma with 13 pairs of dorsal setae, each one broadly palmate, with extensive venation; 2 species, western Palearctic (Egypt), Afrotropical (Mauritius) (Fig. 36) ............................ .............................................................................................................................. Phyllotetranychus Sayed 1938

A

B

FIGURE 36. A) Phyllotetranychus aegyptium female dorsum (after Sayed 1938); B) Phyllotetranychus romaine female dorsum and palp (after Pritchard & Baker 1958). 33b.

Palp 5-segmented; opisthosoma with 12–13 pairs of variously shaped dorsal setae which, if broadly palmate, then not with extensive venation ....................................................................................................................... 34

34a (33b) Opisthosoma with 12 (rarely 13 when e2 is present) pairs of dorsal setae; 2 or 3 pairs of setae inserted in the dorsosublateral position (c2, d2, e2); if with 13 pairs of setae, then e2 and f2 inserted on the lateral margin, thus appearing to have 8 pairs of dorsolateral setae; with 3 pairs of anal setae; 6 species, Holarctic, Afrotropical (Fig. 2E, 37) .......................................................................................................Pentamerismus McGregor 1949

FIGURE 37. Pentamerismus erythreus female dorsum and venter (after Baker & Tuttle 1987).

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129

34b.

Opisthosoma with 13 (rarely 12, setae e2 present or absent) pairs of dorsal setae; 4 pairs of ... setae inserted in the dorsosublateral position (c2, d2, e2, f2); e2 and f2 in dorsosublateral position, not inserted on lateral margin, thus with 6 pairs of setae in dorsolateral position; with 2–3 pairs of anal setae .............................................. 35

35a (34b) With 2 pairs of anal setae; some species with 3 pairs of legs; 15 species, Neotropical, Afrotropical, Oriental, Eastern Palearctic (Fig. 2I, 38)............................................................................. Phytoptipalpus Tragardh 1904

Figure 38. Phytoptipalpus conostegiae female dorsum and venter (after Baker & Tuttle 1987). 35b.

With 3 pairs of anal setae; all species with 4 pairs of legs; 94 species, worldwide except Neotropical region (Fig. 2A, 39) .................................................................................................................... Aegyptobia Sayed 1950

FIGURE 39. Aegyptobia tragardhi female dorsum and venter (after Sayed 1950).

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APPENDIX A. Zoogeographic Distribution of Species of Tenuipalpidae (host plant family and country from which the type specimen(s) were described given as an abbreviation — see page 10 for guide to regions and abbreviations). species danutae plana Total = 2 species abuzabiensis acacia aletes aliartensis allionia alpinensis ambrosiae antenostoma aplopappi arbutusae arenaria arizonensis baccharis baptus beglarovi bibbyi cactaceae cactorum campsis cassiae cedermontana changi crotonae cupressus curtipilis daneshvari delfinadae desertorum edenvillensis ephedrae eremia ericae eriogonum euphratica exarata fallugia flourensia forma franseriae glyptus hamus haplopappus hefeiensis hymenocleae

NA

0

NT

0

WPA

0

AF EPA Acaricis

0 0 Aegyptobia

O

AU

0

AUS AUS 2

UAE AZ CA GRE MEX CA MEX CA AZ MEX JAP AZ AZ CA UKR AZ MEX MEX AZ AZ SAf TAI AZ CA MEX IRN AUS AZ SAf AZ ISR SAf AZ IRQ UKR AZ AZ PAK AZ MEX PAK AZ CHN AZ

A catalog of the Tenuipalpidae of the world with a key to genera

PI

Host Family Cyperaceae Cyperaceae

0 Chenopodiaceae Mimosaceae Cupressaceae Cupressaceae Nyctaginaceae Viscaceae Asteraceae Rosaceae Asteraceae Ericaceae Asteraceae Chenopodiaceae Asteraceae Asteraceae Cupressaceae Chenopodiaceae Cactaceae Cactaceae Bignoniaceae Caesalpineaceae Rubiaceae Cupressaceae Euphorbiaceae Cupressaceae Lamiaceae Salicaceae Casuarinaceae Chenopodiaceae Poaceae Ephedraceae Chenopodiaceae Ericaceae Polygonaceae Cupressaceae Asteraceae Rosaceae Asteraceae Asteraceae Asteraceae Cactaceae Boraginaceae Asteraceae Cupressaceae Asteraceae

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131

species incarnata karystensis kerkyrae kharazii lacida leiahensis lineati macswaini mccormicki menoni mesopotamiensis meyerae monacanthae monitus montana nasicornensis neobaptus nina nomus nothus nummulus odontopilis parcus pavlovskii pennatulae perscia physalis pomaderrisae populus prolixa pseudoleptoides pyramidi salicicola salixi salubris sayedi semper sohanraensis solanum thujae torreyi tragardhi ueckermanni vannus villiensis wainsteini xerophilus xinjiangensis zaitzevi Total = 93 species

NA

NT MEX

WPA

AF

EPA

O

AU

PI

GRE GRE IRN AZ PAK SAf CA WA IND IRQ GRE UAE PAK OR SAf SAf CA ND OK

+ PAK SAf PAK GEO MEX IRN

AZ NZ GRE SAf UT EGY IRQ EGY PAK EGY PAK PAK MEX AZ AZ EGY IRN MEX GRE ARM GEO CHN 32

11

GEO 26

amnicus cliffortiae

132 · Zootaxa 2098 © 2009 Magnolia Press

8 4 Afronychus SAf SAf

10

1

Host Family Nyctaginaceae Cupressaceae Unidentified Cupressaceae Asteraceae Boraginaceae Santalaceae Asteraceae Cupressaceae Cupressaceae Cupressaceae Unidentified Chenopodiaceae Fabaceae Scrophulariaceae Scrophulariaceae Salicaceae Cupressaceae Poaceae Cupressaceae Cupressaceae Scrophuliaceae Anacardiaceae Ephedraceae Mimosaceae Plantaginaceae Solanaceae Rhamnaceae Salicaceae Poaceae Poaceae desert plant Salicaceae Salicaceae Cupressaceae Cupressaceae Unidentified plant Unidentified plant Solanaceae Cupressaceae Poaceae Cupressaceae Cupressaceae Mimosaceae/ Rosaceae Cupressaceae Caryophyllaceae Chenopodiaceae Polygonaceae

1 Unidentified Rosaceae

MESA ET AL.

species diastellae mulibrinus Total = 4 species

NA

NT

WPA

0

0

0

masakii narsikulovi Total = 2 species

0

0

TAJ 1

alphitoniae Total = 1 species

0

0

0

abiesae absens acatlanus acutangliae adelos aeoloides aeolus aepi albus allenrolfeae alni alternatus ambrosiae amecensis andrewsi apalos araucanus arcus ardesiae arizonicae artemesiae arystis asterae asthenes atalantae baccharis bakeri bauhiniae beeri bicolpus borealis bouchea bucerasae bumeliae butcheri californicus cardinalis cassia castillejae ceanothus celtis cercidium

AF EPA SAf SAf 4 0 Amblypalpus SAf

O

AU

PI

0

0

0 Bruniaceae Asteraceae

1 0 Australopalpus

0

0

0

0 0 Brevipalpus

0

AUS 1

0

Rhamnaceae

MEX PRi MEX GUY PAK AZ CA MEX MEX AZ MEX MEX MEX MEX HON PAK CHL FL MEX MEX AZ PAK AZ CA GRE MEX PRi HON CA MD UKR MEX PRi FL FL CA AZ

+

+

+

MEX MEX AZ MEX MEX

A catalog of the Tenuipalpidae of the world with a key to genera

+

Host Family Proteaceae Poaceae

+

+

+

Pinaceae Melastomataceae Salicaceae Lauraceae Malvaceae Asteraceae Asteraceae Asteracaeae Fagaceae Chenopodiaceae Betulaceae Combretaceae Asteraceae Pinaceae Solanaceae Caesalpiniaceae Philesciaceae Asteraceae Myrsinaceae Fagaceae Asteraceae Anacardiaceae Asteraceae Hydrophyllaceae Oleaceae Asteraceae Meliaceae Caesalpiniaceae Asteraceae Caricaceae Pinaceae Verbenaceae Combretaceae Sapotaceae Amaranthaceae Rutaceae Oleaceae Caesalpiniaceae Scrophulariaceae Rhamnaceae Ulmaceae Caesalpiniaceae

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133

species chalkidicus chamaedorea chilensis chucumayi clypealis cnidosculos cochlospermi coffeae coldenia colens colpodes columbiensis combreti conocarpi cordiae cortesi crataegus creber cromroyi crotonellae crotoni cucurbitae cuneatus daqingis delus dentatae desmodium dilaccus dipholisi disparis docimas dosis edax edurensis ehsanii edwinae emarginatae encelia encinarius ennsi erectus ericae essigi euphorbiae ewpristori favus fenghuangis filifer filifolia fleschneri floridianus formosus frankenia

NA

NT

WPA GRE

AF

EPA

O

MEX CHL MEX PAK MEX MEX ETH MEX PAK FL MO MEX FL MEX CHL MEX PAK HON MEX MEX IND ITA CHN CA MEX MEX CA FL PAK NC PAK MEX NY PAK MEX MEX MEX MEX MO MEX SAf CA IND MEX PAK CHN MEX MEX HON FL MEX MEX

134 · Zootaxa 2098 © 2009 Magnolia Press

AU

PI

Host Family Oleaceae Arecaceae Rutaceae Stryracaceae Punicaceae Euphorbiaceae Cochlospermaceae Rubiaceae Boraginaceae Anacardiaceae Asteraceae Platanaceae Combretaceae Combretaceae Boraginaceae Escalloniaceae Rosaceae Tiliaceae Moraceae Euphorbiaceae Euphorbiaceae Cucurbitaceae hedge plant Verbenaceae Cupressaceae Boraginaceae Fabaceae Rosaceae Sapotaceae Solanaceae Juglandaceae Oleaceae Boraginaceae Tiliaceae Solanaceae Asteraceae Capparaceae Asteraceae Fagaceae Asteraceae Rubiaceae Ericaceae Cornaceae Euphorbiaceae Asteraceae Celastraceae Caesalpiniaceae Lauraceae Asteraceae Lauraceae Lauraceae Lauraceae Frankeniaceae

MESA ET AL.

species franxini galliprodiens garmani gauhatiensis germanium gliricidiae glomeratus glymma gortiniensis grandis grewiae guihuansis hafizii hainanensis hamelrectae hamzaii hashmii hellenicus hernandiae homalus hondurani huananis hybus hypti incarnatae inermis insinuatus ipomoeae iraquiensis jambhiri janeae johnstoni jordani juncus junicus juniperus khalidae karackiensis kasuriensis keiferi lagasceae lantanae lepidium levis lewisi lilium linki lippiae lochmius longisetosus lotus lupinus lyallpuriensis

NA GA

NT

WPA

AF

EPA

O

AU

CRi CT IND MEX MEX VA FL GRE OR IND CHN PAK CHN MEX PAK PAK GRE JAM CA HON CHN FL MEX MEX PAK MEX AZ IRQ GA

+ MEX JOR PAK CHN MEX PAK PAK PAK

CA MEX MEX MEX MEX CA WA FL

+

+

+ +

MEX CA PRi MEX MEX

A catalog of the Tenuipalpidae of the world with a key to genera

PAK

PI

Host Family Oleaceae Cochlospermaceae Caprifoliaceae Oleaceae Geraniaceae Fabaceae Fagaceae Verbenaceae Oleaceae Pinaceae Tiliaceae Oleaceae Meliaceae Unidentified plant Rubiaceae Asteraceae Cucurbitaceae Oleaceae Oleaceae Asteraceae Melastomataceae Caesalpiniaceae Unidentified weed Lamiaceae Nyctaginaceae Cucurbitaceae Fagaceae Convolvulaceae Euphorbiaceae Rutaceae Scrophulariaceae Arecaceae Rutaceae Lamiaceae Rutaceae Cupressaceae Aizoaceae Convolvulaceae Unidentified plant Asteraceae Asteraceae Verbenaceae Brassicaceae Boraginaceae Rutaceae Liliaceae Fagaceae Verbenaceae Unidentified plant Unidentified plant Fabaceae Fabaceae Malvaceae

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135

species lysilomae macedonicus mallorquensis mcgregori melichrus mexicanus mingoraesnsis mitrofanovi moreliensis mori mumai nangalensis nannus neoardisiae neobicolpus neohyptis neoreligiosae nocivus nodiflorae oaxacensis obovatus obovoides ochaliensis ochoai ocoteae ogcotus ogmellus ogmus oleae olearius oleasteri olivicola oncidii oreopanacis ornantis ornatus orobos ortizi pachucensis parthenium patulus perseae persicanus phoenicis physali pini piniceltis pinicola piniwaltheriae plexus pluchea pocillator porca

NA FL

NT

WPA

AF

EPA

O

AU

PI

+

+

+

+

GRE SPA CA CA MEX PAK ITA MEX MEX AZ IND CA MEX MEX MEX MEX PAK

+ FL

MEX MEX +

FRA

+

+

+ PAK

HON FL CA FL CA

+ MOR EGY GRE ITA ENG

+ MEX

PAK MEX PAK CRi MEX AZ PAK MEX + CA

+ MEX + MEX

NET

+

+

PAK +

FL MEX PAK MEX MEX UT

136 · Zootaxa 2098 © 2009 Magnolia Press

Host Family Fabaceae Oleaceae Rosaceae Rutaceae Aceraceae Cupressaceae Unidentified plant Fagaceae Fagaceae Moraceae Rosaceae Oleaceae Ericaceae Asteraceae Fabaceae Lamiaceae Pinaceae Boraginaceae Verbenaceae Fagaceae Phytolaccaceae Oleaceae Acanthaceae Asteraceae Lauraceae Hydrophyllaceae Fagaceae Fagaceae Oleaceae Oleaceae Oleaceae Oleaceae Orchidaceae Araliaceae Unidentified plant Chenopodiaceae Asteraceae Arecaceae Fabaceae Asteraceae Solanaceae Lauraceae Rosaceae Arecaceae Asteraceae Pinaceae Ulmaceae Pinaceae Sterculiaceae Moraceae Asteraceae Asteraceae Loranthaceae

MESA ET AL.

species portalis portheo potentillae pritchardi proboscidius psephoides pseudoleptoides pseudolilium pseudophoenicis pseudopini pseudopinicolus pseudostriatus psilostropheae psychotriae punicans pycnanthemi qianniunis quercicolus querensis rapii recki recula religiosae rhanis rica rostratus rotai rubus ruelliae rugosus rugulosus russulus sahii salasi salicis salix salignae salpizo salviae salviella sayedi selas serratus shafii sida similis simlensis sipho solidus spaticus spatulatus spitzeri spurcus

NA AZ

NT +

WPA

AF

EPA

O PAK

AZ AZ MEX CA MEX CUB MEX MEX MEX CRi AZ FL CA NC CHN MEX MEX PAK UKR PAK MEX PAK PAK MEX ITA MEX MEX MEX PAK FRA PAK CRi AZ MEX SAf PAK CA CA FL CA MEX PAK MEX MEX TRI PAK PAK GRE MEX MEX

A catalog of the Tenuipalpidae of the world with a key to genera

PAK

AU

PI

Host Family Asteraceae Cupressaceae Rosaceae Euphorbiaceae Asteraceae Garryaceae Sapindaceae Euphorbiaceae Malvaceae Fagaceae Cupressaceae Fabaceae Asteraceae Rubiaceae Asteraceae Lamiaceae Convolvulaceae Fagaceae Fagaceae Brassicaceae Fagaceae rat-ka-raja Pinaceae Asteraceae Asteraceae Myricaceae Oleaceae Rosaceae Acanthaceae Fagaceae Unidentified plant Cactaceae Solanaceae Anacardiaceae Salicaceae Salicaceae Buddlejaceae Asteraceae Lamiaceae Lamiaceae Juglandaceae Rosaceae Unidentified tree Solanaceae Malvaceae Pinaceae Convolvulaceae Cucurbitaceae Unidentified plant Oleaceae Asteraceae Cactaceae Lamiaceae

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137

species stenolobae stipae striatus styxus tagetinae tarus tepicbutilonae tepicensis testudinalis theae thelycraniae tiliae tinsukiaensis tiwanensis trinidadensis tuberellus turrialbensis urtriculus variolatus ventus venutus verbenae viguiera xystus zempoalensis zhenzhumeis ziaii zinniae Total = 282 species

calendulae martima prasadi xeroclei Total = 4 species abaii aratus arbuti bagdasariani bakeri brachypalpus capacis capensis carpini chitraliensis crataegi creticus dignus eriobotryi evini favosus

NA

NT MEX MEX MEX

WPA

AF

EPA

O

AU

PI

0 +4

0 +3

FL MEX CHL MEX MEX MEX CHN UKR TN IND PAK TRI MEX CRi PAK MEX PAK MEX MEX MEX FL MEX CHN PAK 69 +2

MEX 129 +4

23 +1

3 9 +3 +4 Capedulia SAf

49 +3

Asteraceae Plumbaginaceae Fabaceae Aizoaceae

ISR

0

0

1

SAf SAf 3 0 Cenopalpus

0

IRN PAK GRE TAJ TUR GRE PAK SAf UKR PAK IRN CRE PAK GRE IRN

138 · Zootaxa 2098 © 2009 Magnolia Press

Host Family Asteraceae Poaceae mata gusano Solanaceae Asteraceae Unidentified plant Malvaceae Lamiaceae Unidentified tree Theaceae Cornaceae Tiliaceae Lamiaceae Unidentified plant Verbenaceae Lauraceae Unidentified plant Malvaceae Malvaceae Unidentified plant Asteraceae Verbenaceae Asteraceae Fabaceae Pinaceae Rosaceae Malvaceae Asteraceae

PAK

0

0 Fagaceae Solanaceae Ericaceae Oleaceae Rosaceae Rosaceae Pinaceae Asteraceae Betulaceae Rosaceae Rosaceae Lamiaceae Musaceae Rosaceae Salicaceae Pinaceae

MESA ET AL.

species halperini haqii hederae homalos irani japonicus kritos lanceolatisetae limbatus lineola longirostris mespili meyerae mughalii musai natalensis naupakticus officinalis oleunus orakiensis pegazzanoae pennatisetis picitilis piger pistaciae platani populi pritchardi pseudospinosus pterinus pulcher quadricornis ramus rubber saryabiensis scoopsetus spinosus sunniensis tamarixi taygeticus viniferus virgulatus wainsteini xini Total = 60 species creelae Total =1 species lithops Total = 1 species

NA

NT

WPA ISR

AF

EPA

O

AU

PI

17

0

0

0

AUS 1

0

PAK GRE PAK IRN PAK PAK EGY PAK ITA UKR UKR IRN PAK LEB SAf GRE GRE SAf PAK ITA KAZ PAK KAZ GRE GEO GEO TUR UKR SPA ITA ARM

+

PAK TAJ PAK CRE FRA PAK EGY GRE GRE PAK UKR +1

0

39

0

0

0

0

0

0

CHN 3 1 Chaudhripalpus 0 0 Coleacarus NAM 1 0 Crossipalpus

muellerianae

A catalog of the Tenuipalpidae of the world with a key to genera

Host Family Pinaceae Rosaceae Araliaceae Rosaceae Rutaceae Rosaceae Rosaceae Rosaceae Unidentified plant Pinaceae Fagaceae Rosaceae Platanaceae Cucurbitaceae Rosaceae Scrophulariaceae Rosaceae Lamiaceae Oleaceae Rosaceae Pinaceae Salicaceae Rosaceae Unidentified plant Anacardiaceae Platanaceae Salicaceae Rosaceae Rosaceae Rosaceae Unidentified plant Rosaceae Pinaceae Unidentified plant Pinaceae Ericaceae Primulaceae Unidentified plant Tamaricaceae Rosaceae Vitaceae Unidentified plant Pinaceae Salicaceae

Casuarinaceae

Aizoaceae 0

0 AUS

0 Casuarinaceae

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139

species verticillatae Total = 2 species alpinus ancistrus apaches australianus babylonicus bakeri carnea cocos cracens cymosum elatariae floridanus gramineus macer micidus muhlenbergia repenae salinas summersi tenellae vandergooti wiesei zoysiae Total = 23 species

NA

NT

WPA

0

0

0

0 0 Dolichotetranychus

PI

NM SRI OK SAf IND FL

+ TAJ SAf

CO AZ IND CA CA IND JAV SAf

0

0

0

transitans Total = 1 species

0

0

0

0

0

0

queenslandica Total = 1 species

0

0

0

strandtmanni Total = 1 species

0

0

0

0

IRQ ISR 2

0

0

0 MEX MEX

140 · Zootaxa 2098 © 2009 Magnolia Press

0

JAP 3 1 Krugeria SAf 1 0 Larvacarus 0 0 Lisaepalpus 0 0 Macfarlaniella 0 0 Magdalenapalpus 0 0 Obdulia

Host Family Casuarinaceae

0

AUS

ramosa Total = 1 species

abnormis arbutusae

0

AU AUS 2

IRQ EGY

3

rajamohani rimandoi Total = 2 species

O

FL

+1

daadi tamaricis Total = 2 species

EPA

NZ NZ

8

smileyae Total = 1 species

AF

5

1

2

0

0

0

IND 1

0

0

0

AUS 1

0

AUS 1

0

0

AUS 1

0

Asteraceae Scrophulariaceae Bromeliaceae Poaceae Poaceae Unidentified plant Poaceae Arecaceae Poaceae Asteraceae Zingiberaceae Bromeliaceae Unidentified cereal Poaceae Poaceae Poaceae Poaceae Poaceae Poaceae Poaceae Orchidaceae Poaceae Poaceae

Tiliaceae

Rhamnaceae gall maker Rhamnaceae

0 Myrtaceae

Casuarinaceae

Tamaricaceae Tamaricaceae 0 0 Obuloides

0 0 Pentamerismus

0 IND PHI 2

0

0 Malvaceae Tiliaceae

0

0 Cupressaceae Ericaceae

MESA ET AL.

species canadensis collinus coronatus erythreus foetidae foliisetis judiciarius juniperi kunmingensis oregonensis ossianensis retusus shandongensis tauricus taxi Total = 17 species

NA CAN

AF

EPA

O

AU

PI

0

0

0

PHI 1

0

0

SAf IO

+ SAf UKR

NC GEO CHN OR NY SAf CHN

aegyptium romaine Total = 2 species

5

2+1

UKR SWI 5

0

0

0

3 2 Philippipalpus 0 0 Phyllotetranychus

0

0

1

Arecaceae Arecaceae

MAU 1 0 Phytoptipalpus

0

0

0

EGY

Mimosaceae Mimosaceae Bignoniaceae Bombaceae Caesalpinaceae Melastomataceae Mimosaceae Liliaceae Rosaceae Solanaceae Anacardiaceae Mimosaceae Mimosaceae Mimosaceae Mimosaceae Rosaceae Buxaceae

IND NAM MEX MEX MEX SAf SAf CHN PAK PAK SAf SAf SAf SUD TAJ 0

cherretii macropilis thomissus Total = 3 species

3

2

CHN 7 2 Priscapalpus

3

0

0

GUY MEX 0

0

2

0

0

0

Melastomataceae Sapotaceae Sapotaceae

SAf 1 0 Prolixus

0 0 Pseudoleptus

URA

A catalog of the Tenuipalpidae of the world with a key to genera

Host Family Cupressaceae Thymelaeceae Cupressaceae Cupressaceae Zygophyllaceae Cupressaceae Cupressaceae Cupressaceae Cupressaceae Cupressaceae Cupressaceae Zygophyllaceae Ulmaceae Cistaceae Taxaceae

Casuarinaceae

EGY

aegyptetrapodus albizziae alexandriae ceibae cercidium conostegiae harveyi kalahariensis lanzhouensis lepsis lithos multistriatum muukuorum nyala paradoxus rosae xianensis Total = 17 species

arechavaletae

WPA

ITA

agohoi Total = 1 species

corruginus forsteri Total = 2 species

NT

0

0

0 AUS AUS 2

0 Cyperaceae Cyperaceae 0 Poaceae

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141

species bouteloua graminicola graminosus hilaria palustria panicum peshawariensis tridens zelihae Total = 10 species

NA AZ

NT

WPA

O

AU

PI

0

0

SAf AZ CA AZ PAK AZ 5

1

TUR 2

+1

+1

1

allium Total = 1 species

0

0

TUR 1

conicus Total = 1 species

0

0

0

aboharensis abutiloni acaciae acacii acritus acuminatae aethiopicus africanus albae alpinus amatikulensis amygdalusae anacardii angolensis annonae anoplomexus anoplus antipodus

EPA

UKR

australica camur empedos eugenia indica macfarlanei neotericus obelias pandanae phoenica rahii shimapana Total = 12 species

malvae striatus Total = 2 species

AF

1 0 Raoiella

1

AUS

+

+

+

Myrtaceae Arecaceae Arecaceae Myrtaceae Arecaceae Oleaceae Arecaceae Arecaceae Pandanaceae Arecaceae Arecaceae Celastraceae

PAK PAK IND IND

GRE PAK PAK IND SUD PAK

0

0

0

SAf 2+1 0 Raoielllana

8

1

0

0 0 Tegopalpus

0

0

0

0 0 Tenuilichus

0

AUS 1

0

Alliaceae

0 0 Tenuipalpus

IND IND 2

Casuarinaceae

Malvaceae Unidentified tree 0

0

IND SAf SAf IND SAf IND SAf SAf ANG NZ SAf IND GUY ANG

FL

MEX MEX +

142 · Zootaxa 2098 © 2009 Magnolia Press

Host Family Poaceae Unidentified plant Poaceae Poaceae Poaceae Poaceae Unidentified plant Poaceae Poaceae

NZ

Punicaceae Malvaceae Mimosaceae Mimosaceae Urticacaeae Menispermaceae Mimosaceae Caesalpiniaceae Asteracaeae mat plants Asteraceae Rosaceae Anacardiaceae Euphorbiaceae Annonaceae Meliaceae Meliaceae Violaceae

MESA ET AL.

species arbuti argus ariauae athrixiae attiahi aurantiacus auriculatae austrocedri baeri bagdadensis bakerdeleonorum bakeri banahawensis banksiae barticanus bassiae bellulus berkheyae boninensis boyani bucidae bursereae calcarius caledonicus capparis capassae carlosflechtmanni carolinensis caudatus cedrelae celtidis chamaedorea cheladzeae chelinus chiclorum chinariensis chiococcae cissampelosa citus clematidos coccolobicoloides coccolobicolus coimbatorensis comatus combreti comptus costarricensis couroupita coyacus crassulus crassus crescentiae crocopontensis

NA

NT

WPA UKR

AF

EPA

O

AU

FL BRA SAf CON CHN SAf CHL GEO IRQ FL

HON + PHI AUS GUY IND SAf SAf JAP GUY

FL MEX SAf SAf SAf SAf BRA SC FRA MEX FL CRi UKR SAf MEX PAK FL IND PAK CHN JAM FL IND SAf SAf SAf CRi TRI MEX HOL FRA MEX SAf

A catalog of the Tenuipalpidae of the world with a key to genera

PI

Host Family Ericaceae Agavaceae Vochysiaceae Asteraceae Podocarpaceae Violaceae Plumbaginaceae Cupressaceae Mimosaceae Mimosaceae Asteraceae Magnoliaceae Myrsinaceae Proteaceae shrub Chenopodiaceae Mimosaceae Asteraceae Rubiaceae Sapotaceae Combretaceae Burseraceae Asteraceae Fabaceae Capparaceae Fabaceae Vochysiaceae Asteraceae Caprifoliaceae Meliaceae Ulmaceae Arecaceae Taxaceae Fabaceae Sapotaceae Punicaceae Rubiaceae Menispermaceae Salicaceae Ranunculaceae Rubiaceae Polygonaceae Unidentified plant Rhamnaceae Combretaceae Celastraceae Meliaceae Lecythidaceae Arecaceae Crassulaceae Euphorbiaceae Bignoniaceae Ebenaceae

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143

species cupressoides cyatheae daneshvari danxianensis dasples decus dimensus disparilis dominguensis dombeyae dubinini dumus elegans elongatus emeticae englbrechti ephedrae erasus eremitus eriophyoides eucleae eugeniae euonymi falcatus faresianus faveolus feliciae ferosus fici filicicola flacourtiae flechtmanni frondosus ghaii galpiniae garciniae gatoomensis geigeriae granati grevilleae guamensis guettardae gumbolimbonis guptai haripuriensis hastaligni heteropyxis heveae hondurensis hornotinus hurae ilocanus imias

NA

NT

WPA ISR

AF

EPA

O

AU

PI NZ

IRN CHN FL PAK PAK CHN DRe SAf GEO SAf NZ SAf SAf SAf UKR PAK PAK +

IRQ SAf JAM IRN SAf

IND SAf SAf PAK IND CHN MAL BRA PRi IND SAf CAM ZIM SAf EGY AUS TAH FL FL IND PAK FL SAf BRA HON PAK TRI PHI CUB

144 · Zootaxa 2098 © 2009 Magnolia Press

Host Family Cupressaceae Cyatheaceae Solanaceae Sapotaceae Arecaceae Oleaceae Oleaceae Poaceae Bignoniaceae Sterculiaceae Ephedraceae Combretaceae Asteraceae Podocarpaceae Meliaceae Asteraceae Ephedraceae Salvadoraceae Chenopodiaceae Unidentified plant Ebenaceae Myrtaceae Celastraceae Podocarpaceae Mimosaceae Asteraceae Asteraceae Unidentified plant Moraceae Tthelypteridaceae Flacourtiaceae Myrtaceae Osmundaceae Unidentified plant Mimosaceae Clusiaceae Combretaceae Asteraceae Punicaceae Proteaceae Aspleniaceae Rubiaceae Burseraceae Punicaceae Cannabaceae Lauraceae Heterpyxidaceae Euphorbiaceae Pinaceae Punicaceae Euphorbiaceae Lauraceae Euphorbiaceae

MESA ET AL.

species indicus inophylli insularis isabelae ixorae jagatkhanaens jamaicensis japonicus jasmini jawadii jianfengensis jonkeri jordaani jussiaeae kamalii kapoki karrooi keiensis kenos kesari knorri kobachidzei kraussianae lalbaghensis laminasetae lanceae latiseta lawrencei legatus leipoldti leonarae leucospermi lineosetosus lucumae ludhianaensis lulinicus lunatus lustrabilis lycioides lygodii magalismontani mahoensis malligae mallotae mandraensis mansoni mansoniculus matthyssei melhaniae menglunensis metis metopii micheli

NA

NT

WPA

AF

EPA

O IND

AU

PI NCa

SAf BRA IND IND JAM JAP PAK PAK CHN SAf SAf FL IRN MEX SAf SAf PAK IND ARG GEO SAf IND IND SAf BRA CON PAK SAf SAf SAf CHN MEX IND CHN SAf PAK ZIM JAM SAf NZ IND IND PAK JAM AUS PHI SAf CHN PAK FL SAf

A catalog of the Tenuipalpidae of the world with a key to genera

Host Family Mimosaceae Euphorbiaceae Unidentified tree Euphorbiaceae Rubiaceae Sapotaceae Melastomataceae Ebenaceae Oleaceae Salvadoraceae Unidentified plant Lamiaceae Unidentified plant Onagraceae Oleaceae Bombacaceae Mimosaceae Acanthaceae Rosaceae Brassicaceae Salicaceae Ranunculaceae Mimosaceae Moraceae Unidentified plant Anacardiaceae Celastraceae Unidentified tree Fabaceae Anacardiaceae Mimosaceae Proteaceae Salicaceae Sapotaceae Rosaceae Acanthaceae Mimosaceae Punicaceae Ebenaceae Lygodiaceae Sapotaceae Violaceae Unidentified plant Euphorbiaceae Unidentified plant Bignoniaceae Rhamnaceae Euphorbiaceae Sterculiaceae Unidentified tree Rosaceae Anacardiaceae Ulmaceae

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145

species microphylli mkuziensis molinae montanus mopaneae moraesi morianus mourerae muguanicus mustus myrtus namaensis nenaxi niekerkae nigerianus obvelatus oliveirai ombrensis orchidofilo oribiensis orilloi ortus ovalis oxalis pacificus pagesae pagina palosapis panici papiothalensis pareriophyioides parsii pedrus pernicis persicae philippinensis pieteri pigrus pisinnus placitus platycaryae podocarpi populi portulacae proctori proteae protectus protumidus pruni prunioides pseudocedrelae punicae punjabensis

NA

NT

WPA

AF SAf SAf

EPA

O

AU

PI

HON NZ SAf BRA SAf GUY CHN PAK IRQ SAf SAf SAf NIG CHN BRA SAf BRA SAf PHI PAK SAf BRA PAN PHI PAK PHI PRi SAf ISR IRN HON PAK IND PHI SAf CA PAK PAK CHN SAf IRQ IRN JAM SAf SAf SAf IND SAf CUB

146 · Zootaxa 2098 © 2009 Magnolia Press

PAL

+ IND

Host Family Combretaceae Combretaceae Asteraceae Thymenaeaceae Fabaceae Euphorbiaceae Combretaceae Melastomataceae? Caricaceae Caesalpiniaceae Myrtaceae Anacardiaceae Rubiaceae Icacinaceae Asteraceae Cyperaceae Apocynaceae Mimosaceae 2UFKLGDFHDH Anacardiaceae Rutaceae Moraceae Unidentified shrub Oxalidaceae Orchidaceae Lythraceae Zygophyllaceae Dipterocarpaceae Poaceae Arecaceae Arecaceae Plantaginaceae Unidentified tree Asteraceae Rosaceae Fabaceae Combretaceae Lauraceae Boraginaceae Punicaceae Juglandaceae Podocarpaceae Salicaceae Portulacaceae Bromeliaceae Proteaceae Caesalpiniaceae Asteraceae Rosaceae Combretaceae Meliaceae Punicaceae Apocynaceae

MESA ET AL.

species pyroides pyrusae qingchengensis rangiorae raphiae raptor rarus reticulus rhagicus rhizophorae rhusi rhysus robustae rodionovi rosae rusapensis sagittus salicis sanblasensis sandyi santae sanyaensis scitulus sclerocaryae senecionis sharmai simarubae simplex simplychus smithi solanensis sophiae sparsus spatulatus stativus stefani striolatus tabebuiae taonicus tauricus tectonae tepicanus terminaliae tortulus toowongi tetrazygiae transvaalensis trichiliae trifoliatae trisegmentus trisetosus tuttlei ueckermanni

NA

NT

WPA

AF SAf

EPA

O

AU

PI

IND CHN NZ CAM PAK PAK PAK CA TRI SAf FL SAf AZE UKR ZIM SAf IRQ MEX GUY CRi CHN SAf SAf NZ IND PRi GER PRi SAf IND SAf PAK CHN PAK SAf SAf MEX CHN UKR IND MEX GUY SAf AUS FL SAf JAM IND PAK AZ MEX SAf

A catalog of the Tenuipalpidae of the world with a key to genera

Host Family Anacardiaceae Rosaceae Caprifoliaceae Asteraceae Arecaceae Unidentified plant Zygophyllaceae Vitaceae Ericaceae Rhizophoraceae Anacardiaceae Cyrillaceae Tiliaceae Rosaceae Rosaceae Caesalpiniaceae Restionaceae Salicaceae Solanaceae Humiriaceae Unidentified tree Anacardiaceae Caesalpiniaceae Anacardiaceae Asteraceae Asteraceae Burseraceae Rosaceae Boraginaceae Mimosaceae Punicaceae Mimosaceae Convolvulaceae Asteraceae Amaranthaceae Euphorbiaceae Combretaceae Bignoniaceae Rosaceae Lamiaceae Verbenaceae Ericaceae Combretaceae Monimiaceae Myrtaceae Melastomataceae Capparaceae Meliaceae Unidentified plant Moraceae Agavaceae Arecaceae Caesalpiniaceae

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species umarii unimerus unonopsonis uvae velitor venustus vernoniae vexus victoriae vitexi viticola vridddiriensis waqasii xerocolus xylosmae yarensis yousefi zanthus zeyheri zhengzhouensis zhizhilashviliae zuluensis Total = 306 species delhiensis dehlonensis karachiensis panajiensis psidi serratus Total = 6 species aberrans acharis arboreus asteliae asteliicola baltazarae canelae carpodeti charlini coprosmae coprosmicus cyathodis filicicola hainanensis lacorpuzrarosae meekeri nothofagi pteriphilus quadrisetosus rubi umtataensis

NA

NT

WPA

AF

EPA

O PAK

AU

PI

MEX GUY MEX PAK NZ SAf DRe TRI SAf IRQ IND PAK SAf JAM PAK IND JAM SAf CHN GEO 19

0

64+2

0

29

0

SAf 91 19 Terminalichus

0 0 Ultratenuipalpus

69+2 IND IND PAK IND IND IND 6

4

11 Combretaceae Myrtaceae Combretaceae Combretaceae Myrtaceae Myrtaceae

0

0 NZ

CHL NZ NZ NZ PHI CHL NZ CHL NZ NZ NZ CHN CHN PHI MEX

148 · Zootaxa 2098 © 2009 Magnolia Press

NZ SAf SAf NZ SAf

Host Family Punicaceae Lauraceae Annonaceae Unidentified tree Unidentified plant Cupressaceae Asteraceae Bignoniaceae Unidentified plant Verbenaceae Vitaceae Unidentified plant Amaranthaceae Unidentified herb Flacourtiaceae Arecaceae Myrtaceae Rutaceae Flacourtiaceae Magnoliaceae Ebenaceae Ebenaceae

Rosaceae Proteaceae Phyllocladaceae Asteliaceae Asteliaceae fern Winteraceae Carpotetaceae Aextoxicaceae Rubiaceae Rubiaceae Epacridaceae Thelypteridaceae Lygodiaceae fern fern Fagaceae Dennstaedtiaceae Lauraceae Rosaceae Unidentified plant

MESA ET AL.

species womersleyi younguisti Total = 22 species bunyai Total = 1 species Total Genera = 34 Total species = 891

NA

NT

WPA

0

MEX 5

0

0

0

0

6 138

8 217

14 136

AF

EPA

O

AU AUS

PI

3 2 Urigersonus

2

1

9

0 0 Total 17 8 136 40

0

AUS 1

0

15 180

15 43

5 26

Host Family Myrtaceae Unidentified plant

Araucariaceae

References Ahmad, I. & Akbar, S. (1984) Two new species of the genus Brevipalpus (Acarina: Tenuipalpidae) from Faisalabad. Pakistan Entomologist, 6(1–2), 37–44. Akbar, S. (1990) New species of genus Raoiella (Acarina: Tenuipalpidae) from Pakistan and their phenetic affinities. Pakistan Entomologist, 12(1–2), 75–81. Akbar, S. & Aheer, G.M. (1990) False spider mite (Acarina: Tenuipalpidae) from summer vegetables in Pakistan. Pakistan Journal of Agricultural Science, 27(3), 257–261. Akbar, S. & Aheer, G.M. (1994) Mite fauna of summer vegetables in Punjab. Pakistan Journal of Zoology, 26(4), 339–345. Akbar, S. & Chaudhri, W.M. (1981) New mite species of genus Tenuipalpus (Tenuipalpidae: Acarina) from Pakistan. Pakistan Entomologist, 3(1–2), 1–20. Akbar, S. & Chaudhri, W.M. (1983) Genus Pseudoleptus (Acarina: Tenuipalpidae) from Pakistan. Pakistan Entomologist, 5(1–2), 47–50. Akbar, S. & Chaudhri, W.M. (1985) New species of the genus Cenopalpus (Acari: Tenuipalpidae) from Pakistan and their phenetic affinities. Acarologia, 26(3), 269–287. Akbar, S. & Chaudhri, W.M. (1987) A new species of genus Raoiella (Acarina: Tenuipalpidae) from date palm. Pakistan Entomologist, 9(1–2), 41–44. Akbar, S. & Khalid, S. (1999) New species of the genus Brevipalpus (Acarina: Tenuipalpidae) from Pakistan. Acarologia, 40(2), 155–163. Akbar, S. & Mushtaq, S.M. (1993) Two new species of the genus Aegyptobia (Acarina: Tenuipalpidae) from Pakistan and their phenetic affinities. Pakistan Journal Zoology, 25(2), 153–157. Al-Gboory, I. (1987) Taxonomic studies of false spider mites (Acari: Tenuipalpidae) in central Iraq. 1987. 205 p. Tese (Doctor in Agronomy)—Institut für angewandte Zoologie der Rheinischen Friedrich-Wilhelms—Universitat Bonn, Bonn, 1987. André, M. (1953a) Acariens Phytoptiplapidae parasites des Orchidées, Cactées et plantes grasses cultivées en serres. I. Tenuipalpus orchidarum Parfitt. Bulletin du Muséum d' Histoire Naturelle, Paris. 2o. Série, 25(5), 463–468. André, M. (1953b) Acariens Phytoptiplapidae parasites des Orchidées, Cactées et plantes grasses cultivées en serres (suites). II. Brevipalpus cactorum Oud., et B. confusus Baker. Bulletin du Muséum d' Histoire Naturelle, Paris. 2o. Série, 25(6), 563–571. André, M. (1954) Brevipalpus geisenheyneri (Rübsaamen), Acarien parasite des arbres fruitiers. Bulletin du Muséum d' Histoire Naturelle, Paris. 2o. Série, 26(3), 326–333. André, M. (1958) Contribution to the study of some phytophagous acarina and their predators in Mauritius. Bulletin of Entomological Research, 49(1), 59–75. André, M. (1959) Brevipalpus australis (Tucker), parasite des orangers (Citrus aurantium L.), du Senegal. L'Agronomie Tropicale, 14(2), 239–241. Anwarullah, M. & Khan, B.A. (1973) Terminalichus karachiensis n. gen., n. sp. a new mite from Pakistan (Acarina: Tenuipalpidae). Portugalie Acta Biologica (B), 12(1/4), 153–158. Aranda, B.R. (1969) Contribução para o conhecimento de ácaros de plantas do Paraguay. Universidade de São Paulo-Escola Superior de Agricultra “Luiz de Queiroz -. Piracicaba, Tese (Doutorado), 1–41. Attiah, H.H. (1956) The genus Brevipalpus in Egypt (Acarina: Tenuipalpidae). Bulletin de la Société Entomologique, d'Egypte, 40, 433–477. Bagdasarian, A.T. (1962) Contribution to the fauna false spider mites from Armenia. Izvestiia Akademii Nauk SSR, Moscow, 15, 49–58. Bajwa, W.I., Krantz, G.W. & Kogan, M. (2001) Discovery of Cenopalpus pulcher (C.& F.) (Acari: Tenuipalpidae) in the New World. Proceedings of the Entomological Society of Washington, 103(3), 754–756. Baker, E.W. (1945) Mites of the genus Tenuipalpus (Acarina: Trichadenidae). Proceedings of the Entomological Society of Washington, 47(2), 33–38.

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Baker, E.W. (1948) A new tetranychid mite which further indicates a phylogenetic relationship between the Tetranychidae and Eriophyidae. Proceedings of the Entomological Society of Washington, 50(3), 59–60. Baker, E.W. (1949) The genus Brevipalpus (Acarina: Pseudoleptidae). The American Midland Naturalist, 42(2): 350–402. Baker, E.W. & Pritchard, A.E. (1952a) The geisenheyneri species group of the genus Brevipalpus (Acarina: Phytoptipalpidae). Annals and Magazine of Natural History, Series, 12(5), 609–613. Baker, E.W. & Pritchard, A.E. (1952b) The false spider mite genus Pseudoleptus Bruyant (Acarina: Phytoptipalpidae). The PanPacific Entomologist, 23(2), 112–117. Baker, E.W. & Pritchard, E.A. (1952c) Larvacarus, a new genus of false spider mites (Acarina, Phytoptipalpidae). Proceedings of the Entomological Society of Washington, 54(3), 130–132. Baker, E.W. & Pritchard, A.E. (1953a) The family categories of tetranychoid mites, with a review of the new families Linotetranidae and Tuckerellidae. Annals of the Entomological Society of America, 46(2), 243–257. Baker, E.W. & Pritchard, A.E. (1953b) A review of the false spider mite genus Tenuipalpus Donnadieu (Acarina: Phytoptipalpidae). Annals of the Entomological Society of America, 46(3), 317–336. Baker, E.W. & Pritchard, E. (1953c) A key to mites of the genus Pentamerismus, with descriptions of three new species (Acarina: Phytoptipalpidae). The Wasmann Journal of Biology, 11(3), 353–366. Baker, E.W. & Pritchard, A.E. (1956) False spider mites of the genus Dolichotetranychus (Acarina: Tenuipalpidae). Hilgardia, 24(13), 359–381. Baker, E.W. & Pritchard, A.E. (1960). The tetranychoid mites of Africa. Hilgardia, 29(11), 455–574. Baker, E.W. & Pritchard, A.E. (1962) Macfarlaniella, a new genus of false spider mites (Acarina: Tenuipalpidae). Fieldiana Zoology, 44(15), 123–125. Baker, E.W. & Yin, S.-G. (1988) A catalog of the false spider mites (Tenuipalpidae: Acari) of the United States. International Journal of Acarology, 14(3), 143–155. Baker, E.W. & Tuttle, D.M. (1964) The false spider mite of Arizona (Acarina: Tenuipalpidae). Agricultural Experiment Station, University Arizona, Technical Bulletin, 163, 1–80. Baker, E.W. & Tuttle, D.M. (1972) New species and further notes on the Tetranychoidea mostly from Southwestern United States (Acarina: Tetranychidae and Tenuipalpidae). Smithsonian Contributions to Zoology, 116, 1–36. Baker, E.W. & Tuttle, D.M. (1975) A new genus of Tenuipalpidae (Acarina) from India. USDA Cooperative Economic Institute Report, 25(2), 453–455. Baker, E.W. & Tuttle, D.M. (1987) The false spider mites of Mexico (Tenuipalpidae: Acari). United States Department of Agriculture, Agricultural Research Service, Technical Bulletin, 1706, 1–236. Baker, E.W., Tuttle, D.M. & Abbatiello, M.J. (1975) The false spider mites of northwestern and north central Mexico (Acarina: Tenuipalpidae). Smithsonian Contributions to Zoology, 194, 1–22. Baker, E.W. & Wharton, G.W. (1952) An Introduction to Acarology. The MacMillan Company, New York, 465 pp. Banks, N. (1900) The red spiders mites of the United States (Tetranychus and Stigmaeus). USDA. Division of Entomology Technical Bulletin, Series, 8, 65–67. Banks, N. (1904) Four new species of injurious mites. Journal of the New York Entomological Society, 12, 53–56. Banks, N. (1906) Descriptions of some new mites. Proceedings of the Entomological Society of Washington,. 7, 133–142. Banks, N. (1907) A catalogue of the acarina or mites of the United States. Proceedings of the United States National Museum, 32(1553), 595–625. Banks, N. (1912) New American mites. Proceedings of the Entomological Society of Washington, 14, 96–102. Banks, N. (1915) The Acarina or mites. A review of the group for the use of economic entomologists. United States Department of Agriculture Office of the Secretary. Report, No. 108, 1–153. Beard, J.J., and Gerson, U. (2009) A new flat mite genus, Acaricis (Prostigmata: Tenuipalpidae), from Australian sedges (Cyperaceae). Zootaxa, 2073, 31–44. Beard, J.J., Fan Q.H. & Walter D.E. (2005) A new genus and two new species of Tenuipalpidae (Prostigmata: Tetranychoidea) from an Australian sedge. Acarologia, 45, 161–181. Berlese, A. (1881) Complete Acarological works. Subfamilia Tetranychina. In: Complete Acarological Works, Varia, 1881–1923. L. van der Hammen (ed), 16–19. Berlese, A. (1881) Acari Dannosi alle piante coltivate. In: Complete Acarological Works, Varia, 1881–1923. L. van der Hammen (ed). 2, 142–148. Berlese, A. (1913). Acarotheca Italica. Tipografia di M. Ricci, Firenze, Italy,. 221 p. Blanchard, E.E. (1940) Tres ácaros dañinos para los cultivos argentinos. Revista de la Facultad de Agronomía de La Plata, 24, 11–18. Bodenheimer, F.S. (1930) Die Schädlingsfauna Palästinas. Zeitschrift fur Angewandte Entomologie, 36, 10. Boisduval, A. (1867) Essai sur l’entomologie horticole, France. 648 pp. Bowman, C. E. & Southgate, B. J. (1980) Pentamerismus taxi (Haller) (Acari: Tenuipalpidae) infesting yew in England. Plant Pathology, 29(3), 144–145. Bozai, J. (1970) Tenuipalpus szarvasensis sp. n., a new mite species from Hungary (Acari: Tenuipalpidae). Acta Zoologica Academiae Scientiarum Hungaricae, 16(3–4), 367–369.

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Bruyant, L. 1911. Pseudoleptus arechavaletae n. gen., n. sp., nouvel Acarien Chélétiné de l'Uruguay. Zoologischer Anzeige, 38, 340–345. Bury, R.J. & Krantz, G.W.(1977) A new species of Aegyptobia (Acari: Tenuipalpidae) from a high altitude habitat in Oregon. Acarologia, 19(3), 417–421. Canestrini, G. & Fanzago, F. (1876) Nuovi acari Italiani. Atti Academia Cientifico Veneto, Trentino, Istriana, Pádua, Italy, 5, 130–142 Canestrini, G. (1890) Prospetto dell’Acarofauna Italiana, Familie: Tetranychini. Atti delle adunanze dell’ I.R. Instituto Veneto di Scienze, Lettere ed Arti, Italy. Serie, 6/7, 491–537. Cao, J. Una nueva especie del género Tenuipalpus (Acarina: Tenuipalpidae) sobre Hippomane mancinella. Agrotecnia de Cuba, La Habana, 14(1), 13–20, 1982. Castagnoli, M. (1974) La spermateca in alcune specie di Brevipalpini (Acarina, Tenuipalpidae). Redia, 55, 85–88. Castagnoli, M. (1987) Due nuove specie di Tenuipalpidi delle conifere: Cenopalpus (Cenopalpus) halperini e C. (C.) pegazzanoae (Acarina: Tenuipalpidae). Redia, 70, 105–119. Castagnoli, M. & Pegazzano, F. (1979). Rassegna degli acari Tenuipalpidi dell'Olivo com descrizione di Hystripalpus rotai sp. nov. Redia, 62, 281–297. Cavalcante, R., Almeida e Silva, Q., Pedrosa, F.N., Cavalcante, M.L. S. & Castro, Z. B. (1977) Tenuipalpus anacardii De Leon 1965, em cajueiro, no Estado do Ceará, Brasil. Fitossanidade, Fortaleza, 2(1), 26–27. Chalilova, S.G. (1953) Opisanie devuk novick tetranickovick kleschei ez semeist Bryobiidae I Trichadenidae. Soobshcheniya Akademii Nauk Gruzinskoĭ SSR., Tbilisi, 14(9), 549–551. Chandra, B.K.N. & Channabavanna, G.P. (1974a) Faunistic study of false spider mites (Acarina: Tenuipalpidae) of India. Proceedings of the 4th International Congress of Acarology, Saalfelden, Austria, 174–183. Chandra, B.K.N. & Channabavanna, G.P. (1974b). Raoiella macfarlanei Pritchard & Baker, a new false spider mite from India and its immature stages (Acarina: Tenuipalpidae). Mysore Journal of Agricultural Science, India. 8: 390–395 Channabasavanna, G.P. & Lakkundi, N.H. (1977) Tenuipalpus lalbaghensis (Acarina: Tenuipalpidae), a new species from Southern India. Indian Journal of Acarology, 1, 19–22. Charanasri, V., Saringkapaibul, C. & Kongchuensin, M. (1986). Mites injurious to orchids in Thailand. In: Progress in Acarology (Ed.) Channabasavanna, G.P. & Viraktamath, C.A.. Published VII International Congress of Acarology, Bangalore, India, 9.32, 201–206. Chaudhri, W.M. (1971a) Four new species of the mites of the genus Cenopalpus from Pakistan (Acarina: Tenuipalpidae). Pakistan Journal of Zoology, 3(1), 71–77. Chaudhri, W.M. (1971b) The genus Tenuipalpus in Pakistan—I. Descriptions of five new species (Acarina: Tenuipalpidae). Pakistan Journal of Zoology, 3(2), 203–212. Chaudhri, W.M. (1972a). Five new species of mites of the genus Aegyptobia from Pakistan (Acarina: Tenuipalpidae). Pakistan Journal of Scientific Research, 24(1/2), 18–24. Chaudhri, W.M. 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SPECIES INDEX Names in italics are synonyms aberrans, Ultratenuipalpus abiesae, Brevipalpus abnormis, Pentamerismus aboharensis, Tenuipalpus absens, Brevipalpus abutiloni, Tenuipalpus abuzabiensis, Aegyptobia acacia, Aegyptobia acaciae, Tenuipalpus acacia, Tenuipalpus Acaricis danutae Acaricis plana Acarus, tini acatlanus, Brevipalpus acharis, Ultratenuipalpus acritus, Tenuipalpus acuminatae, Tenuipalpus acutangliae, Brevipalpus adelos, Brevipalpus aegyptetrapodus, Phytoptipalpus aegyptium, Phyllotetranychus Aegyptobia abuzabiensis Aegyptobia acacia Aegyptobia aletes Aegyptobia aliartensis Aegyptobia allionia Aegyptobia alpinensis Aegyptobia ambrosiae Aegyptobia antenostoma Aegyptobia aplopappi Aegyptobia arbutusae Aegyptobia arenaria Aegyptobia arizonensis Aegyptobia baccharis Aegyptobia baptus Aegyptobia beglarovi Aegyptobia bibbyi Aegyptobia cactaceae Aegyptobia cactorum Aegyptobia campsis Aegyptobia cassiae Aegyptobia cedermontana Aegyptobia changi Aegyptobia crotonae Aegyptobia cupressus Aegyptobia curtipilis Aegyptobia daneshvari Aegyptobia delfinadae Aegyptobia desertorum Aegyptobia edenvillensis Aegyptobia ephedrae Aegyptobia eremia

863 101 483 551 102 552 1 2 553 554 890 891 597a 103 864 555 556 104 105 503 501 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31

Aegyptobia ericae Aegyptobia eriogonum Aegyptobia euphratica Aegyptobia exarata Aegyptobia fallugia Aegyptobia flourensia Aegyptobia forma Aegyptobia franseriae Aegyptobia glyptus Aegyptobia hamus Aegyptobia haplopappus Aegyptobia hefeiensis Aegyptobia hymenocleae Aegyptobia incarnata Aegyptobia karystensis Aegyptobia kerkyrae Aegyptobia kharazii Aegyptobia lacida Aegyptobia leiahensis Aegyptobia lineati Aegyptobia macswaini Aegyptobia mccormicki Aegyptobia menoni Aegyptobia mesopotamiensis Aegyptobia meyerae Aegyptobia meyerae Aegyptobia monacanthae Aegyptobia monitus Aegyptobia montana Aegyptobia nasicornensis Aegyptobia neobaptus Aegyptobia nina Aegyptobia nomus Aegyptobia nothus Aegyptobia nummulus Aegyptobia odontopilis Aegyptobia parcus Aegyptobia pavlovskii Aegyptobia pennatulae Aegyptobia perscia Aegyptobia physalis Aegyptobia pomaderrisae Aegyptobia populus Aegyptobia prolixa Aegyptobia pseudoleptoides Aegyptobia pyramidi Aegyptobia salicicola Aegyptobia salixi Aegyptobia salubris Aegyptobia sayedi Aegyptobia semper Aegyptobia semper

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Aegyptobia sohanraensis Aegyptobia solanum Aegyptobia thujae Aegyptobia torreyi Aegyptobia tragardhi Aegyptobia ueckermanni Aegyptobia vannus Aegyptobia villiensis Aegyptobia wainsteini Aegyptobia xerophilus Aegyptobia xerophilus Aegyptobiam xinjiangensis Aegyptobia zaitzevi aeoloides, Brevipalpus aeolus, Brevipalpus aepi, Brevipalpus aethiopicus, Tenuipalpus africanus, Tenuipalpus Afronychus amnicus Afronychus cliffortiae Afronychus diastellae Afronychus mulibrinus agohoi, Philippipalpus albae, Tenuipalpus albizziae, Phytoptipalpus albus, Brevipalpus aletes, Aegyptobia alexandriae, Phytoptipalpus aliartensis, Aegyptobia allenrolfeae, Brevipalpus allionia, Aegyptobia allium, Raoiellana alni, Brevipalpus alphitoniae, Australopalpus alpinensis, Aegyptobia alpinus, Dolichotetranychus alpinus, Tenuipalpus alternatus, Brevipalpus amatikulensis, Tenuipalpus Amblypalpus masakii Amblypalpus narsikulovi ambrosiae, Brevipalpus ambrosiae, Aegyptobia amecensis, Brevipalpus amicus, Brevipalpus amnicus, Afronychus amygdalusae, Tenuipalpus anacardii, Tenuipalpus ancistrus, Dolichotetranychus andrewsi, Brevipalpus angolensis, Tenuipalpus annonae, Tenuipalpus anoplomexus, Tenuipalpus anoplus, Tenuipalpus antenostoma, Aegyptobia

82 83 84 85 86 87 88 89 90 91 91 92 93 106 107 108 557 558 94 95 96 97 500 559 504 109 3 505 4 110 5 547 111 100 6 451 560 112 561 98 99 113 7 114 269a 94 562 563 452 115 564 565 566 567 8

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antipodus, Tenuipalpus apaches, Dolichotetranychus apalos, Brevipalpus aplopappi, Aegyptobia aratus, Cenopalpus araucanus, Brevipalpus arboreus, Ultratenuipalpus arbuti, Cenopalpus arbuti, Tenuipalpus arbutusae, Aegyptobia arbutusae, Pentamerismus arcus, Brevipalpus ardesiae, Brevipalpus arechavaletae, Pseudoleptus arenaria, Aegyptobia argus, Tenuipalpus ariauae, Tenuipalpus arizonensis, Aegyptobia arizonicae, Brevipalpus artemesiae, Brevipalpus arystis, Brevipalpus assamensis, Brevipalpus asteliae, UItratenuipalpus asteliicola, Ultratenuipalpus asterae, Brevipalpus asthenes, Brevipalpus asyntactus, Brevipalpus atalantae, Brevipalpus athrixiae, Tenuipalpus attiahi, Tenuipalpus aurantiacus, Tenuipalpus auriculatae, Tenuipalpus australianus, Dolichotetranychus australica, Raoiella australis, Tenuipalpus Australopalpus alphitoniae austrocedri, Tenuipalpus babylonicus, Dolichotetranychus baccharis, Brevipalpus baccharis, Aegyptobia baeri, Tenuipalpus bagdadensis, Tenuipalpus bagdasariani, Cenopalpus bakerdeleonorum, Tenuipalpus bakeri, Brevipalpus bakeri, Tenuipalpus bakeri , Cenopalpus bakeri , Dolichotetranychus bakeriana, Brevipalpus baltazarae, Ultratenuipalpus banahawensis, Tenuipalpus banksiae, Tenuipalpus baptus, Aegyptobia barticanus, Tenuipalpus bassiae, Tenuipalpus

568 453 116 9 388 117 865 389 569 10 484 118 119 525 11 570 571 12 120 121 122 269b 866 867 123 124 412a 125 572 573 574 575 454 535 136d 100 576 455 126 13 577 578 390 579 127 580 391 456 391a 868 581 582 14 583 584

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bauhiniae, Brevipalpus beeri, Brevipalpus beglarovi, Aegyptobia bellulus, Tenuipalpus berkheyae, Tenuipalpus bibbyi, Aegyptobia bicolpus, Brevipalpus bioculatus, Tenuipalpus bodenheimeri, Tenuipalpus boninensis, Tenuipalpus borealis, Brevipalpus bouchea, Brevipalpus bouteloua, Pseudoleptus boyani, Tenuipalpus brachypalpus, Cenopalpus Brevipalpus abiesae Brevipalpus absens Brevipalpus acatlanus Brevipalpus acutangliae Brevipalpus adelos Brevipalpus aeoloides Brevipalpus aeolus Brevipalpus aepi Brevipalpus albus Brevipalpus allenrolfeae Brevipalpus alni Brevipalpus alternatus Brevipalpus ambrosiae Brevipalpus amecensis Brevipalpus amicus Brevipalpus andrewsi Brevipalpus apalos Brevipalpus araucanus Brevipalpus arcus Brevipalpus ardesiae Brevipalpus arizonicae Brevipalpus artemesiae Brevipalpus arystis Brevipalpus assamensis Brevipalpus asterae Brevipalpus masthenes Brevipalpus asyntactus Brevipalpus atalantae Brevipalpus baccharis Brevipalpus bakeri Brevipalpus bakeriana Brevipalpus bauhiniae Brevipalpus beeri Brevipalpus bicolpus Brevipalpus borealis Brevipalpus bouchea Brevipalpus browningi Brevipalpus bucerasae Brevipalpus bumeliae Brevipalpus butcheri

128 129 15 585 586 16 130 269e 433c 587 131 132 526 588 392 101 102 103 104 105 106 107 108 109 110 111 112 113 114 269a 115 116 117 118 119 120 121 122 269b 123 124 412a 125 126 127 391a 128 129 130 131 132 136a 133 134 135

Brevipalpus californicus Brevipalpus cardinalis Brevipalpus cassia Brevipalpus castillejae Brevipalpus ceanothus Brevipalpus celtis Brevipalpus cercidium Brevipalpus chalkidicus Brevipalpus chamaedorea Brevipalpus chilensis Brevipalpus chucumayi Brevipalpus ciferrii Brevipalpus clypealis Brevipalpus cnidosculos Brevipalpus cochlospermi Brevipalpus coffeae Brevipalpus coldenia Brevipalpus colens Brevipalpus colpodes Brevipalpus columbiensis Brevipalpus combreti Brevipalpus confusus Brevipalpus conocarpi Brevipalpus cordiae Brevipalpus cortesi Brevipalpus crataegus Brevipalpus creber Brevipalpusm cromroyi Brevipalpus crotonellae Brevipalpus crotoni Brevipalpus cucurbitae Brevipalpus cuneatus Brevipalpus daqingis Brevipalpus deleoni Brevipalpus delus Brevipalpus dentatae Brevipalpus desmodium Brevipalpus dilaccus Brevipalpus dipholisi Brevipalpus disparis Brevipalpus docimas Brevipalpus dosis Brevipalpus edax Brevipalpus edurensis Brevipalpus edwinae Brevipalpus ehsanii Brevipalpus emarginatae Brevipalpus encelia Brevipalpus encinarius Brevipalpus ennsi Brevipalpus erectus Brevipalpus ericae Brevipalpus essigi Brevipalpus euphorbiae Brevipalpus ewpristori

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136 137 138 139 140 141 142 143 144 145 146 433a 147 148 149 150 151 152 153 154 155 136b 156 157 158 159 160 161 162 163 164 165 166 292a 167 168 169 170 171 172 173 174 175 176 178 177 179 180 181 182 183 184 185 186 187

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Brevipalpus favus Brevipalpus fenghuangis Brevipalpus filifer Brevipalpus filifolia Brevipalpus fleschneri Brevipalpus floridianus Brevipalpus formosus Brevipalpus frankenia Brevipalpus fraxini Brevipalpus galliprodiens Brevipalpus garmani Brevipalpus gauhatiensis Brevipalpus geranium Brevipalpus gliricidiae Brevipalpus glomeratus Brevipalpus glymma Brevipalpus gortiniensis Brevipalpus grandis Brevipalpus grewiae Brevipalpus guihuanis Brevipalpus hafizii Brevipalpus hafizii Brevipalpus hainanensis Brevipalpus hamelrectae Brevipalpus hamzaii Brevipalpus hashmii Brevipalpus hellenicus Brevipalpus hernandiae Brevipalpus homalus Brevipalpus hondurani Brevipalpus huananis Brevipalpus hybus Brevipalpus hypti Brevipalpus incanum Brevipalpus incarnatae Brevipalpus inermis Brevipalpus insinuatus Brevipalpus ipomoeae Brevipalpus iraquiensis Brevipalpus jambhiri Brevipalpus janeae Brevipalpus johnstoni Brevipalpus jordani Brevipalpus juncus Brevipalpus juncus Brevipalpus junicus Brevipalpus juniperus Brevipalpus kalandadzei Brevipalpus karackiensis Brevipalpus kasuriensis Brevipalpus keiferi Brevipalpus khalidae Brevipalpus lagasceae Brevipalpus lantanae Brevipalpus lepidium

188 189 190 191 192 193 194 195 196 197 198 199 200 201 202 203 204 205 206 207 208a 208 209 210 211 212 213 214 215 216 217 218 219 302a 220 221 222 223 224 225 226 227 228 229a 229 230 231 412b 233 234 235 232 236 237 238

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Brevipalpus levis Brevipalpus lewisi Brevipalpus lilium Brevipalpus linki Brevipalpus lippiae Brevipalpus lochmius Brevipalpus longisetosus Brevipalpus lotus Brevipalpus lupinus Brevipalpus lyallpuriensis Brevipalpus lysilomae Brevipalpus macedonicus Brevipalpus mallorquensis Brevipalpus mcbridei Brevipalpus mcgregori Brevipalpus melichrus Brevipalpus mexicanus Brevipalpus mingoraensis Brevipalpus mitrofanovi Brevipalpus moreliensis Brevipalpus mori Brevipalpus mumai Brevipalpus nangalensis Brevipalpus nannus Brevipalpus neoardisiae Brevipalpus neobicolpus Brevipalpus neohyptis Brevipalpus neoreligiosae Brevipalpus nocivus Brevipalpus nodiflorae Brevipalpus oaxacensis Brevipalpus obovatus Brevipalpus obovoides Brevipalpus ochaliensis Brevipalpus ochoai Brevipalpus ocoteae Brevipalpus ogcotus Brevipalpus ogmellus Brevipalpus ogmus Brevipalpus oleae Brevipalpus olearius Brevipalpus oleasteri Brevipalpus olivicola Brevipalpus oncidii Brevipalpus oreopanacis Brevipalpus origanum Brevipalpus ornantis Brevipalpus ornatus Brevipalpus orobos Brevipalpus ortizi Brevipalpus pachucensis Brevipalpus papayensis Brevipalpus parthenium Brevipalpus patulus Brevipalpus pereger

239 240 241 242 243 244 245 246 247 248 249 250 251 292b 252 253 254 255 256 257 258 259 260 261 262 263 264 265 266 267 268 269 270 271 272 273 274 275 276 277 278 279 280 281 282 269c 283 284 285 286 287 292c 288 289 269d

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Brevipalpus perseae Brevipalpus persicanus Brevipalpus phoenicis Brevipalpus phoenicoides Brevipalpus physali Brevipalpus pini Brevipalpus piniceltis Brevipalpus pinicola Brevipalpus piniwaltheriae Brevipalpus plexus Brevipalpus pluchea Brevipalpus pocillator Brevipalpus porca Brevipalpus portalis Brevipalpus portheo Brevipalpus portheo Brevipalpus potentillae Brevipalpus pritchardi Brevipalpus proboscidius Brevipalpus psephoides Brevipalpus pseudoleptoides Brevipalpus pseudolilium Brevipalpus pseudophoenicis Brevipalpus pseudopini Brevipalpus pseudopinicolus Brevipalpus pseudostriatus Brevipalpus psilostrophe Brevipalpus psilostropheae Brevipalpus psychotriae Brevipalpus punicans Brevipalpus pycnanthemi Brevipalpus pyri Brevipalpus qianniunis Brevipalpus quercicolus Brevipalpus querensis Brevipalpus rapii Brevipalpus recki Brevipalpus recki Brevipalpus recula Brevipalpus religiosae Brevipalpusmrhanis Brevipalpus rica Brevipalpus rostratus Brevipalpus rotai Brevipalpus rubus Brevipalpus ruelliae Brevipalpus rugosus Brevipalpus rugulosus Brevipalpus russulus Brevipalpus sahii Brevipalpus salasi Brevipalpus salicis Brevipalpus salignae Brevipalpus salix Brevipalpus salpizo

290 291 292 292d 293 294 295 296 297 298 299 300 301 302 303a 303 304 305 306 307 308 309 310 311 312 313 359a 314 315 316 317 433b 318 319 320 321 322a 322 323 324 325 326 327 328 329 330 331 332 333 334 335 336 338 337 339

Brevipalpus salviae Brevipalpus salviella Brevipalpus sayedi Brevipalpus selas Brevipalpus serratus Brevipalpus sexflagellatus Brevipalpus shafii Brevipalpus sida Brevipalpusm similis Brevipalpus simlensis Brevipalpus sipho Brevipalpus solanum Brevipalpus solidus Brevipalpus solidus Brevipalpus spaticus Brevipalpus spatulatus Brevipalpus spitzeri Brevipalpus spurcus Brevipalpus stenolobae Brevipalpus stipae Brevipalpus striatus Brevipalpus styxus Brevipalpus tagetinae Brevipalpus tarus Brevipalpus tepicbutilonae Brevipalpus tepicensis Brevipalpus testudinalis Brevipalpus tetraflagellatus Brevipalpus theae Brevipalpus thelycraniae Brevipalpus tiliae Brevipalpus tinsukiaensis Brevipalpus tiwanensis Brevipalpus tlaxcensis Brevipalpus trinidadensis Brevipalpus tuberellus Brevipalpus turrialbensis Brevipalpus utriculus Brevipalpus variolatus Brevipalpus ventus Brevipalpus venutus Brevipalpus verbenae Brevipalpus viguiera Brevipalpus woglumi Brevipalpus xystus Brevipalpus yothersi Brevipalpus zempoalensis Brevipalpus zhenzhumeis Brevipalpus ziaii Brevipalpus zinniae browningi, Brevipalpus bucerasae, Brevipalpus bucidae, Tenuipalpus bumeliae, Brevipalpus bunyai, Ultratenuipalpus

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340 341 342 343 344 258a 345 346 347 348 349 163a 350a 350 351 352 353 354 355 356 357 358 359 360 361 362 363 112a 364 365 366 367 368 294a 369 370 371 372 373 374 375 376 377 136c 378 292e 379 380 381 382 136a 133 589 134 869

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bunyai, Yurigeronus burserae, Tenuipalpus butcheri, Brevipalpus cactaceae, Aegyptobia cactorum, Tenuipalpus cactorum, Aegyptobia calcarius, Tenuipalpus caledonicus, Tenuipalpus calendulae, Capedulia californicus, Brevipalpus Caligonus calyx calyx, Caligonus campsis, Aegyptobia camur, Raoiella camur, Raoiella canadensis, Pentamerismus canelae, Ultratenuipalpus capacis, Cenopalpus capassae, Tenuipalpus Capedulia calendulae Capedulia maritima Capedulia prasadi Capedulia xeroclei capensis, Cenopalpus capparis, Tenuipalpus cardinalis, Brevipalpus carlosflechtmanni, Tenuipalpus carnea, Dolichotetranychus carolinensis, Tenuipalpus carpini, Cenopalpus carpodeti, Ultratenuipalpus cassia, Brevipalpus cassiae, Aegyptobia castillejae, Brevipalpus caudatus, Tenuipalpus ceanothus, Brevipalpus cedermontana, Aegyptobia cedrelae, Tenuipalpus cedrelae, Tenuipalpus ceibae, Phytoptipalpus celtidis, Tenuipalpus celtis, Brevipalpus Cenopalplus xini Cenopalpus abaii Cenopalpus aratus Cenopalpus arbuti Cenopalpus bagdasariani Cenopalpus bakeri Cenopalpus brachypalpus Cenopalpus capacis Cenopalpus capensis Cenopalpus carpini Cenopalpus chitraliensis Cenopalpus crataegi Cenopalpus creticus

890 590 135 17 333a 18 591 592 383 136 597b 597b 19 536a 536 485 870 393 594 383 384 385 386 394 593 137 595 457 596 395 871 138 20 139 597 140 21 598 778a 506 599 141 446 387 388 389 390 391 392 393 394 395 396 397 398

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Cenopalpus dignus Cenopalpus eriobotryi Cenopalpus evini Cenopalpus favosus Cenopalpus fewstrii Cenopalpus halperini Cenopalpus haqii Cenopalpus hederae Cenopalpus homalos Cenopalpus irani Cenopalpus japonicus Cenopalpus kritos Cenopalpus lanceolatisetae Cenopalpus limbatus Cenopalpus lineola Cenopalpus longirostris Cenopalpus mespili Cenopalpus meyerae Cenopalpus mughalii Cenopalpus musai Cenopalpus natalensis Cenopalpus naupakticus Cenopalpus officinalis Cenopalpus oleunus Cenopalpus orakiensis Cenopalpus pegazzanoae Cenopalpus pennatisetis Cenopalpus picitilis Cenopalpus piger Cenopalpus pistaciae Cenopalpus platani Cenopalpus populi Cenopalpus pritchardi Cenopalpus pseudospinosus Cenopalpus pterinus Cenopalpus pulcher Cenopalpus quadricornis Cenopalpus ramus Cenopalpus ruber Cenopalpus saryabiensis Cenopalpus scoopsetus Cenopalpus spinosus Cenopalpus sunniensis Cenopalpus tamarixi Cenopalpus taygeticus Cenopalpus viniferus Cenopalpus virgulatus Cenopalpus wainsteini cercidium, Brevipalpus cercidium, Phytoptipalpus chalkidicus, Brevipalpus chamaedorea, Brevipalpus chamaedorea, Tenuipalpus chamaedoreae, Tenuipalpus change, Aegyptobia

399 400 401 402 445a 403 404 405 406 407 408 409 410 411 412 413 414 415 416 417 418 419 420 421 422 423 424 425 426 427 428 429 430 431 432 433 434 435 436 437 438 439 440 441 442 443 444 445 142 507 143 144 600 833a 22

MESA ET AL.

charlini, Ultratenuipalpus Chaudhripalpus creelae cheladzeae, Tenuipalpus chelinus, Tenuipalpus cherretti, Priscapalpus chiclorum, Tenuipalpus chilensis, Brevipalpus chinariensis, Tenuipalpus chiococcae, Tenuipalpus chitraliensis, Cenopalpus chucumayi, Brevipalpus ciferrii, Brevipalpus cissampelosa, Tenuipalpus citri, Tenuipalpus citus, Tenuipalpus clematidos, Tenuipalpus cliffortiae, Afronychus clypealis, Brevipalpus cnidosculos, Brevipalpus coccolobicoloides, Tenuipalpus coccolobicolus, Tenuipalpus cochlospermi, Brevipalpus cocos, Dolichotetranychus cocosae, Rarosiella coffeae, Brevipalpus coimbatorensis, Tenuipalpus coldenia, Brevipalpus Coleacarus lithops colens, Brevipalpus collinus, Pentamerismus Colopalpus mansoni colpodes, Brevipalpus columbiensis, Brevipalpus comatus, Tenuipalpus combreti, Brevipalpus combreti, Tenuipalpus comptus, Tenuipalpus confuses, Brevipalpus conicus, Tegopalpus conocarpi, Brevipalpus conostegiae , Phytoptipalpus coprosmae, Ultratenuipalpus coprosmicus, Ultratenuipalpus cordiae, Brevipalpus coronatus, Pentamerismus corruginus, Prolixus cortesi, Brevipalpus costarricensis, Tenuipalpus couroupita, Tenuipalpus coyacus, Tenuipalpus cracens, Dolichotetranychus crassulus, Tenuipalpus crassus, Tenuipalpus crataegi, Cenopalpus crataegus, Brevipalpus

872 447 601 602 520 603 145 604 605 396 146 433a 606 636a 607 608 95 147 148 609 610 149 458 539a 150 611 151 448 152 486 721a 153 154 612 155 613 614 136b 548 156 508 873 874 157 487 523 158 615 616 617 459 618 619 397 159

creber, Brevipalpus creelae, Chaudhripalpus crescentiae, Tenuipalpus creticus, Cenopalpus crocopontensis, Tenuipalpus cromroyi, Brevipalpus Crossipalpus, muellerianae Crossipalpus, verticillatae crotonae, Aegyptobia crotonellae, Brevipalpus crotoni, Brevipalpus cucurbitae, Brevipalpus cuneatus, Brevipalpus cupressoides, Tenuipalpus cupressus, Aegyptobia curtipilis, Aegyptobia cyatheae, Tenuipalpus cyathodis, Ultratenuipalpus cymosum, Dolichotetranychus daadi, Obdulia daneshvari, Aegyptobia daneshvari, Tenuipalpus danutae, Acaricis danxianensis, Tenuipalpus daqingis, Brevipalpus dasples, Tenuipalpus decus, Tenuipalpus dehlonensis, Terminalichus deleoni, Brevipalpus delfinadae, Aegyptobia delhiensis, Terminalichus delus, Brevipalpus dentatae, Brevipalpus desertorum, Aegyptobia desmodium, Brevipalpus diastellae, Afronychus dignus, Cenopalpus dilaccus, Brevipalpus dimensus, Tenuipalpus dipholisi, Brevipalpus disparilis, Tenuipalpus disparis, Brevipalpus docimas, Brevipalpus Dolichotetranychus alpinus Dolichotetranychus.ancistrus Dolichotetranychus.apaches Dolichotetranychus.australianus Dolichotetranychus.babylonicus Dolichotetranychus.bakeri Dolichotetranychus.carnea Dolichotetranychus.cocos Dolichotetranychus.cracens Dolichotetranychus.cymosum Dolichotetranychus elatariae Dolichotetranychus floridanus

A catalog of the Tenuipalpidae of the world with a key to genera

160 447 620 398 621 161 449 450 23 162 163 164 165 622 24 25 623 875 460 479 26 624 890 625 166 626 627 858 292a 27 857 167 168 28 169 96 399 170 628 171 629 172 173 451 452 453 454 455 456 457 458 459 460 461 462

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Dolichotetranychus gramineus Dolichotetranychus macer Dolichotetranychus micidus Dolichotetranychus muhlenbergia Dolichotetranychus repenae Dolichotetranychus salinas Dolichotetranychus summersi Dolichotetranychus tenellae Dolichotetranychus vandergooti Dolichotetranychus wiesei Dolichotetranychus zoysiae dombeyae, Tenuipalpus dominguensis, Tenuipalpus dosis, Brevipalpus dubinini, Tenuipalpus dumus, Tenuipalpus edax, Brevipalpus edenvillensis, Aegyptobia edurensis, Brevipalpus edwinae, Brevipalpus ehsanii, Brevipalpus elatariae, Dolichotetranychus elegans, Tenuipalpus elongates, Tenuipalpus emarginatae, Brevipalpus emeticae, Tenuipalpus empedos, Raoiella encelia, Brevipalpus encinarius, Brevipalpus engelbrechti, Tenuipalpus ennsi, Brevipalpus ephedrae, Aegyptobia ephedrae, Tenuipalpus erasus, Tenuipalpus erectus, Brevipalpus eremia, Aegyptobia eremitus, Tenuipalpus ericae, Brevipalpus ericae, Aegyptobia eriobotryi, Cenopalpus eriogonum, Aegyptobia eriophyoids, Tenuipalpus erythreus, Pentamerismus essigi, Brevipalpus eucleae, Tenuipalpus Eugenia, Raoiella eugeniae, Tenuipalpus euonymi, Tenuipalpus euphorbiae, Brevipalpus euphratica, Aegyptobia evini, Cenopalpus ewpristori, Brevipalpus exarata, Aegyptobia falcatus, Tenuipalpus fallugia, Aegyptobia

463 464 465 466 467 468 469 470 471 472 473 631 630 174 632 633 175 29 176 178 177 461 634 635 179 636 537 180 181 637 182 30 638 639 183 31 640 184 32 400 33 641 488 185 642 538 643 644 186 34 401 187 35 645 36

168 · Zootaxa 2098 © 2009 Magnolia Press

faresianus, Tenuipalpus faveolus, Tenuipalpus favosus, Cenopalpus favus, Brevipalpus feliciae, Tenuipalpus fenghuangis, Brevipalpus ferosus, Tenuipalpus fewstrii, Cenopalpus fici, Tenuipalpus filicicola, Tenuipalpus filicicola, Ultratenuipalpus filifer, Brevipalpus filifolia, Brevipalpus flacourtiae, Tenuipalpus flechtmanni, Tenuipalpus fleschneri, Brevipalpus floridanus, Dolichotetranychus floridianus, Brevipalpus flourensia, Aegyptobia foetidae, Pentamerismus foliisetis, Pentamerismus forma, Aegyptobia formosus, Brevipalpus forsteri, Prolixus frankenia, Brevipalpus franseriae, Aegyptobia fraxini, Brevipalpus frondosus, Tenuipalpus galliprodiens, Brevipalpus galpiniae, Tenuipalpus garciniae, Tenuipalpus garmani, Brevipalpus gatoomensis, Tenuipalpus gauhatiensis, Brevipalpus geigeriae, Tenuipalpus geisenheyneri, Tenuipalpus geranium, Brevipalpus ghaii, Tenuipalpus ghanii, Tenuipalpus gibbus, Tenuipalpus glaber, Tenuipalpus gliricidiae, Brevipalpus glomeratus, Brevipalpus glymma, Brevipalpus glyptus, Aegyptobia gonianaensis, Ultratenuipalpus gortiniensis, Brevipalpus gramineus, Dolichotetranychus graminicola, Pseudoleptus graminosus, Pseudoleptus granati, Tenuipalpus grandis, Brevipalpus grevilleae, Tenuipalpus grewiae, Brevipalpus guamensis, Tenuipalpus

646 647 402 188 648 189 649 445a 650 651 876 190 191 652 653 192 462 193 37 489 490 38 194 524 195 39 196 654 197 656 657 198 658 199 659 439a 200 655 628a 881a 439b 201 202 203 40 889 204 463 527 528 660 205 661 206 662

MESA ET AL.

guettardae, Tenuipalpus guihuanis, Brevipalpus gumbolimbonis, Tenuipalpus guptai, Tenuipalpus gurdaspurensis, Priscapalpus hafizii, Brevipalpus hafizii, Brevipalpus hainanensis, Brevipalpus hainanensis, Ultratenuipalpus halperini, Cenopalpus hamelrectae, Brevipalpus hamus, Aegyptobia hamzaii, Brevipalpus haplopappus, Aegyptobia haqii, Cenopalpus haripuriensis, Tenuipalpus harveyi, Phytoptipalpus hashmii, Brevipalpus hastaligni, Tenuipalpus hederae, Cenopalpus hefeiensis, Aegyptobia hellenicus, Brevipalpus hernandiae, Brevipalpus heteropyxis, Tenuipalpus heveae, Tenuipalpus hilaria, Pseudoleptus homalos, Cenopalpus homalus, Brevipalpus hondurani, Brevipalpus hondurensis, Tenuipalpus hornotinus, Tenuipalpus huananis, Brevipalpus hurae, Tenuipalpus hybus, Brevipalpus hymenocleae, Aegyptobia hypti, Brevipalpus ilocanus, Tenuipalpus imias, Tenuipalpus incanum, Brevipalpus incarnate, Aegyptobia incarnatae, Brevipalpus indica, Raoiella indicus, Tenuipalpus inermis, Brevipalpus inophylli, Tenuipalpus inornatus, Tenuipalpus insinuatus, Brevipalpus insularis, Tenuipalpus ipomoeae, Brevipalpus irani, Cenopalpus iraquiensis, Brevipalpus isabelae, Tenuipalpus ixorae, Tenuipalpus jagatkhanaens, Tenuipalpus jamaicensis, Tenuipalpus

663 207 664 665 887 208 208a 209 877 403 210 41 211 42 404 666 509 212 667 405 43 213 214 668 669 529 406 215 216 670 671 217 672 218 44 219 673 674 302a 45 220 539 675 221 676 269f 222 677 223 407 224 678 679 680 681

jambhiri, Brevipalpus janeae, Brevipalpus japonicus, Tenuipalpus japonicus, Cenopalpus jasmine, Tenuipalpus jawadii, Tenuipalpus jianfengensis, Tenuipalpus johnstoni, Brevipalpus jonkeri, Tenuipalpus jordaani, Tenuipalpus jordani, Brevipalpus judiciaries, Pentamerismus juncos, Brevipalpus juncos, Brevipalpus junicus, Brevipalpus juniperi, Pentamerismus juniperus, Brevipalpus jussiaeae, Tenuipalpus kalahariensis, Phytoptipalpus kalandadzei, Brevipalpus kamalii, Tenuipalpus kapoki, Tenuipalpus karachiensis, Terminalichus karackiensis, Brevipalpus karrooi, Tenuipalpus karystensis, Aegyptobia kasuriensis, Brevipalpus keiensis, Tenuipalpus keiferi, Brevipalpus kenos, Tenuipalpus kerkyrae, Aegyptobia kesari, Tenuipalpus khalidae, Brevipalpus kharazii, Aegyptobia knorri, Tenuipalpus kobachidzei, Tenuipalpus kraussianae, Tenuipalpus kritos, Cenopalpus Krugeria ramosa kunmingensis, Pentamerismus lacida, Aegyptobia lacorpuzrarosae, Ultratenuipalpus lagasceae, Brevipalpus lalbaghensis, Tenuipalpus laminasetae, Tenuipalpus lanceae, Tenuipalpus lanceolatisetae, Cenopalpus lantanae, Brevipalpus lanzhouensis, Phytoptipalpus lapidosus, Tenuipalpus Larvacarus transitans latiseta, Tenuipalpus lawrencei, Tenuipalpus legatus, Tenuipalpus leiahensis, Aegyptobia

A catalog of the Tenuipalpidae of the world with a key to genera

225 226 682 408 683 684 685 227 686 687 228 491 229a 229 230 492 231 688 510 412b 689 690 859 233 691 46 234 692 235 693 47 694 232 48 695 696 697 409 474 493 49 878 236 698 699 700 410 237 511 641a 475 701 702 703 50

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leipoldti, Tenuipalpus leonorae, Tenuipalpus lepidium, Brevipalpus lepsis, Phytoptipalpus leucospermi, Tenuipalpus levis, Brevipalpus lewisi, Brevipalpus lilium, Brevipalpus limbatus, Cenopalpus lineate, Aegyptobia lineola, Cenopalpus lineosetosus, Tenuipalpus linki, Brevipalpus lippiae, Brevipalpus Lisaepalpus, smileyae lithops, Coleacarus lithos, Phytoptipalpus lochmius, Brevipalpus longirostris, Cenopalpus longisetosus, Brevipalpus lotus, Brevipalpus lucumae, Tenuipalpus ludhianaensis, Tenuipalpus lulinicus, Tenuipalpus lunatus, Tenuipalpus lupinus, Brevipalpus lustrabilis, Tenuipalpus lyallpuriensis, Brevipalpus lycioides, Tenuipalpus lygodii , Tenuipalpus lysilomae, Brevipalpus macedonicus, Brevipalpus macer, Dolichotetranychus macfarlanei, Raoiella Macfarlaniella queenslandica macropilis, Priscapalpus macswaini, Aegyptobia magalismontani, Tenuipalpus Magdalenapalpus strandtmanni mahoensis, Tenuipalpus malligai, Tenuipalpus mallorquensis, Brevipalpus mallotae, Tenuipalpus malvae, Tenuilichus mandraensis, Tenuipalpus mansoni, Colopalpus mansoni, Tenuipalpus mansoniculus, Tenuipalpus maritime, Capedulia masakii, Amblypalpus matthyssei, Tenuipalpus mcbridei, Brevipalpus mccormicki, Aegyptobia mcgregori, Brevipalpus meekeri, Ultratenuipalpus

704 705 238 512 706 239 240 241 411 51 412 707 242 243 476 448 513 244 413 245 246 708 709 710 711 247 712 248 713 714 249 250 464 540 477 521 52 715 478 716 717 251 718 549 719 721a 720 721 384 98 722 292b 53 252 879

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melhaniae, Tenuipalpus melichrus, Brevipalpus menglunensis, Tenuipalpus menoni, Aegyptobia mesopotamiensis, Aegyptobia mespili, Cenopalpus metis, Tenuipalpus metopii, Tenuipalpus mexicanus, Brevipalpus meyerae, Aegyptobia meyerae, Aegyptobia meyerae, Cenopalpus micheli, Tenuipalpus micidus, Dolichotetranychus microphylli, Tenuipalpus mingoraensis, Brevipalpus mitrofanovi, Brevipalpus mkuziensis, Tenuipalpus molinai, Tenuipalpus monacanthae, Aegyptobia monitus, Aegyptobia Montana, Aegyptobia montanus, Tenuipalpus mopaneae, Tenuipalpus moraesi, Tenuipalpus moreliensis, Brevipalpus mori, Brevipalpus morianus, Tenuipalpus morishitai, Pentamerismus mourerae, Tenuipalpus muellerianae, Crossipalpus mughalii, Cenopalpus muguanicus, Tenuipalpus muhlenbergia, Dolichotetranychus mulibrinus, Afronychus multistriatum, Phytoptipalpus mumai, Brevipalpus musai, Cenopalpus mustus, Tenuipalpus muukuorum, Phytoptipalpus myrtus, Tenuipalpus namaensis, Tenuipalpus nangalensis, Brevipalpus nannus, Brevipalpus narsikulovi, Amblypalpus nasicornensis, Aegyptobia natalensis, Cenopalpus naupakticus, Cenopalpus nenaxi, Tenuipalpus neoardisiae, Brevipalpus neobaptus, Aegyptobia neobicolpus, Brevipalpus neohyptis, Brevipalpus neoreligiosae, Brevipalpus neotericus, Raoiella

723 253 724 54 55 414 725 726 254 48a 56 415 727 465 728 255 256 729 730 57 58 59 731 732 733 257 258 734 499a 735 449 416 736 466 97 514 259 417 737 515 738 739 260 261 99 60 418 419 740 262 61 263 264 265 541a

MESA ET AL.

neotericus, Raoiella niekerkae, Tenuipalpus nigerianus, Tenuipalpus nina, Aegyptobia nocivus, Brevipalpus nodiflorae, Brevipalpus nomus, Aegyptobia nothofagi, Ultratenuipalpus nothus, Aegyptobia nummulus, Aegyptobia nyala, Phytoptipalpus oaxacensis, Brevipalpus Obdulia daadi Obdulia tamaricis obelias, Raoiella obovatus, Brevipalpus obovoides, Brevipalpus Obuloides rajamohani obvelatus, Tenuipalpus ochaliensis, Brevipalpus ochoai, Brevipalpus ocoteae, Brevipalpus odontopilis, Aegyptobia officinalis, Cenopalpus ogcotus, Brevipalpus ogmellus, Brevipalpus ogmus, Brevipalpus oleae, Brevipalpus olearius, Brevipalpus oleasteri, Brevipalpus oleunus, Cenopalpus oliveirai, Tenuipalpus olivicola, Brevipalpus ombrensis, Tenuipalpus oncidii, Brevipalpus orakiensis, Cenopalpus orchidofilo, Tenuipalpus oregonensis, Pentamerismus oreopanacis, Brevipalpus oribiensis, Tenuipalpus origanum, Brevipalpus orilloi, Tenuipalpus ornantis, Brevipalpus ornatus, Brevipalpus orobos, Brevipalpus ortizi, Brevipalpus ortus, Tenuipalpus ossianensis, Pentamerismus oudemansi, Tenuipalpus ovalis, Tenuipalpus oxalis, Tenuipalpus pachucensis, Brevipalpus pacificus, Tenuipalpus pagesae, Tenuipalpus pagina, Tenuipalpus

541 741 742 62 266 267 63 880 64 65 516 268 479 480 542 269 270 481 743 271 272 273 66 420 274 275 276 277 278 279 421 744 280 745 281 422 746 494 282 747 269c 748 283 284 285 286 749 495 433d 750 751 287 752 753 754

palmatus, Tenuipalpus palosapis, Tenuipalpus palustria, Pseudoleptus panajiensis, Terminalichus pandanae, Raoiella panici, Tenuipalpus panicum, Pseudoleptus papayensis, Brevipalpus papiothalensis, Tenuipalpus paradoxus, Phytoptipalpus parcus, Aegyptobia pareriophyiodes, Tenuipalpus parsii, Tenuipalpus parthenium, Brevipalpus patulus, Brevipalpus pavlovskii, Aegyptobia pedrus, Tenuipalpus pegazzanoae, Cenopalpus pennatisetis, Cenopalpus pennatulae, Aegyptobia Pentamerismus abnormis Pentamerismus arbutusae Pentamerismus canadensis Pentamerismus collinus Pentamerismus coronatus Pentamerismus erythreus Pentamerismus foetidae Pentamerismus foliisetis Pentamerismus judiciarius Pentamerismus juniperi Pentamerismus kunmingensis Pentamerismus morishitai Pentamerismus oregonensis Pentamerismus ossianensis Pentamerismus retusus Pentamerismus shandongensis Pentamerismus tauricus Pentamerismus taxi pereger, Brevipalpus pernicis, Tenuipalpus perscia, Aegyptobia perseae, Brevipalpus persicae, Tenuipalpus persicanus, Brevipalpus peshawariensis, Pseudoleptus philippinensis, Tenuipalpus Philippipalpus agohoi phoenica, Raoiella phoenicis, Brevipalpus phoenicoides, Brevipalpus Phyllotetranychus aegyptium Phyllotetranychus romaine physali, Brevipalpus physalis, Aegyptobia Phytoptipalpus aegyptetrapodus

A catalog of the Tenuipalpidae of the world with a key to genera

597c 755 530 860 543 756 531 292c 757 517 67 758 759 288 289 68 760 423 424 69 483 484 485 486 487 488 489 490 491 492 493 499a 494 495 496 497 498 499 269d 761 70 290 762 291 532 763 500 544 292 292d 501 502 293 71 503

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Phytoptipalpus albizziae Phytoptipalpus alexandriae Phytoptipalpus ceibae Phytoptipalpus cercidium Phytoptipalpus conostegiae Phytoptipalpus harveyi Phytoptipalpus kalahariensis Phytoptipalpus lanzhouensis Phytoptipalpus lepsis Phytoptipalpus lithos Phytoptipalpus multistriatum Phytoptipalpus muukuorum Phytoptipalpus nyala Phytoptipalpus paradoxus Phytoptipalpus rosae Phytoptipalpus xianensis piarai, Priscapalpus picitilis, Cenopalpus pieteri, Tenuipalpus piger, Cenopalpus pigrus, Tenuipalpus pini, Brevipalpus piniceltis, Brevipalpus pinicola, Brevipalpus piniwaltheriae, Brevipalpus pisinnus, Tenuipalpus pistaciae, Cenopalpus placitus, Tenuipalpus plana, Acaricis platani, Cenopalpus platycaryae, Tenuipalpus plexus, Brevipalpus pluchea, Brevipalpus pocillator, Brevipalpus podocarpi, Tenuipalpus pomaderrisae, Aegyptobia populi, Tenuipalpus populi, Cenopalpus populus, Aegyptobia porca, Brevipalpus portalis, Brevipalpus portheo, Brevipalpus portheo, Brevipalpus portulacae, Tenuipalpus potentillae, Brevipalpus prasadi, Capedulia Priscapalpus cherretti Priscapalpus gurdaspurensis Priscapalpus macropilis Priscapalpus piarai Priscapalpus thomissus pritchardi, Brevipalpus pritchardi, Cenopalpus proboscidius, Brevipalpus proctori, Tenuipalpus

504 505 506 507 508 509 510 511 512 513 514 515 516 517 518 519 888 425 764 426 765 294 295 296 297 766 427 767 891 428 768 298 299 300 769 72 770 429 73 301 302 303a 303 771 304 385 520 887 521 888 522 305 430 306 772

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prolixa, Aegyptobia Prolixus corruginus Prolixus forsteri proteae, Tenuipalpus protectus, Tenuipalpus protumidus, Tenuipalpus pruni, Tenuipalpus prunioides, Tenuipalpus psephoides, Brevipalpus pseudocedrelae, Tenuipalpus pseudocuneatus, Tenuipalpus pseudoleptoides, Brevipalpus pseudoleptoides, Aegyptobia Pseudoleptus arechavaletae Pseudoleptus bouteloua Pseudoleptus graminicola Pseudoleptus graminosus Pseudoleptus hilaria Pseudoleptus palustria Pseudoleptus panicum Pseudoleptus peshawariensis Pseudoleptus tridens Pseudoleptus zelihae pseudolilium, Brevipalpus pseudophoenicis, Brevipalpus pseudopini, Brevipalpus pseudopinicolus, Brevipalpus pseudospinosus, Cenopalpus pseudostriatus, Brevipalpus psidi, Terminalichus psilostrophe, Brevipalpus psilostropheae, Brevipalpus psychotriae, Brevipalpus pterinus, Cenopalpus pteriphilus, Ultratenuipalpus pulcher, Cenopalpus punicae, Tenuipalpus punicans, Brevipalpus punjabensis, Tenuipalpus pycnanthemi, Brevipalpus pyramidi, Aegyptobia pyri, Brevipalpus pyroides, Tenuipalpus pyrusae, Tenuipalpus qianniunis, Brevipalpus qingchengensis, Tenuipalpus quadricornis, Cenopalpus quadrisetosus, Ultratenuipalpus queenslandica, Macfarlaniella queenslandica, Raoiella quercicolus, Brevipalpus querensis, Brevipalpus rahii, Raoiella rajamohani, Obuloides ramose, Krugeria

74 523 524 773 774 775 776 777 307 778 269g 308 75 525 526 527 528 529 530 531 532 533 534 309 310 311 312 431 313 861 359a 314 315 432 881 433 779 316 780 317 76 433b 781 782 318 783 434 882 477 477 319 320 545 481 474

MESA ET AL.

ramus, Cenopalpus rangiorae, Tenuipalpus Raoiella, australica Raoiella camur Raoiella camur Raoiella empedos Raoiella eugenia Raoiella indica Raoiella macfarlanei Raoiella neotericus Raoiella neotericus Raoiella obelias Raoiella pandanae Raoiella phoenica Raoiella queenslandica Raoiella rahii Raoiella shimapana Raoiellana allium raphiae, Tenuipalpus rapii, Brevipalpus raptor, Tenuipalpus Rarosiella cocosae rarus, Tenuipalpus recki, Brevipalpus recki, Brevipalpus recula, Brevipalpus religiosae, Brevipalpus repenae, Dolichotetranychus reticulus, Tenuipalpus retusus, Pentamerismus rhagicus, Tenuipalpus rhanis, Brevipalpus rhizophorae, Tenuipalpus rhusi, Tenuipalpus rhysus, Tenuipalpus rica, Brevipalpus rillus, Tenuipalpu robustae, Tenuipalpus rodionovi, Tenuipalpus romaine, Phyllotetranychus rosae, Tenuipalpus rosae, Phytoptipalpus rostratus, Brevipalpus rotai, Brevipalpus rubber, Cenopalpus rubi, Ultratenuipalpus rubus, Brevipalpus ruelliae, Brevipalpus rugosus, Brevipalpus rugulosus, Brevipalpus rusapensis, Tenuipalpus russulus, Brevipalpus sagittus, Tenuipalpus sahii, Brevipalpus salasi, Brevipalpus

435 784 535 536a 536 537 538 539 540 541a 541 542 543 544 477 545 546 547 785 321 786 539a 787 322a 322 323 324 467 788 496 789 325 790 791 792 326 620a 793 794 502 795 518 327 328 436 883 329 330 331 332 796 333 797 334 335

salicicola, Aegyptobia salicis, Brevipalpus salicis, Tenuipalpus salignae, Brevipalpus Salinas, Dolichotetranychus salix, Brevipalpus salixi, Aegyptobia salpizo, Brevipalpus salubris, Aegyptobia salviae, Brevipalpus salviella, Brevipalpus sanblasensis, Tenuipalpus sandyi, Tenuipalpus santae, Tenuipalpus sanyaensis, Tenuipalpus saryabiensis, Cenopalpus sayedi, Brevipalpus sayedi, Aegyptobia scitulus, Tenuipalpus sclerocaryae, Tenuipalpus scoopsetus, Cenopalpus selas, Brevipalpus simper, Aegyptobia simper, Aegyptobia senecionis, Tenuipalpus serratus, Brevipalpus serratus, Terminalichus sexflagellatus, Brevipalpus shafii, Brevipalpus shandongensis, Pentamerismus sharmai, Tenuipalpus shimapana, Raoiella sida, Brevipalpus simarubae, Tenuipalpus similes, Brevipalpus simlensis, Brevipalpus simplex, Tenuipalpus simplychus, Tenuipalpus siphon, Brevipalpus smileyae, Lisaepalpus smithi, Tenuipalpus sohanraensis, Aegyptobia solanensis, Tenuipalpus solanum, Brevipalpus solanum, Aegyptobia solidus, Brevipalpus solidus, Brevipalpus sophiae, Tenuipalpus sparsus, Tenuipalpus spathiphyllus, Tenuipalpus spaticus, Brevipalpus spatulatus, Brevipalpus spatulatus, Tenuipalpus spinosus, Cenopalpus spitzeri, Brevipalpus

A catalog of the Tenuipalpidae of the world with a key to genera

77 336 798 338 468 337 78 339 79 340 341 799 800 801 802 437 342 80 803 804 438 343 81a 81 805 344 862 258a 345 497 806 546 346 807 347 348 808 809 349 476 810 82 811 163a 83 350a 350 812 813 748a 351 352 814 439 353

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spurcus, Brevipalpus stativus, Tenuipalpus stefani, Tenuipalpus stenolobae, Brevipalpus stipae, Brevipalpus strandtmanni. Magdalenapalpus striatus, Brevipalpus striatus, Tenuilichus striolatus, Tenuipalpus styxus, Brevipalpus summersi, Dolichotetranychus sunniensis, Cenopalpus swieteniae, Tenuipalpus szarvasensis, Tenuipalpus tabebuiae, Tenuipalpus tagetinae, Brevipalpus tamaricis, Obdulia tamarixi, Cenopalpus taonicus, Tenuipalpus tapirirae, Tenuipalpus tarus, Brevipalpus tauricus, Pentamerismus tauricus, Tenuipalpus taxi, Pentamerismus taygeticus, Cenopalpus tectonae, Tenuipalpus Tegopalpus conicus tenellae, Dolichotetranychus Tenuilichus malvae Tenuilichus striatus Tenuipalpu rillus Tenuipalpus aboharensis Tenuipalpus abutiloni Tenuipalpus acaciae Tenuipalpus acacii Tenuipalpus acritus Tenuipalpus acuminatae Tenuipalpus aethiopicus Tenuipalpus africanus Tenuipalpus albae Tenuipalpus alpinus Tenuipalpus amatikulensis Tenuipalpus amygdalusae Tenuipalpus anacardii Tenuipalpus angolensis Tenuipalpus annonae Tenuipalpus anoplomexus Tenuipalpus anoplus Tenuipalpus antipodus Tenuipalpus arbuti Tenuipalpus argus Tenuipalpus ariauae Tenuipalpus athrixiae Tenuipalpus attiahi Tenuipalpus aurantiacus

354 815 816 355 356 478 357 550 817 358 469 440 567a 601a 818 359 480 441 819 820 360 498 821 499 442 822 548 470 549 550 620a 551 552 553 554 555 556 557 558 559 560 561 562 563 564 565 566 567 568 569 570 571 572 573 574

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Tenuipalpus auriculatae Tenuipalpus australis Tenuipalpus austrocedri Tenuipalpus baeri Tenuipalpus bagdadensis Tenuipalpus bakerdeleonorum Tenuipalpus bakeri Tenuipalpus banahawensis Tenuipalpus banksiae Tenuipalpus barticanus Tenuipalpus bassiae Tenuipalpus bellulus Tenuipalpus berkheyae Tenuipalpus bioculatus Tenuipalpus bodenheimeri Tenuipalpus boninensis Tenuipalpus boyani Tenuipalpus bucidae Tenuipalpus burserae Tenuipalpus cactorum Tenuipalpus calcarius Tenuipalpus caledonicus Tenuipalpus capassae Tenuipalpus capparis Tenuipalpus carlosflechtmanni Tenuipalpus carolinensis Tenuipalpus caudatus Tenuipalpus cedrelae Tenuipalpus cedrelae Tenuipalpus celtidis Tenuipalpus chamaedorea Tenuipalpus chamaedoreae Tenuipalpus cheladzeae Tenuipalpus chelinus Tenuipalpus chiclorum Tenuipalpus chinariensis Tenuipalpus chiococcae Tenuipalpus cissampelosa Tenuipalpus citri Tenuipalpus citus Tenuipalpus clematidos Tenuipalpus coccolobicoloides Tenuipalpus coccolobicolus Tenuipalpus coimbatorensis Tenuipalpus comatus Tenuipalpus combreti Tenuipalpus comptus Tenuipalpus costarricensis Tenuipalpus couroupita Tenuipalpus coyacus Tenuipalpus crassulus Tenuipalpus crassus Tenuipalpus crescentiae Tenuipalpus crocopontensis Tenuipalpus cupressoides

575 136d 576 577 578 579 580 581 582 583 584 585 586 269e 433c 587 588 589 590 333a 591 592 594 593 595 596 597 598 778a 599 600 833a 601 602 603 604 605 606 636a 607 608 609 610 611 612 613 614 615 616 617 618 619 620 621 622

MESA ET AL.

Tenuipalpus cyatheae Tenuipalpus daneshvari Tenuipalpus danxianensis Tenuipalpus dasples Tenuipalpus decus Tenuipalpus dimensus Tenuipalpus disparilis Tenuipalpus dombeyae Tenuipalpus dominguensis Tenuipalpus dubinini Tenuipalpus dumus Tenuipalpus elegans Tenuipalpus elongatus Tenuipalpus emeticae Tenuipalpus engelbrechti Tenuipalpus ephedrae Tenuipalpus erasus Tenuipalpus eremitus Tenuipalpus eriophyoides Tenuipalpus eucleae Tenuipalpus eugeniae Tenuipalpus euonymi Tenuipalpus falcatus Tenuipalpus faresianus Tenuipalpus faveolus Tenuipalpus feliciae Tenuipalpus ferosus Tenuipalpus fici Tenuipalpus filicicola Tenuipalpus,flacourtiae Tenuipalpus flechtmanni Tenuipalpus frondosus Tenuipalpus galpiniae Tenuipalpus garciniae Tenuipalpus gatoomensis Tenuipalpus geigeriae Tenuipalpus geisenheyneri Tenuipalpus ghaii Tenuipalpus ghanii Tenuipalpus gibbus Tenuipalpus glaber Tenuipalpus granati Tenuipalpus grevilleae Tenuipalpus guamensis Tenuipalpus guettardae Tenuipalpus gumbolimbonis Tenuipalpus guptai Tenuipalpus haripuriensis Tenuipalpus hastaligni Tenuipalpus heteropyxis Tenuipalpus heveae Tenuipalpus hondurensis Tenuipalpus hornotinus Tenuipalpus hurae Tenuipalpus ilocanus

623 624 625 626 627 628 629 631 630 632 633 634 635 636 637 638 639 640 641 642 643 644 645 646 647 648 649 650 651 652 653 654 656 657 658 659 439a 655 628a 881a 439b 660 661 662 663 664 665 666 667 668 669 670 671 672 673

Tenuipalpus imias Tenuipalpus indicus Tenuipalpus inophylli Tenuipalpus inornatus Tenuipalpus insularis Tenuipalpus isabelae Tenuipalpus ixorae Tenuipalpus jagatkhanaens Tenuipalpus jamaicensis Tenuipalpus japonicus Tenuipalpus jasmini Tenuipalpus jawadii Tenuipalpus jianfengensis Tenuipalpus jonkeri Tenuipalpus jordaani Tenuipalpus jussiaeae Tenuipalpus kamalii Tenuipalpus kapoki Tenuipalpus karrooi Tenuipalpus keiensis Tenuipalpus kenos Tenuipalpus kesari Tenuipalpus knorri Tenuipalpus kobachidzei Tenuipalpus kraussianae Tenuipalpus lalbaghensis Tenuipalpus laminasetae Tenuipalpus lanceae Tenuipalpus lapidosus Tenuipalpus latiseta Tenuipalpus lawrencei Tenuipalpus legatus Tenuipalpus leipoldti Tenuipalpus leonorae Tenuipalpus leucospermi Tenuipalpus lineosetosus Tenuipalpus lucumae Tenuipalpus ludhianaensis Tenuipalpus lulinicus Tenuipalpus lunatus Tenuipalpus lustrabilis Tenuipalpus lycioides Tenuipalpus lygodii Tenuipalpus magalismontani Tenuipalpus mahoensis Tenuipalpus malligai Tenuipalpus mallotae Tenuipalpus mandraensis Tenuipalpus mansoni Tenuipalpus mansoniculus Tenuipalpus matthyssei Tenuipalpus melhaniae Tenuipalpus menglunensis Tenuipalpus metis Tenuipalpus metopii

A catalog of the Tenuipalpidae of the world with a key to genera

674 675 676 269f 677 678 679 680 681 682 683 684 685 686 687 688 689 690 691 692 693 694 695 696 697 698 699 700 641a 701 702 703 704 705 706 707 708 709 710 711 712 713 714 715 716 717 718 719 720 721 722 723 724 725 726

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Tenuipalpus micheli Tenuipalpus microphylli Tenuipalpus mkuziensis Tenuipalpus molinai Tenuipalpus montanus Tenuipalpus mopaneae Tenuipalpus moraesi Tenuipalpus morianus Tenuipalpus mourerae Tenuipalpus muguanicus Tenuipalpus mustus Tenuipalpus myrtus Tenuipalpus namaensis Tenuipalpus nenaxi Tenuipalpus niekerkae Tenuipalpus nigerianus Tenuipalpus obvelatus Tenuipalpus oliveirai Tenuipalpus ombrensis Tenuipalpus orchidofilo Tenuipalpus oribiensis Tenuipalpus orilloi Tenuipalpus ortus Tenuipalpus oudemansi Tenuipalpus ovalis Tenuipalpus oxalis Tenuipalpus pacificus Tenuipalpus pagesae Tenuipalpus pagina Tenuipalpus palmatus Tenuipalpus palosapis Tenuipalpus panici Tenuipalpus papiothalensis Tenuipalpus pareriophyiodes Tenuipalpus parsii Tenuipalpus pedrus Tenuipalpus pernicis Tenuipalpus persicae Tenuipalpus philippinensis Tenuipalpus pieteri Tenuipalpus pigrus Tenuipalpus pisinnus Tenuipalpus placitus Tenuipalpus platycaryae Tenuipalpus podocarpi Tenuipalpus populi Tenuipalpus portulacae Tenuipalpus proctori Tenuipalpus proteae Tenuipalpus protectus Tenuipalpus protumidus Tenuipalpus pruni Tenuipalpus prunioides Tenuipalpus pseudocedrelae Tenuipalpus pseudocuneatus

727 728 729 730 731 732 733 734 735 736 737 738 739 740 741 742 743 744 745 746 747 748 749 433d 750 751 752 753 754 597c 755 756 757 758 759 760 76 762 763 764 765 766 767 768 769 770 771 772 773 774 775 776 777 778 269g

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Tenuipalpus punicae Tenuipalpus punjabensis Tenuipalpus pyroides Tenuipalpus pyrusae Tenuipalpus qingchengensis Tenuipalpus rangiorae Tenuipalpus raphiae Tenuipalpus raptor Tenuipalpus rarus Tenuipalpus reticulus Tenuipalpus rhagicus Tenuipalpus rhizophorae Tenuipalpus rhusi Tenuipalpus rhysus Tenuipalpus robustae Tenuipalpus rillus Tenuipalpus rodionovi Tenuipalpus rosae Tenuipalpus rusapensis Tenuipalpus sagittus Tenuipalpus salicis Tenuipalpus sanblasensis Tenuipalpus sandyi Tenuipalpus santae Tenuipalpus sanyaensis Tenuipalpus scitulus Tenuipalpus sclerocaryae Tenuipalpus senecionis Tenuipalpus sharmai Tenuipalpus simarubae Tenuipalpus simplex Tenuipalpus simplychus Tenuipalpus smithi Tenuipalpus solanensis Tenuipalpus sophiae Tenuipalpus sparsus Tenuipalpus spathiphyllus Tenuipalpus spatulatus Tenuipalpus stativus Tenuipalpus stefani Tenuipalpus striolatus Tenuipalpus swieteniae Tenuipalpus szarvasensis Tenuipalpus tabebuiae Tenuipalpus taonicus Tenuipalpus tapirirae Tenuipalpus tauricus Tenuipalpus tectonae Tenuipalpus tepicanus Tenuipalpus terminaliae Tenuipalpus tetrazygiae Tenuipalpus toowongi Tenuipalpus tortulus Tenuipalpus transvaalensis Tenuipalpus trichiliae

779 780 781 782 783 784 785 786 787 788 789 790 791 792 793 620a 794 795 796 797 798 799 800 801 802 803 804 805 806 807 808 809 810 811 812 813 748a 814 815 816 817 567a 601a 818 819 820 821 822 823 824 827 826 825 828 829

MESA ET AL.

Tenuipalpus trifoliatae Tenuipalpus trisegmentus Tenuipalpus trisetosus Tenuipalpus tuttlei Tenuipalpus ueckermanni Tenuipalpus umarii Tenuipalpus unimerus Tenuipalpus unonopsonis Tenuipalpus uvae Tenuipalpus velitor Tenuipalpus venustus Tenuipalpus vernoniae Tenuipalpus vexus Tenuipalpus victoriae Tenuipalpus vitexi Tenuipalpus viticola Tenuipalpus vitis Tenuipalpus vriddagiriensis Tenuipalpus waqasii Tenuipalpus xerocolus Tenuipalpus xylosmae Tenuipalpus yarensis Tenuipalpus yousefi Tenuipalpus zanthus Tenuipalpus zeyheri Tenuipalpus zhengzhouensis Tenuipalpus zhizhilashviliae Tenuipalpus zuluensis tepicanus Tenuipalpus tepicbutilonae Brevipalpus tepicensis Brevipalpus terminaliae Tenuipalpus Terminalichus dehlonensis Terminalichus delhiensis Terminalichus karachiensis Terminalichus panajiensis Terminalichus psidi Terminalichus serratus testudinalis, Brevipalpus tetraflagellatus, Brevipalpus tetrazygiae, Tenuipalpus theae, Brevipalpus thelycraniae, Brevipalpus thomissus, Priscapalpus thujae, Aegyptobia tiliae, Brevipalpus tini, Acarus tinsukiaensis, Brevipalpus tiwanensis, Brevipalpus tlaxcensis, Brevipalpus toowongi, Tenuipalpus torreyi, Aegyptobia tortulus, Tenuipalpus tragardhi, Aegyptobia transitans, Larvacarus

830 831 832 833 834 835 836 837 838 839 840 841 842 843 844 845 136e 846 847 848 849 850 851 852 853 854 855 856 823 361 362 824 858 857 859 860 861 862 363 112a 827 364 365 522 84 366 597a 367 368 294a 826 85 825 86 475

transvaalensis, Tenuipalpus trichiliae, Tenuipalpus tridens, Pseudoleptus trifoliatae, Tenuipalpus trinidadensis, Brevipalpus trisegmentus, Tenuipalpus trisetosus, Tenuipalpus tuberellus, Brevipalpus turrialbensis, Brevipalpus tuttlei, Tenuipalpus ueckermanni, Tenuipalpus ueckermanni, Aegyptobia UItratenuipalpus asteliae Ultratenuipalpus aberrans Ultratenuipalpus acharis Ultratenuipalpus arboreus Ultratenuipalpus asteliicola Ultratenuipalpus baltazarae Ultratenuipalpus bunyai Ultratenuipalpus canelae Ultratenuipalpus carpodeti Ultratenuipalpus charlini Ultratenuipalpus coprosmae Ultratenuipalpus coprosmicus Ultratenuipalpus cyathodis Ultratenuipalpus filicicola Ultratenuipalpus gonianaensis Ultratenuipalpus hainanensis Ultratenuipalpus lacorpuzrarosae Urigeronus bunyai wainsteini, Cenopalpus waqasii, Tenuipalpus wiesei, Dolichotetranychus woglumi, Brevipalpus womersleyi, Ultratenuipalpus xeroclei, Capedulia xerocolus, Tenuipalpus xerophilus, Aegyptobia xerophilus, Aegyptobia xianensis, Phytoptipalpus xini, Cenopalplus xinjiangensis, Aegyptobia xylosmae, Tenuipalpus xystus, Brevipalpus yarensis, Tenuipalpus yothersi, Brevipalpus younguisti, Ultratenuipalpus yousefi, Tenuipalpus zaitzevi, Aegyptobia zanthus, Tenuipalpus zelihae, Pseudoleptus zempoalensis, Brevipalpus zeyheri, Tenuipalpus zhengzhouensis, Tenuipalpus zhenzhumeis, Brevipalpus

A catalog of the Tenuipalpidae of the world with a key to genera

828 829 533 830 369 831 832 370 371 833 834 87 866 863 864 865 867 868 869 870 871 872 873 874 875 876 889 877 878 890 445 847 472 136c 885 386 848 91 91 519 446 92 849 378 850 292e 886 851 93 852 534 379 853 854 380

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zhizhilashviliae, Tenuipalpus ziaii, Brevipalpus zinniae, Brevipalpus zoysiae, Dolichotetranychus zuluensis, Tenuipalpus Ultratenuipalpus meekeri Ultratenuipalpus nothofagi Ultratenuipalpus pteriphilus Ultratenuipalpus quadrisetosus Ultratenuipalpus rubi Ultratenuipalpus umtataensis Ultratenuipalpus womersleyi Ultratenuipalpus younguisti umarii, Tenuipalpus umtataensis, Ultratenuipalpus unimerus, Tenuipalpus unonopsonis, Tenuipalpus utriculus, Brevipalpus uvae, Tenuipalpus vandergooti, Dolichotetranychus vannus, Aegyptobia variolatus, Brevipalpus velitor, Tenuipalpus

855 381 382 473 856 879 880 881 882 883 884 885 886 835 884 836 837 372 838 471 88 373 839

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MESA ET AL.

HOST INDEX Abies alba, Pinaceae: 423 Abies grandis, Pinaceae: 205 Abies nordmanniana, Pinaceae: 601 Abies religiosa, Pinaceae: 101, 114, 265, 324, 347, 379 Abutilon austroafricanum, Malvaceae: 552a Abutilon sp., Malvaceae: 361 Acacia arabica, Fabaceae: 554 Acacia farnesiana, Fabaceae: 646 Acacia farnesiana, Fabaceae: 675 Acacia galpinii, Fabaceae; 656 Acacia gerrardii, Fabaceae: 745 Acacia greggii, Fabaceae; 2 Acacia karroo, Fabaceae: 553, 514, 515, 691, 705, 812 Acacia nigrescens, Fabaceae: 711 Acacia nilotica, Fabaceae: 503, 697 Acacia pennatula, Fabaceae: 69 Acacia sp., Fabaceae: 517 Acacia tortilis, Fabaceae: 810 Acacia xanthophloea, Fabaceae: 557 Acantholimon lepturoides, Plumbaginaceae: 91 Acer negundo, Aceraceae: 253 Achras sapota, Sapotaceae: 603, 680 Actinostemon sp., Euphorbiaceae: 678 Acuba sp., Cornaceae: 185 Adenostoma fasciculatum, Rosaceae: 8 Adenostoma sp., Rosaceae: 251, 432 Adhatoda vasica, Acanthaceae: 271 Aeluropus sp., Poaceae: 534 Aerva javanica, Amaranthaceae: 815 Aerva sp., Amaranthaceae: 847 Aextoxicon punctatum, Aextoxicaceae: 872 Albizia harveyi, Fabaceae: 509, 585 Albizia procera, Fabaceae: 504 Alhagi maurorum, Fabaceae: 703 Allamanda cathartica, Apocynaceae: 217 Allenrolfea occidentalis, Chenopodiaceae: 110 Allionia incarnate, Nyctaginaceae: 5, 45, 220 Allium pulchellum, Alliaceae: 547 Allocasuarina luehmannii, Casuarinaceae: 447, 478 Allocasuarina muelleriana, Casuarinaceae: 27, 449 Allocasuarina verticillata, Casuarinaceae: 450 Alnus arguta, Betulaceae: 111 Alphitonia excelsa, Rhamnaceae: 100, 476 Amaranthus sp., Amaranthaceae: 135 Ambrosia confertiflora, Asteraceae: 7 Ambrosia ilicifolia, Asteraceae: 39 Ambrosia sp., Asteraceae: 113 Amygdalus communis Rosaceae: 562 Anacardium sp., Anacardiaceae: 563 Andrachne sp., Euphorbiaceae: 269 Anisoptera thurifera, Dipterocarpaceae: 755 Annona sp., Annonaceae: 565 Antennaria plantaginifolia, Asteraceae182

Anthospermum aethiopicum, Rubiaceae 21 Apodytes dimidiate, Icacinaceae: 741 Aptosimum marlothii, Scrophulariaceae: 66c Aptosimum spinescens, Scrophulariaceae: 60a Araucaria bidwilli, Araucariaceae: 869, 890d Arbutus sp., Ericaceae: 10, 484 Arbutus unedo, Ericaceae: 389, 569 Arctostaphylos bicolor, Ericaceae: 261 Ardisia revolute, Myrsinaceae: 119 Aristida namaquensis, Poaceae: 464 Artemisia capillaries, Asteraceae: 11 Artemisia carruthii, Asteraceae: 121 Artemisia filifolia, Asteraceae: 191 Artemisia scoparia, Asteraceae: 38 Artemisia sp., Asteraceae: 35 Artocarpus integrifolia, Moraceae: 698 Arundina graminifolia, Orchidaceae: 746 Aspalathus acuminate, Fabaceae: 592 Aspalathus linearis, Fabaceae: 385 Aspilia africana, Asteraceae: 742 Aspilia natalensis, Asteraceae: 561 Astelia sp., Asteliaceae: 866, 867 Aster alpinus: Asteraceae: 99 Aster exilis, Asteraceae: 123 Atamisquea emarginata, Capparaceae: 179 Athrixia elata, Asteraceae: 572 Atraphaxis spinosa, Polygonaceae: 93 Atriplex canescens, Chenopodiaceae: 12, 16, 28 Atriplex crassifolia, Chenopodiaceae: 640 Atriplex sp., Chenopodiaceae: 284 Aureolaria flava, Scrophulariaceae: 226 Austrocedrus chilensis, Cupressaceae: 576 Azadirachta indica, Meliaceae: 208 Baccharis conferta, Asteraceae: 187 Baccharis glutinosa, Asteraceae: 13 Baccharis heterophylla, Asteraceae: 126 Baccharis heterophylla, Asteraceae: 262 Baccharis pilularis, Asteraceae: 235a Baccharis sp., Asteraceae: 153 Banksia spinulosa, Proteaceae: 582 Bassia latifolia, Chenopodiaceae: 584 Bauhinia purpurea, Fabaceae: 217, 737 Bauhinia sp., Fabaceae: 128 Bequaertiodendron magalismontanum, Sapotaceae: 715 Berkheya carlinopsis, Asteraceae 775 Berkheya radula, Asteraceae: 586 Biota orientalis, Cupressaceae: 43, 493 Boscia albitrunca, Capparaceae: 828 Bouchea prismatica, Verbenaceae: 132 Bouteloua gracilis, Poaceae: 63, 75, 526 Brachyglottis repanda, Asteraceae: 784 Brachystegia manga, Fabaceae: 796 Brassica oleracea, Brassicaceae: 694

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Brassica rapa, Brassicaceae: 321 Bucida buceras, Combretaceae: 133 Bucida buceras, Combretaceae: 589 Buddleja asiatica, Buddlejaceae: 323 Buddleja saligna, Buddlejaceae: 338 Bumelia sp., Sapotaceae: 134 Bursera simaruba, Burseraceae: 664, 807 Bursera sp., Burseraceae: 590 Buxus microphylla, Buxaceae: 519 Camelli sinensis, Theaceae: 217 Campomanesia sp., Myrtaceae: 653 Campsis radicans, Bignoniaceae: 19 Cannabis sativa, Cannabaceae: 666a Capparis sepiaria, Capparaceae: 593 Capsicum frutescens, Solanaceae: 334 Carex sp., Cyperaceae: 890 Carica papaya, Caricaceae: 736 Carica sp., Caricaceae: 130 Carpinus orientalis, Betulaceae: 395 Carpodetus serratus, Carpodetaceae: 871 Caryota ochlandra, Arecaceae: 217 Cassia covesii, Fabaceae: 20 Cassia crotolarioides, Caesalpiniaceae: 138 Cassia fistula, Fabaceae: 116 Cassia sp., Fabaceae: 803 Cassine transvaalensis, Celastraceae: 546 Castilleja sp., Scrophulariaceae: 139 Casuarina equisetifolia, Casuarinaceae; 500 Casuarina sp., Casuarinaceae: 548 Catalpa longissima, Bignoniaceae: 630 Catophractes alexandri, Bignoniaceae: 505 Ceanothus fendleri, Rhamnaceae: 140 Cedrela mexicana, Meliaceae: 778 Cedrela sp., Meliaceae: 598, 615 Ceiba acuminate, Bombacaceae: 373 Ceiba pentandra, Bombacaceae: 690 Ceiba sp.Bombacaceae: 506a Celmisia laricifolia, Asteraceae: 451 Celtis pallida, Ulmaceae: 141 Celtis reticulate, Ulmaceae: 295 Celtis sinensis, Ulmaceae: 217, 497 Celtis sp.Ulmaceae: 599 Cerastium argenteum, Caryophyllaceae: 91 Cercidium floridum, Caesalpiniaceae: 142, 507 Chaetachme aristata. Ulmaceae: 727 Chamaedorea sp., Arecaceae: 144, 227, 286, 600, 833, 833 Chiococca pinetorum, Rubiaceae: 605 Chionochloa sp., Poaceae: 451 Chromolaena odorata, Asteraceae: 579 Chrysalidocarpus lutescens, Arecaceae: 217a Chrysanthemum sp., Asteraceae: 761 Cissampelos pareira, Menispermaceae: 606 Cistus tauricus, Cistaceae: 498 Citrus sp., Rutaceae: 136, 136d, 228a, 636, 748 Citrus jambhiri, Rutaceae: 225

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Citrus limon, Rutaceae: 252, 889 Citrus reticulate, Rutaceae: 230, 292 Citrus? sp., Rutaceae: 269 Clematis heracleifolia, Ranunculaceae: 608 Clematis vitalba, Ranunculaceae: 696 Clerodendrum cyrtophyllum, Verbenaceae: 166d Cliffortia ramosissima, Rosaceae: 95 Cnidosculos sp., Euphorbiaceae: 148a Coccoloba aurifolia, Polygonaceae: 610 Cochlospermum sp., Cochlospermaceae: 149 Cochlospermum vitifolium, Cochlospermaceae: 197 Cocos nucifera, Arecaceae: 458, 539 Codiaeum inophyllum, Euphorbiaceae: 676 Codiaeum variegatum, Euphorbiaceae: 224, 309a Coffea arabica, Rubiaceae: 150 Coldenia greggii, Boraginaceae: 151 Colophospermum mopane, Fabaceae: 732 Combretum apiculatum, Combretaceae: 633 Combretum farinosum, Combretaceae: 155 Combretum molle, Combretaceae: 613, 734 Combretum paniculatum, Combretaceae: 728 Combretum sp., Combretaceae: 658 Conocarpus erecta, Combretaceae: 112, 156 Conostegia xalapensis, Melastomataceae: 508 Convolvulus sp., Convolvulaceae: 233 Convolvulus umbellatus, Convolvulaceae: 348 Coprosma sp., Rubiaceae: 873, 874 Corchorus capsularis, Tiliaceae: 160 Cordia alba, Boraginaceae: 168 Cordia boissieri, Boraginaceae: 157 Cordia elaeagnoides, Boraginaceae: 175 Cordia glabra, Boraginaceae: 239 Cordia rothii, Boraginaceae: 266 Cordia sulcata, Boraginaceae: 809 Cornulaca monacantha, Chenopodiaceae: 1, 57 Cornus sanguinea, Cornaceae: 365, 439 Cotoneaster sp., Rosaceae: 425 Couroupita guianensis, Lecythidaceae: 616 Crassula lycopodioides, Crassulaceae: 618 Crataegus oxyacantha, Rosaceae: 391, 397a Crataegus sp., Rosaceae: 159 Croton ciliatoglandulifer, Euphorbiaceae: 162, 163 Croton corymbulosus, Euphorbiaceae: 23 Croton sp., Euphorbiaceae: 186 Cryptocarya woodii, Lauraceae: 881, 882 Cucurbita maxima, Cucurbitaceae: 164 Cucurbita pepo, Cucurbitaceae: 212 Cupressus forbesii, Cupressaceae: 24 Cupressus sempervirens, Cupressaceae: 34a, 65, 80, 622 Cupressus sp., Cupressaceae: 46, 54, 55, 87 Cyathea medullaris, Cyatheaceae: 623 Cyathodes fasciculate, Epacridaceae: 875 Cyclosorus sp., Thelypteridaceae: 651, 876 Cydonia sp., Rosaceae: 794 Cynodon dactylon, Poaceae: 454

MESA ET AL.

Cyrilla racemiflora, Cyrillaceae: 792 Datura inoxia, Solanaceae: 512 Delonix regia, Caesalpiniaceae: 189 Dendrobium sp., Orchidaceae: 471 Desmodium sp., Fabaceae: 169 Diastella divaricata, Proteaceae: 96 Dichrostachys cinerea, Fabaceae: 516 Diospyros kaki, Ebenaceae: 682, 855 Diospyros lycioides, Ebenaceae: 713 Dipholis salicifolia, Sapotaceae: 171 Distichlis scoparia, Poaceae: 525 Distichlis spicata, Poaceae: 468, 530 Ditaxis lanceolata, Euphorbiaceae: 305 Dodonea viscose, Sapindaceae: 308 Dombeya rotundifolia, Sterculiaceae: 631 Drymis winteri, Winteraceae: 870 Echinocactus sp., Cactaceae: 333 Eclipta prostrate, Asteraceae: 559 Elettaria cardamomum, Zingiberaceae: 461 Elytropappus gnaphaloides, Asteraceae: 394 Encelia farinose, Asteraceae: 180 Ephedra distachya, Ephedraceae: 638 Ephedra fasciculate, Ephedraceae: 30 Ephedra procera, Ephedraceae: 68, 632b Eragrostis tenella, Poaceae: 470 Erica manipuliflora, Ericaceae: 438 Erica sp., Ericaceae: 184 Erica walkeria, Ericaceae: 32 Eriobotrya japonica, Rosaceae: 400, 408 Eriodictyon californicum, Hydrophyllaceae: 124, 274 Eriogonum wrightii, Polygonaceae: 33 Erythrina sp., Fabaceae: 313 Escallonia sp., Escalloniaceae: 158 Eucalyptus micrantha, Myrtaceae: 477 Eucalyptus sp., Myrtaceae: 535, 826 Euclea crispa, Ebenaceae: 621 Euclea divinorum, Ebenaceae: 642, 856 Eugenia biflora, Myrtaceae: 643 Eugenia sp., Myrtaceae: 538 Euonymus sp., Celastraceae: 644 Eupatorium glabratum, Asteraceae: 178 Eupatorium hemiteropodum, Asteraceae: 108 Euphorbia echinus, Euphorbiaceae: 619 Euphorbia sp., Euphorbiaceae: 816 Fagonia cretica, Zygophyllaceae: 754 Fallugia paradoxa, Rosaceae: 36 Felicia hirsuta, Asteraceae: 648 Ferocactus sp., Cactaceae: 40 Ficus carica, Moraceae: 161, 650 Flacourtia indica, Flacourtiaceae: 652 Flourensia cernua, Asteraceae: 37 Forestiera porulosa, Oleaceae: 270 Frankenia palmeri, Frankeniaceae: 195 Franseria deltoidea, Asteraceae: 106 Fraxinus excelsior, Oleaceae: 689

Fraxinus profunda, Oleaceae: 196 Fraxinus sp., Oleaceae: 390 Garcinia sp., Clusiaceae: 657 Garrya flavescens, Garryaceae: 307 Geigeria aspera, Asteraceae: 659 Geranium sp., Geraniaceae: 200 Gevuina avellana, Proteaceae: 864 Gliricidia sepium, Fabaceae: 201 Gossypium hirsutum, Malvaceae: 381 Grevillea sericea, Proteaceae: 661 Grewia robusta, Tiliaceae: 793 Grewia bicolor, Tiliaceae: 474 Grewia microcos, Tiliaceae: 206 Guarea ramiflora, Meliaceae: 127 Guazuma tomentosa, Sterculiaceae: 258 Guettarda scabra, Rubiaceae: 663 Gutierrezia californica, Asteraceae: 52, 316a Gutierrezia lucida, Asteraceae: 49 Gymnosporia royleana, Celastraceae: 188 Halleria lucida, Scrophulariaceae: 418 Hamelia erecta, Rubiaceae: 183, 210 Hammada scoparia, Chenopodiaceae: 31 Haplopappus acradenius, Asteraceae9 Haplopappus spinulosus, Asteraceae: 9, 42 Haplopappus venutus, Asteraceae: 375 Hebe salicifolia, Scrophulariaceae: 452 Hedera sp., Araliaceae: 405 Helianthus annuus, Asteraceae: 211, 325, 285, 339, 806 Helichrysum cymosum, Asteraceae: 460 Helichrysum petiolare, Asteraceae: 647 Helichrysum rugulosum, Asteraceae: 637 Heliotropium sp., Boraginaceae: 41, 50 Heliotropium strigosum, Boraginaceae: 766 Hemizonia virgata, Asteraceae: 14 Hernandia catalpifolia, Hernandiaceae: 214 Heterocentron subtriplinervium, Melastomataceae: 216 Heteropyxis natalensis, Heteropyxidaceae: 668 Hevea brasiliensis, Euphorbiaceae : 217 Hibiscus esculentus, Malvaceae: 105, 248 Hibiscus sp., Malvaceae: 481 Hilaria rigida, Poaceae: 529 Hippomane mancinella, Euphorbiaceae: 674 Hohenbergia proctori, Bromeliaceae: 772 Humiria balsamifera, Humiriaceae: 800 Hura crepitans, Euphorbiaceae: 672 Hymenoclea pentalepis, Asteraceae: 44 Hypoestes sp., Acanthaceae: 692 Hyptis albida, Labiatae: 219 Hyptis sp., Lamiaceae: 264, 362 Ichthyomethia piscipula, Fabaceae: 378 Inula vulgaris, Asteraceae: 322 Ipomoea biloba, Convolvulaceae: 813 Ipomoea cairica, Convolvulaceae : 217 Ipomoea plummerae, Convolvulaceae: 223 Ipomoea sp. , Convolvulaceae : 217

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Ixora parviflora, Rubiaceae: 679 Jasminum grandiflorum, Oleaceae: 199 Jasminum humile, Oleaceae: 628 Jasminum sambac, Oleaceae: 260 Jasminum sp., Oleaceae: 683 Juniperus chinensis, Cupressaceae: 22 Juniperus communis, Cupressaceae: 15, 48, Juniperus deppeana, Cupressaceae: 231a, 254, 312 Juniperus mexicanus, Cupressaceae: 231 Juniperus oxycedrus, Cupressaceae: 15 Juniperus sp., Cupressaceae: 64, 90, 483, 492 Jussiaea sp., Onagraceae: 688 Kochia prostrata, Chenopodiaceae: 92 Lagascea angustifolia, Asteraceae: 236 Lagerstroemia speciosa, Lythraceae: 753 Lagonychium farctum, Fabaceae: 577, 578 Lannea grandis, Anacardiaceae: 802 Lantana camara, Verbenaceae: 166 Lantana sp., Verbenaceae: 237 Lapageria rosea, Philesciaceae: 117 Lasianthaea fruticosa, Asteraceae: 272 Latania loddigesii, Arecaceae: 502 Lepidium sp., Brassicaceae: 238 Leptospermum sp., Myrtaceae: 885 Leucospermum conocarpodendron, Proteaceae: 706 Liabum glabrum, Asteraceae: 306 Libocedrus bidwillii, Cupressaceae: 840 Libocedrus decurrens, Cupressaceae: 3, 62, 167, 494 Licaria sp., Lauraceae: 194 Ligustrum amurense, Oleaceae: 269 Lilium regale, Liliaceae: 241e Limonium meyeri, Plumbaginaceae: 384 Lippia sp., Verbenaceae: 243 Lithops erniana, Aizoaceae: 448 Litsea glutinosa, Lauraceae: 673 Lonchocarpus capassa, Fabaceae: 594 Lotus sp., Fabaceae: 246 Loxostylis alata, Anacardiaceae: 747 Lucuma salicifolia, Sapotaceae: 708 Luffa acutangula, Cucurbitaceae: 349 Lupinus sp., Fabaceae: 247, 263, 287 Lygodium volubile, Lygodiaceae: 714 Lygodium japonicum, Lygodiaceae: 877 Lysiloma bahamensis, Fabaceae: 249 Magnolia denudata, Magnoliaceae: 854 Magnolia sp., Magnoliaceae: 580 Mallotus sp., Euphorbiaceae: 718 Malus sp., Rosaceae: 407 Malvastrum sp., Malvaceae: 372 Mammillaria sp., Cactaceae: 333 Mangifera indica, Anacardiaceae: 67, 122, 152, 513 Manilkara zapota, Sapotaceae: 625 Maytenus heterophylla, Celastraceae: 614 Medicago sp., Fabaceae: 58 Melanolepis multiglandulosa, Euphorbiaceae: 722

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Melhania forbesii, Sterculiaceae: 723 Melicytus ramiflorus, Violaceae: 568, 716 Mespilus sp., Rosaceae: 414 Metopium toxiferum, Anacardiaceae: 726 Miconia foveolata, Melastomataceae: 102 Miconia sp., Melastomataceae: 520 Momordica charantia, Cucurbitaceae: 221, 416 Morinda boninensis, Rubiaceae: 587 Morus alba, Moraceae: 258, 298, 749, 831 Mourera sideroxylon, Melastomataceae: 735 Muhlenbergia asperifolia, Poaceae: 465 Muhlenbergia torreyi, Poaceae: 85, 466 Musa paradisiaca, Musaceae: 399 Myrica mexicana, Myricaceae: 327 Myrtus communis, Myrtaceae: 738 Nebelia sp., Bruniaceae: 98 Nenax microphylla, Rubiaceae: 740 Neottopteris nidus, Aspleniaceae: 662 Nerium indicum, Apocynaceae: 780 Nothofagus fusca, Fagaceae: 880 Nothofagus spp., Fagaceae: 875 Ocimum basilicum, Lamiaceae: 229, 354a, 367 Ocotea acutangula, Lauraceae: 104 Ocotea coriacea, Lauraceae: 273, 667 Olea africana, Oleaceae: 421 Olea cuspidata, Oleaceae: 627 Olea europaea, Oleaceae: 125, 174, 204, 213, 278, 279, 280, 328, 351 Oncidium sp., Orchidaceae: 281 Oreopanax peltatum, Araliaceae: 282 Origanum sp., Labiatae: 269b Osmanthus fragrans, Oleaceae: 207 Osteospermum calendulaceum, Asteraceae: 383c Oxalis sp., Oxalidaceae: 751 Pandanus sp., Pandanaceae: 543 Panicum maximum, Poaceae: 756 Panicum obtusum, Poaceae: 531 Panicum repens, Poaceae: 467 Parthenium incanum, Asteraceae: 288, 302, 302c, 352a Passerina falcifolia, Thymelaeaceae: 486 Peganum harmala, Zygophyllaceae: 787 Peltophorum africanum, Fabaceae: 558, 834 Penstemon davidsonii, Scrophulariaceae: 59 Persea hintonii, Lauraceae: 290 Persea popenoei, Lauraceae: 192 Persea schiedeana, Lauraceae: 190 Persea (Tamala) borbonia, Lauraceae: 193 Peschiera fuchsiaefolia, Apocynaceae: 744 Petrea sp., Verbenaceae: 292 Petrophytum caespitosum, Rosaceae: 259 Phalaenopsis stuartiana, Orchidaceae: 752a Pharbitis nil, Convolvulaceae: 217, 318 Phoebe tampicensis, Lauraceae: 370 Phoenix dactylifera, Arecaceae: 536, 537a, 541, 542a, 544, 545, 641, 758, 850a

MESA ET AL.

Phoenix sp., Arecaceae: 292 Phoradendron bolleanum, Viscaceae: 6 Phoradendron sp., Viscaceae: 301 Photinia arbutifolia, Rosaceae: 343 Photinia sp., Rosaceae: 217 Phragmites australis, Poaceae: 455 Phyla nodiflora, Verbenaceae: 267 Phylica sp., Rhamnaceae: 612 Phyllanthus sp., Euphorbiaceae: 564 Phyllocladus alpinus, Podocarpaceae: 865 Physalis wrightii, Solanaceae: 71 Phytolacca sp., Phytolaccaceae: 269 Pimelea sericeovillosa, Thymelaeaceae: 731 Pinus halepensis, Pinaceae: 403a Pinus hamata, Pinaceae: 412 Pinus oocarpa, Pinaceae: 670 Pinus radiate, Pinaceae: 294b Pinus silvestris, Pinaceae: 131 Pinus sp., Pinaceae: 294, 402, 437a, 445 Pinus wallichiana, Pinaceae: 393a Pistacia lentiscus, Anacardiaceae: 427 Plantago lanceolata, Plantaginaceae: 70, 759 Platanus occidentalis, Platanaceae: 154 Platanus sp., Platanaceae: 415, 428 Platycarpa strobilacea, Juglandaceae: 768 Pluchea odorata, Asteraceae: 299 Pluchea purpurascens, Asteraceae: 118 Pluchea sericea, Asteraceae: 107 Plumbago auriculata, Plumbaginaceae: 575 Plumeria rubra, Apocynaceae: 217 Podocarpus elongate, Podocarpaceae: 635 Podocarpus falcate, Podocarpaceae: 645, 769 Podocarpus usambarensis, Podocarpaceae: 573 Pomaderris rugosa, Rhamnaceae: 72 Pomaderris sp., Rhamnaceae: 721 Populus alba, Salicaceae: 61a, 401 Populus nigra, Salicaceae: 446, 770 Populus sp., Salicaceae: 26, 73, 424, 429, 707 Portulaca oleracea, Portulacaceae: 771 Potentilla hippiana, Rosaceae: 304 Pouteria sp., Sapotaceae: 588 Pouzolzia mixta, Urticaceae: 555 Primula vulgaris, Primulaceae: 439 Prosopis glandulosa, Fabaceae: 88 Protea nana, Proteaceae: 773e Prunus amygdalus, Rosaceae: 406, 725 Prunus amygdalus, Rosaceae: 725 Prunus armeniaca, Rosaceae: 410 Prunus domestica, Rosaceae: 410 Prunus persica, Rosaceae: 291, 693, 762, 776, 819 Psidium guajava, Myrtaceae: 851, 858, 861, 862, 888 Psilostrophe cooperi, Asteraceae: 314 Psilostrophe tagetina, Asteraceae: 359, 359a Psychotria grandis, Rubiaceae: 609 Psychotria sp., Rubiaceae: 315a

Pteleopsis myrtifolia, Combretaceae: 729 Pteridium aquilinum, Dennstaedtiaceae: 881 Pterocarpus indicus, Fabaceae: 763 Punica gramatum, Puicaceae: 147, 551, 604, 660, 665, 671, 712, 767, 779, 811, 835 Pycnanthemum pycnanthemoides, Lamiaceae: 317 Pyrus amygdaliformis, Rosaceae: 89 Pyrus communis, Rosaceae: 410, 430, 628, 709 Pyrus malus, Rosaceae: 396, 404, 409, 410a, 422, 782, 863 Qualea grandiflora, Vochysiaceae: 571, 595 Quercus aristata, Fagaceae: 109 Quercus arizonica, Fagaceae: 120 Quercus cerris, Fagaceae: 256 Quercus conzattii, Fagaceae: 268 Quercus lobata, Fagaceae: 276 Quercus magnoliaefolia, Fagaceae: 331 Quercus pubescens, Fagaceae: 322 Quercus sp., Fagaceae: 202, 222, 242ª, 257, 311, 319, 320, 322, 387, 413 Quercus sp.(near castanea), Fagaceae: 181 Raphia sp., Arecaceae: 785 Rhigozum trichotomum, Bignoniaceae: 510 Rhizophora mangle, Rhizophoraceae: 790 Rhoicissus tridentate, Vitaceae: 522 Rhus lancea, Anacardiaceae: 700, 739 Rhus macowanii, Anacardiaceae: 757 Rhus pyroides, Anacardiaceae: 781 Rhus undulate, Anacardiaceae: 704, 791 Rhynchosia monophylla, Fabaceae: 602 Rosa omeiensis, Rosaceae: 511 Rosa sp., Rosaceae: 518, 795 Rosmarinus officinalis, Lamiaceae: 420 Rubus sp., Rosaceae: 269, 322, 392d, 417, 419, 431, 434, 442, 883 Rubus cordifolius, Rosaceae: 329 Ruellia drymophila, Acanthaceae: 710 Ruellia nudiflora, Acanthaceae: 330 Ruellia repens, Acanthaceae : 217 Sabal megacarpa, Arecaceae: 626 Salix alba, Salicaceae: 77, 695, 798 Salix babylonica, Salicaceae: 607 Salix gooddingii, Salicaceae: 336 Salix sp., Salicaceae 78, 103, 337 Salvadora oleoides, Salvadoraceae: 639 Salvadora sp., Salvadoraceae: 684 Salvia africana-caerulea, Lamiaceae: 686 Salvia mellifera, Lamiaceae: 341 Salvia officinalis, Lamiaceae: 398 Salvia sp., Lamiaceae: 25, 340 Sambucus sp., Caprifoliaceae: 198 Scleria sphacelata, Cyperaceae: 891 Sclerocarya birrea ssp. caffra, Anacardiaceae: 804 Scolopia zeyheri, Flacourtiaceae: 853 Sebastiana brasiliensis, Euphorbiaceae: 733 "sedge, Gahnia aspera", Cyperaceae: 523, 524

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Senecio hectori, Asteraceae: 634 Senecio kirkii, Asteraceae: 805 Sida sp., Malvaceae: 310, 346 Solanum bahamense, Solanaceae: 358 Solanum elaeagnifolium, Solanaceae: 83, 163 Solanum melongena, Solanaceae: 345a, 624 Solanum nudum, Solanaceae: 115 Solanum tuberosum, Solanaceae: 177, 388 Sorbaria arborea, Rosaceae: 380Spathiphyllum sp., Araceae: 748 Spondias purpurea, Anacardiaceae: 335a Sporobolus cryptandrus, Poaceae: 459 Sporobolus sp., Poaceae: 457 Stipa ichu, Poaceae: 356 Styrax argenteus, Styracaceae: 146 Swietenia macrophylla, Meliaceae: 112, 566a, 567 Swietenia sp. mahogany, Meliaceae: 567a Syringa sp., Oleaceae: 217 Tabebuia pentaphylla, Bignoniaceae: 818 Tabebuia sp., Bignoniaceae: 720, 842 Tagetes tenuifolia, Asteraceae: 326 Tamarix articulata, Tamaricaceae: 441 Tamarix maris-mortui, Tamaricaceae: 480 Tamarix pentandra, Tamaricaceae: 479 Tapirira guianensis, Anacardiaceae: 820 Tarchonanthus camphoratus, Asteraceae: 591 Taxus baccata, Taxaceae: 499, 601 Taxus sp., Taxaceae: 499a Tectona grandis, Verbenaceae: 822 Terminalia amazonica, Combretaceae: 824 Terminalia arjuna, Combretaceae: 857 Terminalia catappa, Combretaceae: 859 Terminalia prunioides, Combretaceae: 777 Terminalia sericea, Combretaceae: 764, 817 Terminalia sp., Combretaceae: 860 Tetrazygia bicolor, Melastomataceae: 827 Tetrazygia pallens, Melastomataceae: 681 Thea sinensis, Theaceae: 364 Theobroma cacao, Sterculiaceae: 217 Thesium lineatum, Santalaceae: 51 Thuja occidentalis, Cupressaceae 84, 491 Thuja orientalis, Cupressaceae: 86, 303, 303a Thuja sp., Cupressaceae: 4, 485, 487, 495 Thymus sp., Lamiaceae: 821 Tilia americana, Tiliaceae: 176 Tilia heterophylla, Tiliaceae: 366 Tiliacora acuminata, Menispermaceae: 556 Tillandsia sp., Bromeliaceae: 453 Trianthema partulacastrum, Aizoaceae: 232 Trianthema portulacastrum, Aizoaceae: 232 Trianthema sp., Aizoaceae: 386 Trichilia hirta, Meliaceae: 112, 829 Trichilia emetica, Meliaceae: 636a Tridax procumbens, Asteraceae: 293 Tridens pulchellus, Poaceae: 533

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Umbellularia californica, Lauraceae: 765 unidentified, Asteraceae: 129, 215, 730, 814 unidentified, Cactaceae: 17, 18, 333, 353 unidentified, Caesalpiniaceae: 774 unidentified, Celastraceae: 701 unidentified, Cupressaceae: 79, 490 unidentified, Cyperaceae: 743 unidentified, Myrsinaceae: 581 unidentified, Poaceae: 97, 528 unidentified, Restionaceae: 797 unidentified, “apple”, Rosaceae: 433, 433a, 808 unidentified, “apricot”, Rosaceae: 433 unidentified, “Arbor-vitae”, Cupressaceae: 53b, 488d unidentified, “ash tree, bark of”: 137 unidentified, “bamboo”, Poaceae: 629 unidentified, “beech”, Fagaceae: 433 unidentified, “cereal”: 463 unidentified, “capulincillo” (cherry): Ericaceae: 823b unidentified, “chaparral”: 244 unidentified, “cuasticomate”, Bignoniaceae: 620g unidentified, “dates”, Arecaceae: 501 unidentified, “bermuda grass, Poaceae: 469 unidentified, “desert plant”: 76, 456 unidentified, “English walnut”, Juglandaceae: 292 unidentified, “fence tree”: 801 unidentified, “fern”: 868, 878, 879 unidentified, “fern tree”, Osmundaceae: 654 unidentified, “fir”:, Pinaceae: 435 unidentified, “goldenrod”, Asteraceae: 269, 596 unidentified, “grass & in soil”: 29, 74, 472 unidentified, “guamacate”, Lauraceae: 836 unidentified, “hedge plant”: 165, 433, 830, 846 unidentified, “herbaceous plant”: 47, 56 unidentified, “hickory”, Juglandaceae: 173, 342 unidentified, “lantana”, Verbenaceae: 369 unidentified, “large tree”: 702 unidentified, “laurestinus”, Caprifoliaceae: 597 unidentified, “lemon”, Rutaceae: 136, 136e, 145c, 240 unidentified, “malvaceous tree”, Malvaceae: 549 unidentified, “mat plants on rocks”: 560 unidentified, "Mata gusano": 357 unidentified, “mountain mahogany”: 170 unidentified, “naranjilla” Solanaceae: 799 unidentified, “oak”, Fagaceae: 275 unidentified, “oil palm”, Arecaceae: 617 unidentified, “olive”, Oleaceae: 143, 250, 277, 540 unidentified, “orange”, Rutaceae: 136, 407 unidentified, “orchid”, Orchidaceae: 136 unidentified, “ornamental plant”: 360 unidentified, “papaya”, Caricaceae: 292 unidentified, “pears", Rosaceae: 433 unidentified, “Pine”, Pinaceae: 296, 412 unidentified, “pine forest litter with rhododendrons” Ericaceae: 412 unidentified, “pineapple”, Bromeliaceae: 462

MESA ET AL.

unidentified, “plums", Rosaceae: 433 unidentified, “privet”: Oleaceae: 292 unidentified, “rubber tree”, Euphorbiaceae: 669 unidentified, “sapodilla”, Sapotaceae: 521 unidentified, “shrub” 583 unidentified, “unidentified plant”: 81, 81a, 82, 209, 234, 245, 255, 269, 283b, 332, 350, 350a, 368, 371, 374, 411, 426, 436, 440, 444, 527, 532, 597, 611b, 620, 641a, 649, 655, 699, 717, 719, 786, 839, 884, 886 unidentified, “unidentified shrub or young tree”: 843 unidentified, “unidentified tree”: 344, 363, 550, 760 unidentified, “uva” (a large tree): 838a unidentified, “weed”: 218 unidentified, “wild herb”: 848 unidentified, “wild lantana”: 203 unidentified, “wild plant”: 94, 687 unidentified, “wild shrub”: 750 unidentified, “wild tree”: 677 unidentified, “kind of arbor”: 724 unidentified: 439, 391a, 685 Unonopsis guatterioides, Annonaceae: 837 Urena lobata, Malvaceae: 217 Vaccinium ovatum, Ericaceae: 789 Verbena sp., Verbenaceae: 376 Verbesina sp., Asteraceae: 300 Vernonia oligocephala, Asteraceae: 841 Viburnum brachybotryum, Caprifoliaceae: 783 Viburnum tinus, Caprifoliaceae: 597c, 597 Viguiera sp., Asteraceae: 377 Viguiera stenoloba, Asteraceae: 355 Viola philippica, Violaceae: 574 Vitex harveyana, Verbenaceae: 844 Vitis vinifera, Vitaceae: 443, 788, 845, 887 Waltheria sp., Sterculiaceae: 297 Whitania somnifera, Solanaceae: 172, 289 Xylosma affnitida, Flacourtiaceae: 849 Xymalos monospora, Monimiaceae: 825 Yucca elata, Agavaceae: 832 Yucca gloriosa, Agavaceae: 570 Zanthoxylum elephantiasis, Rutaceae: 852 Zinnia acerosa, Asteraceae: 382 Ziziphus mauritiana, Rhamnaceae: 475 Zoysia tenuifolia, Poaceae: 473 Zygophyllum oetidum, Zygophyllaceae: 489 Zygophyllum simplex, Zygophyllaceae: 496

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